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Terrestrial Gartersnake - Thamnophis elegans

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Native Species

Global Rank: G5
State Rank: S5
(see State Rank Reason below)


Agency Status
USFWS:
USFS:
BLM:


 

External Links

State Rank Reason (see State Rank above)
Species is relatively common within suitable habitat and widely distributed across portions of the state
  • Details on Status Ranking and Review
    Terrestrial Gartersnake (Thamnophis elegans) Conservation Status Review
    Review Date = 05/03/2018
    Range Extent

    ScoreG - 200,000-2,500,000 km squared (about 80,000-1,000,000 square miles)

    Comment304,213 square Kilometers from Natural Heritage Program range maps

    Long-term Trend

    ScoreE - Relatively Stable (±25% change)

    CommentHabitat is likely stable within +/- 25% since European settlement, the increasing in stock ponds may have increased the amount of habitat for this species

    Short-term Trend

    ScoreE - Stable. Population, range, area occupied, and/or number or condition of occurrences unchanged or remaining within ±10% fluctuation

    CommentSpecies is regularly found during surveys of lentic sites within its range

    Threats

    ScoreH - Unthreatened. Threats if any, when considered in comparison with natural fluctuation and change, are minimal or very localized, not leading to significant loss or degradation of populations or area even over a few decades’ time. (Severity, scope, and/or immediacy of threat considered Insignificant.)

    CommentNo operational threats in the next 15-20 years identified

    Intrinsic Vulnerability

    ScoreB - Moderately Vulnerable. Species exhibits moderate age of maturity, frequency of reproduction, and/or fecundity such that populations generally tend to recover from decreases in abundance over a period of several years (on the order of 5-20 years or 2-5 generations); or species has moderate dispersal capability such that extirpated populations generally become reestablished through natural recolonization (unaided by humans).

    CommentModerately Vulnerable. Species exhibits moderate age of maturity, frequency of reproduction, and/or fecundity such that populations generally tend to recover from decreases in abundance within 5-20 years or 2-5 generations. Species has good dispersal ca

    Environmental Specificity

    ScoreC - Moderate. Generalist. Broad-scale or diverse (general) habitat(s) or other abiotic and/or biotic factors are used or required by the species but some key requirements are scarce in the generalized range of the species within the area of interest.

    CommentFound across a wide range of habitats

    Raw Conservation Status Score

    Score 3.5 + 0 (geographic distribution) + 0 (environmental specificity) + 0 (short-term trend) + 1 (threats) = 4.5

 
General Description
Adults vary from 16 to 43 inches in length. This snake is distinguished by three yellow stripes (one dorsal, two lateral) running the length of the body and a series of black spots situated between, and somewhat on, the stripes. The background color between the stripes is brownish or greenish. All-black individuals are occasionally found. A series of dark black/brown blotches covers most of the belly. There are normally eight scales on the upper lip.

Species Range
Montana Range Range Descriptions

Native

Western Hemisphere Range

 


Observations in Montana Natural Heritage Program Database
Number of Observations: 3523

(Click on the following maps and charts to see full sized version) Map Help and Descriptions
Relative Density

Recency

 

(Observations spanning multiple months or years are excluded from time charts)



Migration
Non-migratory.

Habitat
Terrestrial Gartersnakes are found in nearly all habitats, but most commonly at lower elevations around water. Females give birth to 6 to 18 live young during summer. They eat a variety of vertebrates and invertebrates. Common near water but also found away from water. At high elevations common on rocky cliffs and brushy talus (Brunson and Demaree 1951 and Franz 1971). On prairie may be more common along brushy bottomland (Mosimann and Rabb 1952 and Lewin 1963).

Ecological Systems Associated with this Species
  • Details on Creation and Suggested Uses and Limitations
    How Associations Were Made
    We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
    1. Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
    2. Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
    3. Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
    4. Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
    Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.  In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.  However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if point observations were associated with that system.  Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.  The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.  If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.

    Suggested Uses and Limitations
    Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.  These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: mtnhp.org/requests) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.  Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.  Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.  Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).  Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.

    Literature Cited
    • Adams, R.A.  2003.  Bats of the Rocky Mountain West; natural history, ecology, and conservation.  Boulder, CO: University Press of Colorado.  289 p.
    • Dobkin, D. S.  1992.  Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34.  Missoula, MT.
    • Foresman, K.R.  2012.  Mammals of Montana.  Second edition.  Mountain Press Publishing, Missoula, Montana.  429 pp.
    • Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998.  Montana atlas of terrestrial vertebrates.  Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT.  1302 p.
    • Hutto, R.L. and J.S. Young.  1999.  Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32.  72 p.
    • Maxell, B.A.  2000.  Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species.  Report to U.S. Forest Service Region 1.  Missoula, MT: Wildlife Biology Program, University of Montana.  161 p.
    • Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath.  2004.  Amphibians and reptiles of Montana.  Missoula, MT: Mountain Press Publishing Company. 262 p.

Food Habits
Generalist predator in aquatic and terrestrial habitats (Anderson 1977). May specialize in amphibians during metamorphosis (Kephart and Arnold 1982, Gregory 1984c). Diet in western MT: leech 44%, slug 24%, fish 10%, amphibian 10%, miscellaneous 12% (Anderson 1977).

Reproductive Characteristics
Birth usually occurs in August to September (Colorado) (Hammerson 1982), late July to early August (Pacific Northwest) (Nussbaum et al. 1983).


