Northern Goshawk - Accipiter gentilis
The Northern Goshawk is a fairly large hawk with a long tail having a broad, dark sub-terminal band and three to four narrower dark bands, rounded wing tips, and a conspicuous pale eyebrow. The sexes are similar with adults having a dark crown, blue-gray back, white underparts with fine, dense gray barring and conspicuous white undertail coverts. The eyes of adults are deep ruby-red and the feet are yellow. Immature Northern Goshawks are brown above, buffy below, with dense, blurry streaking. The undertail coverts are dark-streaked and the tail has wavy dark bands bordered with white and a thin white tip. The eyes of immature Northern Goshawks are yellowish, deepening in color to red as they mature. The total length is 53 to 66 cm, with females averaging larger than males (Squires and Reynolds 1997).
For a comprehensive review of the conservation status, habitat use, and ecology of this and other Montana bird species, please see Marks et al. 2016, Birds of Montana.
The Northern Goshawk is the largest and heaviest bodied of the three North American accipiters. Goshawks are clearly larger than Sharp-shinned Hawks (Accipiter striatus). Male Northern Goshawks can be of similar size to female Cooper's Hawks (Accipiter cooperi), but Northern Goshawks have broader wings and a relatively short tail compared to Cooper's Hawks (Squires and Reynolds 1997). Their ventral surface is pale rather than rust-colored as well. They can be distinguished from both Sharp-shinned and Cooper's Hawks by their whiteish underside as well as a boldly patterned head with a strong, white superciliary line above the eye (Sibley 2000). Juvenile Northern Goshawks can be distinguished from juvenile Cooper's Hawks by their conspicuous pale superciliary line (Squires and Reynolds 1997), overall buffy wash appearance on the breast and belly (Sibley 2000), uneven tail-bands creating a zig-zag pattern when the tail is spread (Squires and Reynolds 1997) and overall paler and more patterned upperside (Sibley 2000). Northern Goshawks can be discerned from falcons by their shorter, more rounded wings, and alternating flap-and-glide flight pattern.
Western Hemisphere Range
The Northern Goshawk has the widest distribution of the world’s 50 species of accipiters. It is a permanent resident in North America from Alaska east to Labrador and Newfoundland and south throughout much of the western U.S., along the northern tier of states from Minnesota to New York, in northern New Jersey and New England, and in the Appalachian Mountains of Maryland and West Virginia. It also winters in the Great Plains and across the central U.S. Disjunct populations occur in Mexico from Sonora and Chihuahua south to Jalisco and in Guerrero. Northern Goshawks also breed across Eurasia between 36º and 65ºN.
Summary of Observations Submitted for Montana
Number of Observations: 2585
(Click on the following maps and charts to see full sized version)
Map Help and Descriptions
(direct evidence "B")
(indirect evidence "b")
No evidence of Breeding
(regular observations "W")
(at least one obs. "w")
(Observations spanning multiple months or years are excluded from time charts)
Earliest, peak and latest migration dates for Northern Goshawks in Montana are not well established for either spring or fall, and spring movements are less obvious than fall migration. Northern Goshawks are much less common at migration watch sites than are the other two accipiters, and their numbers vary more from year to year (Mueller et al. 1977). The number counted annually from late August through late October at the Bridger Mountains watch site ranged from two to 96 between 1991 and 2010 (mean = 32). West of the divide at the Jewel Basin watch site, 50 were counted from 31 August to 1 November 2008, 30 from 30 August to 25 October 2009, and 35 from 28 August to 23 October 2010 (Dan Casey, personal communication). The highest numbers occurred after the middle of October in all three years, and the lower counts in 2009 and 2010 may have resulted from the count periods ending sooner owing to heavy snows (Dan Casey, personal communication). The species is generally considered a year-round resident or partial migrant in Montana as Northern Goshawks have been observed in transit during every month of the year (Montana Bird Distribution Committee 2012). In the northeastern quarter of Montana it is an uncommon migrant and winter resident only. Movements are apparently dependent on prey availability and is often only to lower elevations or into more open habitat types (Squires and Reynolds 1997), which may explain the presence of Northern Goshawks wintering at locations within the state east of documented breeding locations. Migration routes throughout the west are poorly defined and are believed to occur over a broad area (Squires and Reynolds 1997). Beebe (1974) believed significant numbers from Alberta, as well as Montana breeding birds, might migrate south along the Rocky Mountain Front Range. Irruptive movements of northern birds to the south occur in approximately 10-year cycles and are apparently dependent on Snowshoe Hare and grouse population lows (Squires and Reynolds 1997).
