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		Curlycup Gumweed - Grindelia squarrosa
		
		
		
		
Native Species
			Global Rank: 
G5
			State Rank: 
S5
			
				C-value: 
3
			
			
			
			Agency Status
			USFWS: 
			USFS: 
			BLM: 
		
			
			
				
			
			
				External Links
				
			
		 
	 
	
	 
		General Description
		Biennial to perennial, sometimes branched at the caudex. Stems erect, branched above, 10–70 cm. Herbage glabrous, glandular-punctate. Leaf blades linear-oblanceolate to oblong, 1–10 cm long; dentate with amber-colored, resin-tipped teeth. Involucres 5–12 mm high. Rays 24 to 40; ligules 5–12 mm long. Disk corollas 4–6 mm long. Achenes 1–4 mm long, top margin smooth; pappus awns 2 or 3 (
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX).
Species Range
	
		
			Montana Range
		Range Descriptions
			 
			
			
					
						 Native
						Native
					 
				
				
					
			 
			
		 
		
	 
	
    		Range Comments
			Throughout North America. Thought to be native to the Great Plains and Rocky Mountains and adventive elsewhere (Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX).
			
		
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 1607
		
		
(Click on the following maps and charts to see full sized version)
		Map Help and Descriptions
		
		
			
				Relative Density
				 
			 
		
			
				Recency
				 
			 
		
		 
		
			
				 
			
				 
			
					 (Observations spanning multiple months or years are excluded from time charts)
				
			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Habitat
		Margins of wetlands, streambanks, roadsides and other disturbed, vernally moist habitats; plains, valleys (
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX).
Ecology
		POLLINATORS The following animal species have been reported as pollinators of this plant species or its genus where their geographic ranges overlap: 
Bombus bifarius, 
Bombus fervidus, 
Bombus flavifrons, 
Bombus rufocinctus, 
Bombus sylvicola, and 
Bombus occidentalis (Thorp et al. 1983, Williams et al. 2014).
 
		
	
		Stewardship Responsibility
		
		
	
	References
	
		
			- Literature Cited AboveLegend:   View Online Publication View Online Publication Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
 Thorp, R.W., D.S. Horning, and L.L. Dunning. 1983. Bumble bees and cuckoo bumble bees of California (Hymenoptera: Apidae). Bulletin of the California Insect Survey 23:1-79. Thorp, R.W., D.S. Horning, and L.L. Dunning. 1983. Bumble bees and cuckoo bumble bees of California (Hymenoptera: Apidae). Bulletin of the California Insect Survey 23:1-79.
 Williams, P., R. Thorp, L. Richardson, and S. Colla. 2014. Bumble Bees of North America. Princeton, NJ: Princeton University Press. 208 p. Williams, P., R. Thorp, L. Richardson, and S. Colla. 2014. Bumble Bees of North America. Princeton, NJ: Princeton University Press. 208 p.
 