References
  • Literature Cited AboveLegend:   View Online Publication
    • Anderson, M.E. 1977. Aspects of the ecology of two sympatric species of Thamnophis and heavy metal accumulation with the species. M.S. thesis, University of Montana, Missoula. 147 pp.
    • Brunson, R.B. and H.A. Demaree, Jr. 1951. The herpetology of the Mission Mountains, Montana. Copeia (4):306-308.
    • Franz, R. 1971. Notes on the distribution and ecology of the herpetofauna of northwestern Montana. Bulletin of the Maryland Herpetological Society 7: 1-10.
    • Gregory, P.T. 1984c. Habitat, diet and composition of assemblages of garter snakes (Thamnophis) at eight sites on Vancouver Island (Canada). Canadian Journal of Zoology 62(10): 2013-2022.
    • Hammerson, G.A. 1982. Amphibians and reptiles in Colorado. Colorado Division of Wildlife. Denver, CO. 131 pp.
    • Kephart, D.G. and S.J. Arnold. 1982. Garter snake diets in a fluctuating environment: a seven-year study. Ecology 63(5): 1232-1236.
    • Lewin, V. 1963. The herpetofauna of southeastern Alberta. Can. Field-Nat. 77:203-214.
    • Mosimann, J.E. and G.B. Rabb. 1952. The herpetology of Tiber Reservoir Area, Montana. Copeia(1): 23-27.
    • Nussbaum, R.A., E.D. Brodie, Jr. and R.M. Storm. 1983. Amphibians and reptiles of the Pacific Northwest. University of Idaho Press. Moscow, ID. 332 pp.
  • Additional ReferencesLegend:   View Online Publication
    Do you know of a citation we're missing?
    • [VTNWI] VTN Wyoming Incorporated. No Date. Second year's analysis of terrestrial wildlife on proposed mine access and railroad routes in southern Montana and northern Wyoming, March 1979 - February 1980. VTN Wyoming Incorporated. Sheridan, WY. 62 p.
    • [WWPC] Washington Water Power Company. 1995. 1994 wildlife report Noxon Rapids and Cabinet Gorge Reservoirs. Washington Water Power Company. Spokane, WA.
    • Alfaro, M.E. 2003. Sweeping and striking: a kinematic study of the trunk during caprture in three thamnophiine snakes. Journal of Experimental Biology 206(14): 2381-2392.
    • Arnold, S.J. 1977b. Polymorphism and geographic variation in the feeding behavior of the garter snake, T. elegans. Science 197: 676-678.
    • Arnold, S.J. 1980. Inheritance and evolution of reproductive traits in garter snakes. American Zoologist 20(4): 781.
    • Arnold, S.J. 1981a. Behavioral variation in natural populations I. Phenotypec, genetic, and environmental correlations between chemoreceptive responses to prey in the garter snake, T. elegans. Evolution 35: 489-509.
    • Arnold, S.J. 1981b. Behavioral variation in natural populations II. The inheritance of a feeding response in crosses between geographic races of the garter snake, T. elegans. Evolution 35:510-515.
    • Arnold, S.J. 1982. A quantitative approach to antipredator performance: salamander defense against snake attack. Copeia 1982(2): 247-253.
    • Arnold, S.J. 1988. Quantitative genetics and selection in natural populations: microevolution of vertebral numbers in the garter snake Thamnophis elegans. In: Weir, B.S., Eisen, E.J., Goodman, M. & Namkoong, G. [Eds]. Proceedings of the Second International Conference on Quantitative Genetics. Sinauer Associates, Inc., Sunderland, Massachusetts. 724 p.
    • Arnold, S.J. and C.R. Peterson. 2002. A model for optimal reaction norms: the case of the pregnant garter snake and her temperature-sensitive embryos. American Naturalist 160(3):306-316.
    • Arnold, S.J. and P.C. Phillips. 1999. Hierarchical comparison of genetic variance-covariance matrices. II. Coastal-inland divergence in the garter snake, Thamnophis elegans. Evolution 53: 1516-1527.
    • Arnold, S.J. and R.J. Wassersug. 1978. Differential predation on metamorphic anurans by garter snakes (Thamnophis): social behavior as a possible defense. Ecology 59(5): 1014-1022.
    • Arnold, S.J., C.R. Peterson, and J. Gladstone. 1995. Behavioural variation in natural populations. VII. Maternal body temperature does not affect juvenile thermoregulation in a garter snake. Animal Behaviour 50(3): 623-633.
    • Arnold, S.J., K.C. Kelley, and J. Gladstone. 1993. Population variation in locomotor performance of juvenile garter snakes (Thamnophis elegans). American Zoologist 33(5): 25A.
    • Atkinson, E.C. and M.L. Atkinson. 2004. Amphibian and reptile survey of the Ashland and Sioux of the Custer National Forest with special emphasis on the Three-Mile Stewardship Area:2002. Marmot's Edge Conservation. 22 p.
    • Ayres, F.A. and S.J. Arnold. 1983. Behavioral variation in natural populations. IV. Mendelian models and heritability of a feeding response in the garter snake, T. elegans. Heredity 51: 405-413.
    • Baird, S.F. and C.F. Girard. 1853. Catalogue of North American Reptiles., Pt. 1, p. 34-35.