Goshawks nest in a variety of forest types in Montana, including Douglas-fir and Western Larch west of the Continental Divide, Lodgepole Pine in Beaverhead County, and Ponderosa Pine in Powder River and Carter counties. They prefer mature and old-growth forests with a preponderance of large trees, a dense canopy, and a relatively open understory (Hayward and Escano 1989, Squires and Reynolds 1997, Clough 2000). An exception to this generality is in Beaverhead County, where nests commonly occur in Lodgepole Pine stands with an average tree diameter of only 13 cm, although the birds usually place their nests in larger trees within these stands (Kirkley 1996). The nest is a bulky platform of sticks placed near the main trunk of a large tree from 6-20 m off the ground, usually in the lower part of the canopy. Forest stands where Northern Goshawks nest in Montana tend to be mature large-tract conifer forests with a high canopy cover (69%), relatively steep slope (21%), and little to sparse undergrowth (Kirkley 1996). Hayward and Escano (1989) examined nest-site characteristics at 17 territories in western Montana and northern Idaho in 1983. The birds nested preferentially in mature and old-growth stands of conifers that had a closed canopy (75-85% canopy cover) and a large forest opening within 1 km of the nest. Nest heights ranged from 7-17 m, and most nests were placed next to the main trunk in the lower one-third of the canopy. All Northern Goshawk nest trees reported by Kirkley (1996) were either Lodgepole Pine or Douglas-fir with an average DBH (diameter at breast height) of 33.6 cm and average height of 21.9 meters. In another study conducted in Montana, Douglas-fir, Ponderosa Pine and Grand Fir were the trees selected most often for nest building (State of Idaho HCA/CS Dev. Team 1995). Nests were constructed an average 10.9 meters above the ground and were usually located near water (232 m) or a clearing (85 m) (Kirkley 1996). Range-wide nest site characteristics are similar. Almost no information is available regarding Northern Goshawk foraging strategies in Montana. It is known they hunt in closed canopy habitats as well as more open landscapes and over 50 species of identified prey indicate they are generalists in terms of prey selection. Little information exists concerning Northern Goshawk non-breeding or wintering habitat in the state. However, in the Bozeman area, birds coming into the valley are found in forested or thickety areas. Multiple observations of wintering Northern Goshawks are documented in the north-central and northeastern areas of Montana (Montana Bird Distribution Committee 2012), possibly indicating movement toward areas of higher prey availability.
Ecological Systems Associated with this Species
- Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
- Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
- Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
- Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
- Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if
point observations were associated with that system.
Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.
If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: http://mtnhp.org/requests/default.asp
) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.
- Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
- Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
- Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
- Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
- Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
- Maxell, B.A. 2000. Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
- Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
- Commonly Associated with these Ecological Systems
Forest and Woodland Systems
Recently Disturbed or Modified
- Occasionally Associated with these Ecological Systems
Forest and Woodland Systems
Recently Disturbed or Modified
Wetland and Riparian Systems
Goshawks are generalists when it comes to foraging, hunting beneath the forest canopy in dense and open stands and at forest-grassland and forest-shrubland ecotones, where they take a wide variety of prey that includes forest grouse, woodpeckers, corvids, lagomorphs, and squirrels. During winter in Montana, many birds move to grasslands, shrublands, and valley-bottom riparian areas, where they hunt Gray Partridges, Ring-necked Pheasants, Sharp-tailed Grouse, and other prey that frequent open country (J. Marks, pers. obs.). Parratt (1964) observed a nest in Glacier NP in 1956 that was placed in a spruce about 18 m off the ground and contained four young that fledged on 12 August; small mammals (especially chipmunks) were the main prey items. Mammals made up the bulk of the diet of 19 pairs in the Flint Creek Mountains in 1997 and 1998 (Clough 2000), with the major prey items being Snowshoe Hares, Columbian Ground Squirrels, and Red Squirrels. Avian prey included Ruffed Grouse, Dusky Grouse, Northern Flickers, and Gray Jays. Generally, Northern Goshawks forage during short flights alternating with brief prey searches from perches. Elsewhere across their range, dominant mammalian prey includes five species of tree squirrels, four ground squirrels, and lagomorphs. Frequently killed birds include three galliformes, four corvids, six woodpeckers (Piciformes) and the American Robin (Turdus migratorius) (Squires and Reynolds 1997). During the nesting season, the diet can vary with prey availability. For example, as more fledgling passerines become available, they make up a greater portion of the diet (Linden and Wikman 1983, Reynolds and Meslow 1984). The ratio of mammalian prey to avian prey in the diet during the breeding season (in percent) in Arizona was 76:24 and 62:38 (Boal and Mannan 1994, Reynolds et al. 1994). This ratio in Nevada was 67:32 (Younk and Bechard 1994) in New York was 39:61 (Grzybowski and Eaton 1976) and Oregon was 42:59 and 45:55 (Bull and Hohmann 1994, Reynolds and Meslow 1984). Non-breeding season food habits are unknown for North American populations. In Sweden, birds dominate the diet during the nesting season (86 percent of prey), whereas in winter, Red Squirrels (Sciurus vulgaris) comprise the bulk of the diet (79 percent) (Widen 1987).