- Additional ReferencesLegend:   View Online Publication View Online Publication
 Do you know of a citation we're missing? Adhikari, S. 2018. Impacts of dryland farming systems on biodiversity, plant-insect interactions, and ecosystem services. Ph.D. Dissertation. Bozeman, MT: Montana State University. 207 p. Adhikari, S. 2018. Impacts of dryland farming systems on biodiversity, plant-insect interactions, and ecosystem services. Ph.D. Dissertation. Bozeman, MT: Montana State University. 207 p.
 Anderson, N.L. 1951. Field studies on the biology of range grasshoppers of southeastern Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 96 p. Anderson, N.L. 1951. Field studies on the biology of range grasshoppers of southeastern Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 96 p.
 Bartoli, Adriana, and Roberto D. Tortosa. 2012. "Revision of the North American Species of Grindelia (Asteraceae) 1". Annals of the Missouri Botanical Garden. 98 (4): 447-513. Bartoli, Adriana, and Roberto D. Tortosa. 2012. "Revision of the North American Species of Grindelia (Asteraceae) 1". Annals of the Missouri Botanical Garden. 98 (4): 447-513.
 Boggs, K. W. 1984. Succession in riparian communities of the lower Yellowstone River, Montana. M.S. Thesis. Montana State University, Bozeman, 107 pp. Boggs, K. W. 1984. Succession in riparian communities of the lower Yellowstone River, Montana. M.S. Thesis. Montana State University, Bozeman, 107 pp.
 Britton, M. P. 1955. An ecological study of a relict grassland and an adjacent grazed pasture in Beaverhead Valley, Montana. M.S. thesis. Montana State University, Bozeman. 23 pp. Britton, M. P. 1955. An ecological study of a relict grassland and an adjacent grazed pasture in Beaverhead Valley, Montana. M.S. thesis. Montana State University, Bozeman. 23 pp.
 Chadde, S.W. 1985. Initial recovery patterns of southwestern Montana foothill range. M.Sc. Thesis. Bozeman, MT: Montana State University. 103 p. Chadde, S.W. 1985. Initial recovery patterns of southwestern Montana foothill range. M.Sc. Thesis. Bozeman, MT: Montana State University. 103 p.
 Cope, M.G. 1992. Distribution, habitat selection and survival of transplanted Columbian Sharp-tailed Grouse (Tympanuchus phasianellus columbianus) in the Tobacco Valley, Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 60 p. Cope, M.G. 1992. Distribution, habitat selection and survival of transplanted Columbian Sharp-tailed Grouse (Tympanuchus phasianellus columbianus) in the Tobacco Valley, Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 60 p.
 Corr, D.R. 1988. Effects of stress inducing factors on musk thistle (Carduus nutans L,) including--grass competition, Rhinocyllus conicus Froel., terminal flower loss, and insecticides. M.Sc. Thesis. Bozeman, MT: Montana State University. 86 p. Corr, D.R. 1988. Effects of stress inducing factors on musk thistle (Carduus nutans L,) including--grass competition, Rhinocyllus conicus Froel., terminal flower loss, and insecticides. M.Sc. Thesis. Bozeman, MT: Montana State University. 86 p.
 DuBois, K.L. 1979. An inventory of the avifauna in the Long Pines of Southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 113 p. DuBois, K.L. 1979. An inventory of the avifauna in the Long Pines of Southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 113 p.
 Eggers, M.J.S. 2005. Riparian vegetation of the Montana Yellowstone and cattle grazing impacts thereon. M.Sc. Thesis. Montana State University, Bozeman, MT. 125 p. Eggers, M.J.S. 2005. Riparian vegetation of the Montana Yellowstone and cattle grazing impacts thereon. M.Sc. Thesis. Montana State University, Bozeman, MT. 125 p.
 Fritzen, D.E. 1995. Ecology and behavior of Mule Deer on the Rosebud Coal Mine, Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 143 p. Fritzen, D.E. 1995. Ecology and behavior of Mule Deer on the Rosebud Coal Mine, Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 143 p.
 Gobeille, J.E. 1992. The effect of fire on Merriams turkey brood habitat in southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 61 p. Gobeille, J.E. 1992. The effect of fire on Merriams turkey brood habitat in southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 61 p.
 Guenther, G.E. 1989. Ecological relationships of bitterbrush communities on the Mount Haggin Wildlife Management Area. M.Sc. Thesis. Bozeman, MT: Montana State University. 73 p. Guenther, G.E. 1989. Ecological relationships of bitterbrush communities on the Mount Haggin Wildlife Management Area. M.Sc. Thesis. Bozeman, MT: Montana State University. 73 p.
 Harvey, S.J. 1990. Responses of steppe plants to gradients of water soil texture and disturbance in Montana, U.S.A. Ph.D. Thesis. Bozeman, MT: Montana State University. 34 p. Harvey, S.J. 1990. Responses of steppe plants to gradients of water soil texture and disturbance in Montana, U.S.A. Ph.D. Thesis. Bozeman, MT: Montana State University. 34 p.
 Jones, W. W. 1901. Preliminary flora of Gallatin County. M.S. Thesis. Bozeman, MT: Montana State College. 78 pp. Jones, W. W. 1901. Preliminary flora of Gallatin County. M.S. Thesis. Bozeman, MT: Montana State College. 78 pp.
 King, L.A. 1980. Effects of topsoiling and other reclamation practices on nonseeded species establishment on surface mined land at Colstrip, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 129 p. King, L.A. 1980. Effects of topsoiling and other reclamation practices on nonseeded species establishment on surface mined land at Colstrip, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 129 p.
 Kisch, H.R. 2015. An inventory of carbon stocks under native vegetation and farm fields in south-central Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 67 p. Kisch, H.R. 2015. An inventory of carbon stocks under native vegetation and farm fields in south-central Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 67 p.
 Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p.
 Meier, G.A. 1997. The colonization of Montana roadsides by native and exotic plants. M.Sc. Thesis. Bozeman, MT: Montana State University. 45 p. Meier, G.A. 1997. The colonization of Montana roadsides by native and exotic plants. M.Sc. Thesis. Bozeman, MT: Montana State University. 45 p.
 Mussgnug, G.L. 1972. The structure and performance of an adult population of Aulocara elliotti (Thomas) (Orthoptera, Acrididae) near Billings, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 97 p. Mussgnug, G.L. 1972. The structure and performance of an adult population of Aulocara elliotti (Thomas) (Orthoptera, Acrididae) near Billings, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 97 p.
 Nielsen, L.S. 1978. The effects of rest-rotation grazing on the distribution of Sharp-tailed Grouse. M.Sc. Thesis. Bozeman, MT: Montana State University. 52 p. Nielsen, L.S. 1978. The effects of rest-rotation grazing on the distribution of Sharp-tailed Grouse. M.Sc. Thesis. Bozeman, MT: Montana State University. 52 p.
 Peterson, J.G. 1969. The food habits and summer distribution of juvenile sage grouse in central Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 39 p. Peterson, J.G. 1969. The food habits and summer distribution of juvenile sage grouse in central Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 39 p.
 Quire, R.L. 2013. The sagebrush steppe of Montana and southeastern Idaho shows evidence of high native plant diversity, stability, and resistance to the detrimental effects of nonnative plant species. M.Sc. Thesis. Bozeman, MT: Montana State University. 124 p. Quire, R.L. 2013. The sagebrush steppe of Montana and southeastern Idaho shows evidence of high native plant diversity, stability, and resistance to the detrimental effects of nonnative plant species. M.Sc. Thesis. Bozeman, MT: Montana State University. 124 p.
 Rundquist, V.M. 1973. Avian ecology on stock ponds in two vegetational types in north-central Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 112 p. Rundquist, V.M. 1973. Avian ecology on stock ponds in two vegetational types in north-central Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 112 p.
 Sater, S. 2022. The insects of Sevenmile Creek, a pictorial guide to their diversity and ecology. Undergraduate Thesis. Helena, MT: Carroll College. 242 p. Sater, S. 2022. The insects of Sevenmile Creek, a pictorial guide to their diversity and ecology. Undergraduate Thesis. Helena, MT: Carroll College. 242 p.
 Seipel, T.F. 2006. Plant species diversity in the sagebrush steppe of Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 87 p. Seipel, T.F. 2006. Plant species diversity in the sagebrush steppe of Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 87 p.
 Skilbred, Chester L. 1979. Plant succession on five naturally revegetated strip-mined deposits at Colstrip, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 128 pp. Skilbred, Chester L. 1979. Plant succession on five naturally revegetated strip-mined deposits at Colstrip, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 128 pp.
 Skinner, K.F. 1995. Plant and grasshopper community composition: indicators & interactions across three spatial scales. M.Sc. Thesis. Bozeman, MT: Montana State University. 144 p. Skinner, K.F. 1995. Plant and grasshopper community composition: indicators & interactions across three spatial scales. M.Sc. Thesis. Bozeman, MT: Montana State University. 144 p.
 Thompson, W. L. 1993. Ecology of Merriam's Turkeys in relation to burned and logged areas in southeastern Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 195 p. Thompson, W. L. 1993. Ecology of Merriam's Turkeys in relation to burned and logged areas in southeastern Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 195 p.
 Tschache, O.P. 1970. Effects of ecological changes induced by various sagebrush control techniques on small mammal populations. M.Sc. Thesis. Bozeman, MT: Montana State University. 51 p. Tschache, O.P. 1970. Effects of ecological changes induced by various sagebrush control techniques on small mammal populations. M.Sc. Thesis. Bozeman, MT: Montana State University. 51 p.
 Tuinstra, K. E. 1967. Vegetation of the floodplains and first terraces of Rock Creek near Red Lodge, Montana. Ph.D dissertation. Montana State University, Bozeman 110 pp. Tuinstra, K. E. 1967. Vegetation of the floodplains and first terraces of Rock Creek near Red Lodge, Montana. Ph.D dissertation. Montana State University, Bozeman 110 pp.
 Wiman, N.G. 2001. Dynamics of leafy spurge (Euphorbia esula L.) infested plant communities influenced by flea beetles in the Aphthona complex (Colepotera: Chrysomelidae). M.Sc. Thesis. Bozeman, MT: Montana State University. 148 p. Wiman, N.G. 2001. Dynamics of leafy spurge (Euphorbia esula L.) infested plant communities influenced by flea beetles in the Aphthona complex (Colepotera: Chrysomelidae). M.Sc. Thesis. Bozeman, MT: Montana State University. 148 p.
 Wood, A.K. 1987. Ecology of a prairie mule deer population. Ph.D. Dissertation. Bozeman, MT: Montana State University. 205 p. Wood, A.K. 1987. Ecology of a prairie mule deer population. Ph.D. Dissertation. Bozeman, MT: Montana State University. 205 p.
 Zapatka, T.P. 1963. Some results of two limited hunting seasons on hen Pheasants in north central Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 26 p. Zapatka, T.P. 1963. Some results of two limited hunting seasons on hen Pheasants in north central Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 26 p.
 
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