    • Barker, B.S., S.J. Arnold, and M.E. Pfrender. 2003. Comparisons of inheritance matrices in three populations of the garter snake (Thamnophis elegans). Abstract. Northwestern Naturalist 84:94.
    • Beal, M.D. 1951. The occurrence and seasonal activity of vertebrates in the Norris and Gibbon Geyser Basins of Yellowstone National Park. M.S. Thesis. Utah State Agricultural College. Logan, Utah. 61 pp.
    • Bellemin, J.M. and G.R. Stewart. 1977. Diagnostic characters and color convergence of the garter snakes (Thamnophis elegans terrestris) and (Thamnophis couchii atratus) along the central California coast. Bulletin of the Southern California Academy of Science 76(2): 73-84.
    • Benson, K.R. 1978. Herpetology of the Lewis and Clark expedition 1804-1806. Herpetological Review 9(3): 87-91.
    • Berner, N.J. 1988. Molecular basis of the difference in oxygen affinity between maternal and fetal blood in the viviparous garter snake, Thamnophis elegans. M.S. Thesis, University of Idaho. 40 p.
    • Berner, N.J., and R.L. Ingermann. 1988. Molecular basis of the difference in oxygen affinity between maternal and fetal red blood cells in the viviparous garter snake Thamnophis elegans. Journal of Experimental Biology 140(0): 437-454.
    • Boundy, J. 2001. Herpetofaunal surveys in the Clark Fork Valley region, Montana. Herpetological Natural History 8: 15-26.
    • Boundy, J. and D. A. Rossman. 1995. Allocation and status of the garter snake names Coluber infernalis Blainville, Eutaenia sirtalis tetrataenia Cope, and Eutaenia imperialis Coues and Yarrow. Copeia 1995:236-240.
    • Bourguignon, T. 2002. Garter snakes, origins, care and species. Stuttgart:Verlag Eugen Ulmer. 2002.
    • Bronikowski, A.M. 2000. Experimental evidence for the adaptive evolution of growth rate in the garter snake Thamnophis elegans. Evolution 54(5): 1760.
    • Bronikowski, A.M. and S.J. Arnold. 1999. The evolutionary ecology of life history variation in the garter snake Thamnophis elegans. Ecology 80(7): 2314-2325.
    • Bronikowski, A.M. and S.J. Arnold. 2001. Cytochrome b phylogeny does not match subspecific classification in the western terrestrial garter snake, Thamnophis elegans. Copeia (2) 508-513.
    • Brunson, R.B. 1955. Check list of the amphibians and reptiles of Montana. Proceedings of the Montana Academy of Sciences 15: 27-29.
    • Bull, E.L. and M.P. Hayes. 2000. Livestock effects on reproduction of the Columbia spotted frog. Journal of Range Management 53: 293-296.
    • Bureau of Indian Affairs, Department of the Interior. 1981. Draft Environmental Impact Statement. Unpublished report for the Crow/Shell Coal Lease, Crow Indian Reservation, Montana.
    • Burghardt, G.M. 1969. Comparative prey-attack studies in newborn snakes of the genus Thamnophis. Behaviour 33: 77-114.
    • Burroughs, R. D. 1961. Natural history of the Lewis and Clark expedition. Michigan State University Press, East Lansing. 340 p.
    • Butler, R. 1985. The captive care and breeding of Thamnophis elegans terrestris. Herptological Review 10(2): 34-38.
    • Butts, T.W. 1997. Mountain Inc. wildlife monitoring Bull Mountains Mine No. 1, 1996. Western Technology and Engineering. Helena, MT.
    • Campbell, R.W. 1969. Notes on some foods of the wandering garter snake on Mittlenatch Island, British Columbia. Syesis 2: 183-187.
    • Charland, M.B and P.T. Gregory. 1995. Movements and habitat use in gravid and nongravid female garter snakes (Colubridae: Thamnophis). Journal of Zoology (London) 236(4): 543-561.
    • Charland, M.B. 1991. Anesthesia and transmitter implantation effects on gravid garter snakes (Thamnophis sirtalis and Thamnophis elegans). Herpetological Review 22(2): 46-47.
    • Charland, M.B. 1993. Reproductive ecology of female garter snakes (Thamnophis) in southeastern British Columbia. Ph.D. Dissertation, University of Victoria (Canada) 226p. 1991.
    • Charland, M.B. 1995. Thermal consequences of reptilian viviparity: thermoregulation in gravid and nongravid garter snakes (Thamnophis). Journal of Herpetology 29(3): 383-390.
    • Cochran, P.A. 1986a. Opportunistic scavenging by a garter snake in Colorado. Bulletin of the Chicago Herpetological Society 21(3-4): 94-95.
    • Confluence Consulting Inc. 2010. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT.
    • Cooper, J. G. 1869. Notes on the fauna of the Upper Missouri. American Naturalist 3(6):294-299.
    • Cooper, J. G. 1869. The fauna of Montana Territory (concluded) III. Reptiles, IV Fish. American Naturalist 3(3):124-127
    • Cope, E.D. 1872. Report on the recent reptiles and fishes of the survey, collected by Campbell Carrington and C.M. Dawes. pp. 467-469 In: F.V. Hayden, Preliminary report of the United States geological survey of Montana and portions of adjacent territories; being a fifth annual report of progress. 538 pp. 42nd Congress, 2nd Session, House Executive Document Number 326. Serial 1520.