Northern Goshawks are considered to be decreasing in numbers near Fortine (Weydemeyer 1975). All other ecological information regarding this species, from studies in other areas of the species' range, indicate nesting densities of most western U.S. populations range from 6.6 to 10.7 pairs per 100 square kilometers (Squires and Reynolds 1997). The single nesting density estimate for the eastern U.S. is 1.17 pairs per 100 square kilometers (Kimmel and Yahner 1994). Home ranges during nesting vary from 95 to 3500 hectares depending on sex and habitat characteristics (Squires and Reynolds 1997). Home ranges of males are typically larger than those of females (Hargis et al. 1994, Keane and Morrison 1994, Kennedy et al. 1994). Exclusive of nesting areas, home ranges of adjacent pairs are not defended and may overlap (Squires and Reynolds 1997). The core area encompasses the nest site and constitutes 32 percent of the home range (Kennedy et al. 1994). Individuals typically enlarge or sometimes shift location of home ranges after breeding (Hargis et al. 1994, Keane and Morrison 1994). Home ranges of non-breeders are poorly known, but may be larger than those of breeders (Squires and Reynolds 1997). In North America, winter home ranges are unknown. In Sweden, winter home ranges of males and females were similar and averaged 5700 hectares (Widen 1989). In California, 76.5 percent of males and 71.4 percent of females returned to the same nesting area in subsequent years. Males were significantly more likely to return to previously inhabited territories in consecutive years than females (Detrich and Woodbridge 1994). In Arizona, 80 percent of nest areas examined in two consecutive years were re-used the second year by one or both members of the pair banded the first year (Reynolds et al. 1994). Sixty to 72 percent of adults located in consecutive years retained the mate from the previous year (Detrich and Woodbridge 1994, Reynolds et al. 1994). Dispersal of young is not well documented. Detrich and Woodbridge (1994) recaptured two adult females, banded as nestlings 5 to 7 years prior, 16 and 24 kilometers from their natal sites. Three females, banded as nestlings and recaptured as breeding adults, moved an average of 21.5 kilometers from their natal sites, and another female, captured as a breeding adult seven years after being banded as a nestling, moved 100 kilometers from its natal site (Squires and Reynolds 1997). Little is known regarding survivorship in the U.S. In Arizona, annual survivorship of male and females more than 1 year old was estimated to be 68.8 percent and 86.6 percent, respectively (Squires and Reynolds 1997). In Yukon, Canada, an observed population decline was attributed to increased mortality of eggs, nestlings, immatures and adults, as well as to dispersal following a precipitous decline in number of Snowshoe Hares (Doyle and Smith 1994). The maximum lifespan of a wild bird is 11 years (Squires and Reynolds 1997). The sex ratio is 1:1 prior to fledging and among adults (Mueller and Berger 1968, Reynolds et al. 1994). The BBS is inadequate for assessing population trends, as are counts at migration watch sites (Boal et al. 2005).
Information on Northern Goshawk reproduction in Montana is limited. Eggs are laid from early April to late May, young fledge from late June to mid-August and are independent of parental care by early September. Only one brood is raised per year. Most Northern Goshawk studies have focused on habitat and nest-site characteristics. Very few studies provide information on aspects of nesting biology or reproductive effort/success. The Montana Natural Heritage Program's Point Observation Database indicates most documented Northern Goshawk nests had 1 to 3 chicks/juveniles. This information must be used with caution because in many instances these were just observations of nests with no methodological survey techniques applied. Kirkley (1996) did have a total of 13 young fledge from five active nests and Pilliod (1994) had two nestlings but only one fledgling from a single nest. Egg dates are probably similar to those reported for Wyoming: May 10 to June 17 (Johnsgard 1986). Young have been observed in June and August (Davis 1961). Information from other portions of this species' range indicate that usually one clutch is produced per year, occurring from late April through early May (Squire and Reynolds 1997). However, some individuals may not breed during cold, wet springs (DeStefano et al. 1994).