    • Cope, E.D. 1875. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. U.S. Natioanl Museum Bulletin 1: 1-104.
    • Cope, E.D. 1900. The crocodilians, lizards, and snakes of North America. Report of the U.S. National Museum 1898: 153-1270.
    • Corn, J. and P. Hendricks. 1998. Lee Metcalf National Wildlife Refuge bullfrog and painted turtle investigations: 1997. Montana Natural Heritage Program. Helena, MT. 20 pp.
    • Coues, E. and H. Yarrow. 1878. Notes on the herpetology of Dakota and Montana. Bulletin of the U.S. Geological Geographic Survey of the Territories 4: 259-291.
    • Crother, B.I. (ed.) 2008. Scientific and standard English names of amphibians and reptiles of North America north of Mexico. SSAR Herpetological Circular No. 37:1-84.
    • Cunningham, J.D. 1955. Notes on the ecology of Thamnophis e. elegans (Baird and Girard). Herpetologica 11: 152.
    • de Queiroz, A. and R. Lawson. 1994. Phylogenetic relationships of the garter snakes based on DNA sequence and allozyme variation. Biological Journal of the Linnean Society 53: 209-229.
    • Drummond, H. 1983. Aquatic foraging in garter snakes: a comparison of specialists and generalists. Behaviour 86: 1-30.
    • Drummond, H. and C.M. Garcia. 1989. Limitations of a generalist: a field comparison of foraging snakes. Behaviour 108(1-2): 23-43.
    • Drummond, H. and G.M. Burghardt. 1983. Geographic variation in the foraging behavior of the garter snake, Thamnophis elegans. Behaviour of Ecological Sociobiology 12(1): 43-48.
    • Drummond, H.M. 1980. Aquatic foraging in some new world natricine snakes: generalists and specialists, and their behavioral evolution. Ph.D. Dissertation, University of Tennessee 204p. 1980.
    • Farmer, P. and S.B. Heath. 1987. Wildlife baseline inventory, Rock Creek study area, Sanders County, Montana. Western Technology and Engineering, Inc. Helena, MT.
    • Farr, D.R. 1988. The ecology of garter snakes, Thamnophis sirtalis and T. elegans, in southeastern British Columbia. Unpubl. M.S. Thesis, University of Victoria, BC, Canada.
    • Farr, D.R. and P.T. Gregory. 1991. Sources of variation in estimating litter characteristics of the garter snake, Thamnophis elegans. Journal of Herpetology 25(3): 261-268.
    • Feder, M. E. and S.J. Arnold. 1982. Anaerobic metabolism and behavior during predatory encounters between snakes (Thamnophis elegans) and salamanders. (Plethodon Jordani). Oecologia 53(1): 93-97.
    • Feigley, H. P. 1997. Colonial nesting bird survey on the Bureau of Land Management Lewistown District: 1996. Unpublished report, U.S. Bureau of Land Management, Lewistown, Montana.
    • Finley, R.B., Jr., D. Chiszar, and H.M. Smith. 1994. Field observations of salivary digestion of rodent tissue by the wandering garter snake, Thamnophis elegans vagrans. Bulletin of the Chicago Herpetological Society 29(1): 5-6.
    • Fitch, H.S. 1940a. A biogeographical study of the ordinoides artenkreis of garter snakes (genus Thamnophis). University of California Publications in Zoology 44: 1-150.
    • Fitch, H.S. 1948. Further remarks concerning Thamnophis ordinoides and its relatives. Copeia 1948(2): 121-126.
    • Fitch, H.S. 1949. Study of snake populations in central California. American Midland Naturalist 41(3): 513-579.
    • Fitch, H.S. 1980a. Remarks concerning certain western garter snakes of the Thamnophis elegans complex. Transactions of Kansas Academy of Sciences 83: 106-113.
    • Fitch, H.S. 1983. Thamnophis elegans. Catalogue of American Amphibians and Reptiles 320.1-320.4.
    • Flath, D.L. 2002. Reptile and amphibian surveys in the Madison-Missouri River Corridor, Montana. Annual Progress Report. 14pp.
    • Fleharty, E.D. 1967. Comparative ecology of Thamnophis elegans, T. cyrtopsis, and T. rufipunctatus in New Mexico. Southwestern Naturalist 12: 207-230.
    • Flowers, M.A. and B.M. Graves. 1997. Juvenile toads avoid chemical cues from snake predators. Animal Behaviour 53(3): 641-646.
    • Ford, N.B. and G.A. Shuttlesworth. 1986. Effects of variation in food intake on locomotory performance of juvenile garter snakes. Copeia 1986(4): 999-1001.
    • Fox, W. 1948b. Effect of temperature on development of scutellation in the garter snake, Thamnophis elegans atratus. Copeia 1948(4): 252-262.
    • Fox, W. 1951. Relationships among the garter snakes of the Thamnophis elegans rassenkreis. University of California Publication of Zoology 50(5): 485-530.
    • Fox, W. 1952a. Notes on the feeding habits of Pacific coast garter snakes. Herpetologica 8: 4-8.
    • Fox, W. 1952b. Seasonal variation in the male reproductive system of Pacific coast garter snakes. Journal of Morphology 90: 481-554.
    • Fox, W. 1954. Genetic and environmental variation in the timing of the reproductive cycles of male garter snakes. Journal of Morphology 95: 415-450.
    • Fox, W. and H.C. Dessauer. 1964. Collection of garter snakes for blood studies. Yearbook of American Philosophy Society 1964: 263-266.
    • Fox, W., C. Gordon, and M.H. Fox. 1961. Morphological affect of low temperatures during the embryonic development of the garter snake, Thamnophis elegans. Zoologica 46: 57-71.
    • Garland, T., Jr. and S.J. Arnold. 1983. Effects of a full stomach on locomotory performance of juvenile garter snakes (Thamnophis elegans). Copeia 1983(4): 1092-1096.
    • Garner, T.W.J. and K.W. Larsen. 2005. Multiple paternity in the western terrestrial garter snake, Thamnophis elegans. Canadian Journal of Zoology 83(5):656-663.
    • Garner, T.W.J., P.B. Pearman, P.T. Gregory, G. Tomio, S.G. Wischniowski, and D.J. Hosken. 2004. Microsatellite markers developed from Thamnophis elegans and Thamnophis sirtalis and their utility in three species of garter snakes. Molecular Ecology Notes 4(3): 369-371.
    • Gates, M.T. 2005. Amphibian and reptile baseline survey: CX field study area Bighorn County, Montana. Report to Billings and Miles City Field Offices of Bureau of Land Management. Maxim Technologies, Billings, MT. 28pp + Appendices.
    • Goldberg, S.R. 2004b. Reproduction in the western terrestrial garter snake, Thamnophis elegans (Serpentes, Colubridae) from Arizona. Journal of the Arizona-Nevada Academy of Science 36(2):77-80.
    • Gomez, H.F., M. Davis, S. Phillips, P. McKinney, and J. Brent. 1994. Human envenomation from a wandering garter snake. Annals of Emergency Medicine 23(5): 1119-1122.
    • Graves, B.M. and D. Duvall. 1990. Spring emergence patterns of wandering garter snakes and prairie rattlesnakes in Wyoming. Journal of Herpetology 24(4): 351-356.
    • Gregory, P.T. 1967. Feeding habits and diet overlap of three species of garter snakes (Thamnophis) on Vancouver Island. Canadian Journal of Zoology 56: 1967-1974.
    • Gregory, P.T. 1978. Feeding habits and diet overlap of three species of garter snakes (Thamnophis) on Vancouver Island. Canadian Journal of Zoology 56: 1967-1974.
    • Gregory, P.T. 1984b. Correlations between body temperature and environmental factors and their variations with activity in garter snakes (Thamnophis). Canadian Journal of Zoology 62(11): 2244-2249.
    • Gregory, P.T. 1990. Temperature differences between head and body in garter snakes (Thamnophis) at a den in Central British Columbia (Canada). Journal of Herpetology 24(3): 241-245.
    • Gregory, P.T. and A.G.D. McIntosh. 1980. Thermal niche overlap in garter snakes (Thamnophis) on Vancouver Island. Canadian Journal of Zoology 58(3): 351-355.
    • Gregory, P.T. and C.J. Prelypchan. 1994. Analysis of variance of first-year growth in captive garter snakes (Thamnophis elegans) by family and sex. Journal of Zoology 232(2): 313-322.
    • Gregory, P.T. and K.M. Skebo. 1998. Trade-offs between reproductive traits and the influence of food intake during pregnancy in the garter snake, Thamnophis elegans. American Naturalist 151: 477-486.
    • Gregory, P.T., J.M. Macartney, and D.H. Rivard. 1980. Small mammal predation and prey handling behavior by the garter snake Thamnophis elegans. Herpetologica 36(1): 87-93.
    • Grismer, L.L. 1994. Food observations on the endemic Sierra San Pedro Martir garter snake (Thamnophis elegans hueyi) from Baja California, Mexico. Herpetological Natural History 2(1): 107-108.
    • Hammerson, G.A. 1999. Amphibians and reptiles in Colorado. University Press of Colorado & Colorado Division of Wildlife. Denver, CO. 484 p.
    • Hanauska-Brown, L., B.A. Maxell, A. Petersen, and S. Story. 2014. Diversity Monitoring in Montana 2008-2010 Final Report. Montana Fish, Wildlife & Parks. Helena, MT. 78 pp.
    • Hayden, F.V. 1858. Catalogue of the collections in geology and natural history, obtained by the expedition under command of Lieutenant G.K. Warren, Topographical Engineers. pp. 104-105. In: F.N. Shubert (1981) Explorer on the northern plains: Lieutenant Gouverneur K. Warren's preliminary report of explorations in Nebraska and Dakota, in the years 1855-'56-'57. Engineer Historical Studies No. 2. Office of the Chief of Engineers, Washington, DC. 125 p.
    • Hayden, F.V. 1862. On the geology and natural history of the upper Missouri. Transactions of the American Philosophical Society New Series 12(1): 1-218
    • Hebard, W.B. 1950. Relationships and variation in the garter snakes, genus Thamnophis, of the Puget Sound region of Washington state. Herpetologica 6: 97-101.
    • Heller, S. and M. Halpern. 1981. Laboratory observations on conspecific and congeneric scent trailing in garter snakes (Thamnophis). Behavioral Neural Biology 33(3): 372-377.
    • Hendricks, P. 1996. Geographical distribution. Thamnophis elegans vagrans. Herpetological Review 27(2):89.
    • Hendricks, P. 1996. Thamnophis elegans vagrans (wandering garter snake). Herpetological Review 26(2): 89.