Egg-laying may begin later at higher elevations and during cold, wet springs (Henny et al. 1985, Younk and Bechard 1994). The average clutch is typically two to four eggs and rarely one to five (Squires and Reynolds 1997). The average clutch size of 44 North American clutches is 2.7 eggs (Apfelbaum and Seelbach 1983). Eggs are laid every two to three days and incubation usually begins after the second egg is laid. Incubation, conducted principally by the female, takes 28 to 38 days and hatching is asynchronous. Principally the female performs brooding and feeding of nestlings. The male brings food to the nest. The young begin flying at 35 to 42 days and become independent at about 70 days (Boal 1994, Squires and Reynolds 1997). Northern Goshawks maintain one to eight alternate nests within a nest area (Squire and Reynolds 1997). Alternate nests range from 15 to 2066 meters apart (Reynolds and Wight 1978, Woodbridge and Detrich 1994). Nesting by subadults is more frequent in expanding populations and less frequent in stable populations (Reynolds and Wight 1978).
The Northern Goshawk is currently listed as a sensitive species in all National Forests and on all BLM lands in Montana. It is also a Forest Plan Management Indicator Species in the Beaverhead-Deerlodge National Forest. Northern Goshawks are a Species of Management Concern in U.S. Fish and Wildlife Service Region 6 (USFWS 1995). However, Maj (1996) reports Northern Goshawk populations in Region 1 are increasing or stable in most forests. The USFWS was petitioned in 1997 to list goshawks under the Endangered Species Act but determined that such listing was not warranted. An independent committee that was formed to review the status of goshawks in the western U.S. concluded that existing data were not adequate for assessing population trends, and that assessing the status of goshawks solely using distribution of late-successional forests is not appropriate (Boal et al. 2005). The latter conclusion resulted from the fact that goshawk habitat use is not restricted to mature and old-growth forest, even though goshawks nest selectively in these habitats in many areas. On balance, more information is needed on population trends and habitat relations, especially with regard to how forestry practices influence these issues.
Threats or Limiting Factors
Although shooting is not considered an important mortality factor (Squires and Reynolds 1997), the problem has not disappeared entirely given that a radio-tagged goshawk was shot in Montana as recently as Jan 2011 (J. Kirkley, personal communication). Timber harvest is the main threat to nesting birds because of their preference for large trees and forest stands with high canopy cover (Squires and Reynolds 1997), although loss of forest stands to fire and bark beetles could degrade goshawk habitat. The effects of timber harvest on large-scale population trends remain unknown (Squires and Reynolds 1997, Boal et al. 2005).
- Literature Cited AboveLegend: View Online Publication
- Apfelbaum, S. I. and P. Seelbach. 1983. Nest tree, habitat selection and productivity of seven North American raptor species based on the Cornell University Nest Record Card Program. Raptor Research 17:97-113.
- Beebe, F. L. 1974. Goshawk. Pp. 54-62 In: Field studies of the Falconiformes of British Columbia. British Columbia Provincial Museum Occasional Papers. Series No. 17.
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- Boal, C.W., Andersen, D.E. and P.L. Kennedy. 2005. Foraging and nesting habitat of male northern goshawks in the Laurentian mixed forest province, Minnesota. Journal Of Wildlife Management 69(4): 1516-1527.
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- Clough, L. T. 2000. Nesting habitat selection and productivity of Northern Goshawks in west-central Montana. M.S. thesis, University of Montana, Missoula.
- Davis, C.V. 1961. A distributional study of the birds of Montana. Ph.D. dissertation. Oregon State University, Corvallis. 462 pp.
- DeStefano, S., S. K. Daw, S. M. Desimone, and E. C. Meslow. 1994. Density and productivity of northern goshawks: implications for monitoring and management. Studies in Avian Biology 16:88-91.
- Detrich, P. J. and B. Woodbridge. 1994. Territory fidelity, mate fidelity, and movements of color-marked northern goshawks in the southern Cascades of California. Studies in Avian Biology 16:130-132.