    • Hendricks, P. 1997. Lee Metcalf National Wildlife Refuge preliminary amphibian and reptile investigations: 1996. Montana Natural Heritage Program, Helena, MT. 21 p.
    • Hendricks, P. 1999a. Amphibian and reptile survey of the Bureau of Land Management Miles City District, Montana. Montana Natural Heritage Program, Helena, MT. 80 p.
    • Hendricks, P. 2000. Amphibian and reptile survey of the Thompson Chain of Lakes. A report to the Montana Department of Fish, Wildlife, and Parks. Montana Natural Heritage Program, Helena, MT. 15 p.
    • Hendricks, P. and J.D. Reichel. 1996a. Amphibian and reptile survey of the Bitterroot National Forest: 1995. Montana Natural Heritage Program. Helena, MT. 95 p.
    • Hendricks, P. and J.D. Reichel. 1996b. Preliminary amphibian and reptile survey of the Ashland District, Custer National Forest: 1995. Montana Natural Heritage Program, Helena, MT. 79 p.
    • Hendricks, P. and J.D. Reichel. 1998. Amphibian and reptile survey on Montana refuges: 1996. Montana Natural Heritage Program, Helena, MT. 19 p.
    • Hendricks, P. and M. Roedel. 2001. A faunal survey of the Centennial Valley Sandhills, Beaverhead County, Montana. Report to the U.S. Bureau of Land Management and U.S. Fish and Wildlife Service. Montana Natural Heritage Program, Helena, MT. 44 p.
    • Herman, J.K. and R.L. Ingermann. 1996. Effects of hypoxia and hyperoxia on oxygen-transfer properties of the blood of a viviparous snake. Journal of Experimental Biology 199(9): 2061-2070.
    • Hicks, J.W., M.L. Riedesel, and M.L. Glass. 1979. Ventilative and gas exchange in Thamnophis elegans after acute and chronic exposure to various thermal environments. Journal of the Colorado-Wyoming Academy of Science 11(1): 99-100.
    • Highfill, D.R. 1976. Function of the corpora lutea of pregnancy in the garter snake (Thamnophis elegans). Ph. D. Dissertation, University of Idaho; 52p. 1975.
    • Hill, S.R. 1995. Migratory chronology of adult tiger salamanders (Ambystoma tigrinum) and survey of larvae of the tiger salamander in the northern range of Yellowstone National Park. Masters thesis, Montana State University. Bozeman, MT. 72 pp.
    • Hill, S.R. and R.E. Moore. 1994. Herpetological survey in the northern range of Yellowstone National Park. Annual Report, Yellowstone National Park. February 1, 1994. 21 pp.
    • Hill, S.R., Jr. and R.E. Moore. 1994a. Herpetological survey in the northern range of Yellowstone National Park. Investigator's Annual Reports Yellowstone National Park 1993. Yellowstone Center for Resources. pp. 96-97.
    • Hilliard, J., H. Minkus, and M. Weber. 1997. Amphibian survey of the Birch Creek drainage, Beaverhead County. Wildland Studies Project, San Francisco State University, San Francisco, CA. 12 p.
    • Hogg, J.B. 1974. Color convergence in the garter snakes Thamnophis elegans terrestris and Thamnophis couchi atratus along the central California coast. M.S. Thesis, California State Polytechnic University (Pomona). 95 p.
    • Hollenbeck, R.R. 1974. Growth rates and movements within a population of Rana pretiosa pretiosa Baird and Girard in south central Montana. Ph.D. Dissertation. Bozeman, Montana: Montana State University. 66 p.
    • Hosea, R.C., K.Z. Bjurstrom, and E.E. Littrell. 2005. Acute oral and dermal toxicity of aquatic herbicides and a surfactant to garter snakes. California Fish and Game 91(2):119-127.
    • Hossack, B., D. Pilliod, and P.S. Corn. 2001b. Preliminary amphibian surveys of the National Bison Range, Lost Trail National Wildife Refuge, and Swan River National Wildlife Refuge: 2001. USGS Northern Rocky Mountain Science Center, Aldo Leopold Wilderness Research Institute, Missoula, MT. 15 p.
    • Huey, R.B., C.R. Peterson, S.J. Arnold, and W.P. Porter. 1988. The thermal consequences of retreat site selection by garter snakes. American Zoologist 28(4): 46A.
    • Huey, R.B., C.R. Peterson, S.J. Arnold, and W.P. Porter. 1989. Hot rocks and not-so-hot rocks: retreat-site selection by garter snakes and its thermal consequences. Ecology 70(4): 931-944.
    • Ingermann, R.L., N.J. Berner, and F.R. Ragsdale. 1991a. Changes in red cell ATP concentration and oxygen-affinity following birth in the neonatal garter snake, Thamnophis elegans. Journal of Experimental Biology 157: 579-584.
    • Ingermann, R.L., N.J. Berner, and F.R. Ragsdale. 1991b. Effect of pregnancy and temperature on red cell oxygen-affinity in the viviparous snake Thamnophis elegans. Journal of Experimental Biology 156(0): 399-406.
    • James, D.K., L. Petrinovich, T.L. Patterson, and A.H. James. 1983. Predation of white-crowned sparrow nestlings by the western terrestrial garter snake in San Francisco, California. Copeia 1983(2): 511-513.
    • Jansen, D.W. 1980. Possible functions of Duvernoy's gland in the wandering garter snake (Thamnophis elegans vagrans). American Zoologist 20(4): 962.
    • Jansen, D.W. 1981. Allometric growth of the skull of Thamnophis elegans vagrans: adaptive potentials. American Zoologist 21(4): 958.
    • Jansen, D.W. 1987. The myonecrotic effect of Duvernoy's gland secretion of the snake Thamnophis elegans vagrans. Journal of Herpetology 21(1): 81-83.
    • Jean, C., P. Hendricks, M. Jones, S. Cooper, and J. Carlson. 2002. Ecological communities on the Red Rock Lakes National Wildlife Refuge: inventory and review of aspen and wetland systems. Report to Red Rock Lakes National Wildlife Refuge.
    • Jennings, W.B., D.F. Bradford, and D.F. Johnson. 1992. Dependence of the garter snake Thamnophis elegans on amphibians in the Sierra Nevada of California. Journal of Herpetology 26(4): 503-505.
    • Johnson, M.L. 1947. The status of the elegans subspecies of Thamnophis with description of a new subspecies from Washington State. Herpetologica 3: 159-165.
    • Joslin, Gayle, and Heidi B. Youmans. 1999. Effects of recreation on Rocky Mountain wildlife: a review for Montana. [Montana]: Montana Chapter of the Wildlife Society.
    • Kardong, K.V. and D.L. Luchtel. 1986. Ultrastructure of Duvernoy's gland from the wandering garter snake, Thamnophis elegans vagrans (Serpentes, Colubridae). Journal of Morphology 188(1): 1-14.
    • Kasper, S. and S.N. Kasper. 1997. Serpentes: Thamnophis elegans vagrans (wandering garter snake). Paralysis. Herpetological Review 28(1): 46.
    • Kelly, K.C., S.J. Arnold, and J. Gladstone. 1997. The effects of substrate and vertebral number on locomotion in the garter snake Thamnophis elegans. Functional Ecology 11: 189-198.
    • Kephart, D.G. 1981. Populaton ecology and population structure of Thamnophis elegans and Thamnophis sirtalis. Ph.D. Dissertation, Univeristy of Chicago; 1981.
    • Kephart, D.G. 1982. Microgeographic variation in the diets of garter snakes. Oecologia 52(2): 287-291.
    • Kingery, U.C. and H.E. Kingery. 2003. Standoff at a Chatfield marsh: house wrens and a garter snake. Colorado Birds 37(4):190-191.
    • Koch, E.D. and C.R. Peterson. 1989. A preliminary survey of the distribution of amphibians and reptiles in Yellowstone National Park. pp. 47-49. In: Rare, sensitive and threatened species of the Greater Yellowstone Ecosystem, T.W. Clark, A.H. Harvey, R.D. Dorn, D.C. Genter, and C. Groves (eds.), Northern Rockies Conservation Cooperative , Montana Natural Heritage Program, The Nature Conservancy, and Mountain West Environmental Services. 153 p.
    • Koch, E.D. and C.R. Peterson. 1995. Amphibians and reptiles of Yellowstone and Grand Teton national parks. University of Utah Press, Salt Lake City, UT. 188 p.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Big Spring Creek, Lewistown, Montana. Proj. No. 130091.029. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2002: Big Spring Creek, Lewistown, Montana. Proj. No. 130091.029. February 2003. In 2002 Wetland Mitigation Monitoring Reports, Vol. I.
    • Lawson, R. and H.C. Dessauer. 1979. Biochemical genetics and systematics of garter snakes of the Thamnophis-elegans-couchii-ordinoides complex. Occasional Papers of the Museum of Zoology of Louisiana State University 56: 1-24.
    • Lewis, M., Clark, W., and Members of the Corps of Discovery. (2002). The Journals of the Lewis and Clark Expedition (G. Moulton, Ed.). Lincoln: University of Nebraska Press. Retrieved Oct. 1, 2005, from the University of Nebraska Press / University of Nebraska-Lincoln Libraries-Electronic Text Center, The Journals of the Lewis and Clark Expedition.
    • Licht, E.L. 1985. Thamnophis elegans vagrans (gray garter snake), longevity. Bulletin of the Maryland Herpetological Society 21(4): 150.
    • Littrell, E.E. 1983. A study of the effects of bolero 10g on the mountain garter snake, Thamnophis elegans elegans. California Fish and Game 69(3): 186-187.
    • Macartney, J.M. and P.T. Gregory. 1981. Differential susceptibility of sympatric garter snake species to amphibian secretions. American Midland Naturalist 106: 271-281.
    • Manier, M.K. 2003. Does natural variation at neutral molecular markers reflect ecological variation in garter snakes? Abstract. Northwestern Naturalist 84:105.
    • Manier, M.K. 2005. Ecological correlates of population genetic structure for two coexisting garter snake species. Abstract. Northwestern Naturalist 86:106.
    • Manier, M.K. and S.J. Arnold. 2005. Population genetic analysis indentifies source-sink dynamics for two sympatric garter snake species (Thamnophis elegans and Thamnophis sirtalis). Molecular Ecology 14:3965-3976.
    • Manville, R.H. 1957. Amphibians and reptiles of Glacier National Park, Montana. Copeia 1957: 308-309.
    • Marnell, L. E. 1997. Herpetofauna of Glacier National Park. Northwestern Naturalist 78:17-33.