- Doyle, F. I. and J. M. N. Smith. 1994. Population responses of northern goshawks to the 10-year cycle in numbers of snowshoe hares. Studies in Avian Biology 16:122-129.
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- Hayward, G. D. and R. E. Escano. 1989. Goshawk nest-site characteristics in western Montana and northern Idaho. Condor 91:476-479.
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- Johnsgard, P.A. 1986. Birds of the Rocky Mountains: with particular reference to national parks in the northern Rocky Mountain region. Colorado Associated University Press, Boulder, CO.
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- Kimmel, J. T. and R. H. Yahner. 1994. The northern goshawk in Pennsylvania: habitat use, survey protocols, and status. Final Report. School of Forest Resources, Pennsylvania State University, University Park.
- Kirkley, Jack. 1996. Northern goshawk monitoring, Beaverhead, Deerlodge National Forest, Dillon District, Wisdom District and Wise River District. Summer 1996.
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- Marks, J.S., P. Hendricks, and D. Casey. 2016. Birds of Montana. Arrington, VA. Buteo Books. 659 pages.
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- Parratt, L. P. 1964. Birds of Glacier National Park. Special Bulletin No. 9, Glacier Natural History Association, West Glacier, Montana.
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- State of Idaho Habitat Conservation Assessment and Conservation Strategy (HCA/CS) Development Team. 1995. Habitat conservation assessment and strategy for the northern goshawk (Accipiter gentilis). Unpublished report. Idaho State Conservation Effort (Idaho Department of Fish and Game). 35 pp.
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- Additional ReferencesLegend: View Online Publication
Do you know of a citation we're missing?
- Adkisson, C. S. 1990. Accipiters. Pages 63-9 in B. G. Pendleton (editors). Proceedings of the Southeast Raptor Management Symposium and Workshop. Nat. Wildl. Fed. Sci. and Tech. Ser. No. 14.
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- Anonymous. 1993. Northern goshawk management workshop. Proceedings of March 1993 workshop, Madison, Wisconsin. 60 pp.
- Bailey, Vernon and Florence Merriam Bailey. 1918. Wild animals of Glacier National Park: the mammals, with notes on physiography and life zones and the birds. U.S. Department of the Interior, National Park Service. 210 pp.
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- Becker, Dale M., 1980, A Survey of raptors on national forest land in Carter County, Montana. Final Progress Report: 1977-1979.
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- Boal, C. W., and J. E. Bacorn. 1994. Siblicide and cannibalism in northern goshawk nests. Auk 111:748-750.
- Boal, C. W., and R. W. Mannan. 1996. Prey sizes of male and female northern goshawks. Southwestern Naturalist 41:355-358.
- Bosakowski, T., and R. Speiser. 1994. Macrohabitat selection by nesting northern goshawks: implications for managing eastern forests. Studies in Avian Biology 16:46-49
- Bright-Smith, D. J., and R. W. Mannan. 1994. Habitat use by breeding male northern goshawks in northern Arizona. Studies in Avian Biology 16:58-65.
- Cameron, E. S. 1907. The birds of Custer and Dawson counties, Montana. Auk 24(3): 241-270, 389-406.
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- Casey, D. 2000. Partners in Flight Draft Bird Conservation Plan Montana. Version 1.0. 287 pp.
- Casey, Dan. 2008. Personal comunication to the Montana Natural Heritage Program.
- Crocker-Bedford, D. C. 1990. Goshawk reproduction and forest management. Wildl. Soc. Bull., 18:262-269.
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- Dickson, D.C. 1991. Systematic wildlife observations on the Blackfoot-Clearwater Wildlife Management Area. Montana Department of Fish, Wildlife and Parks. Missoula, MT. 14 pp. plus appendices and photographs.
- Dobkin, D. S. 1992. Neotropical migrant landbirds in the Northern Rockies and Great Plains. U.S.D.A. For. Serv. N. Region Publ. R1-93-34. Missoula, Mont.
- Doyle, F.I. 1995. Bald eagle (Haliaeetus leucocephalus) and northern goshawk (Accipiter gentilis) nests apparently preyed upon by a wolverine(s), (Gulo gulo) in the southwestern Yukon Territory. Canadian Field-Naturalist 109:115-116.
- ECON, Inc. (Ecological Consulting Service), Helena, MT., 1975, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1975. Proj. 71-23-A. December 31, 1975.
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