    • Matthews, K.R., R.A. Knapp, and K.L. Pope. 2002. Garter snake distributions in high-elevation aquatic ecosystems: is there a link with declining amphibian populations and nonnative trout introductions? Journal of Herpetology 36: 16-22.
    • Maxell, B.A. 2002a. Amphibian and aquatic reptile inventories in watersheds in the South and Middle Forks of the Flathead River drainage that contain lakes being considered for application of piscicides and subsequent stocking of west slope cutthroat trout. Report to the Region 1 Office of the U.S. Forest Service and the Montana Department of Fish, Wildlife, and Parks. Montana Cooperative Wildlife Research Unit and Wildlife Biology Program, University of Montana, Missoula, MT. 62 pp.
    • Maxell, B.A. 2016. Flammulated Owl surveys on the Big Timber, Bozeman, Gardiner and Livingston Ranger Districts of the Custer Gallatin National Forest: 2013. Report to Custer Gallatin National Forest. Montana Natural Heritage Program. Helena, MT. 27pp + appendices.
    • Maxell, B.A. 2016. Northern Goshawk surveys on the Beartooth, Ashland, and Sioux Districts of the Custer-Gallatin National Forest: 2012-2014. Montana Natural Heritage Program. Helena, MT. 114pp.
    • Maxell, B.A., J.K. Werner, P. Hendricks, and D.L. Flath. 2003. Herpetology in Montana: a history, status summary, checklists, dichotomous keys, accounts for native, potentially native, and exotic species, and indexed bibliography. Society for Northwestern Vertebrate Biology, Northwest Fauna Number 5. Olympia, WA. 135 p.
    • McDowell, S. and M. Manier. 2001. Do hungry snakes move faster than full snakes? Northwestern Naturalist 82(2):77.
    • Mead, R.A., V.P. Eroschenko, and D.R. Highfill 1981. Effects of progesterone and estrogen on the histology of the oviduct of the garter snake, Thamnophis elegans. General Comparison of Endocrinology 45(3): 345-354.
    • Miller, J. D. 1975. Interspecific food relationships of anurans in northwestern Montana and fluoride accumulation in amphibians and reptiles in northwestern Montana. M.S. thesis. University of Montana, Missoula, MT. 105 p.
    • Minton, S.A. and S.A. Weinstein. 1987. Colubrid snake venoms: Immunologic relationships, electrophoretic patterns. Copeia 1987(4): 993-1000.
    • Morey, S.R. 1990. Microhabitat selection and predation in the Pacific Treefrog, Pseudacris regilla. Journal of Herpetology 24(3): 292-296.
    • Moulton, G.E. editor. 1983. The Journals of the Lewis and Clark Expedition. University of Nebraska Press, Lincoln & London.
    • Nicol, J.T., R. Demaree, and D.M. Wootton. 1985. Levinseniella ophidea, new species (Trematoda: Microphallidae) from the western garter snake, Thamnophis elegans and the bullfrog, Rana catesbeiana. Proceedings of The Helminthological Society of Washington 52(2): 180-183.
    • Northrop, Devine and Tarbell, Inc. 1995. Cabinet Gorge and Noxon Rapids Hydroelectric Developments, 1994 Wetland Mapping and Assessment Study, Volume I of II. 27 pp. plus appendices.
    • Oechsli, L.M. 2000. Ex-urban development in the Rocky Mountain West: consequences for native vegetation, wildlife diversity, and land-use planning in Big Sky, Montana. M.Sc. Thesis. Montana State University, Bozeman. 73 p.
    • Osypka, N.M. and S.J. Arnold. 2000. The development effect of sex ratio on a sexually dimorphic scale count in the garter snake Thamnophis elegans. 34(1): 1-5.
    • Panik, H.R. and S. Barrett. 1994. Distributions of amphibians and reptiles along the Truckee River System. Northwest Science 68(3):197-204.
    • Patla, D.A. 1998a. Amphibians and reptiles in the Old Faithful sewage treatment area. Report to Yellowstone Center for Resources, Yellowstone National Park. 10 September, 1998. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 7 p.
    • Patla, D.A. 1998b. Potential effects of native fish restoration projects on amphibians in Yellowstone National Park Part I. Report to National Park Service, Yellowstone National Park. 20 November 1998. 26 pp.
    • Patla, D.A. 1999a. Amphibians and reptiles along the grand loop road in Yellowstone National Park: Canyon Junction to Fishing Bridge Junction. December 11, 1999. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 24 p.
    • Patla, D.A. 1999b. Amphibians and reptiles of the Madison to Norris road improvement project area, Yellowstone National Park. 11 November, 1999. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 17 p.
    • Patla, D.A. 1999c. Amphibians and reptiles, Tower Junction to Canyon Village, Yellowstone National Park; addendum to previous report. 24 November, 1999. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 10 p.
    • Patla, D.A. and C.R. Peterson. 1996a. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Tower Junction to Canyon Village. 24 February, 1996. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 49 p.
    • Patla, D.A. and C.R. Peterson. 1996b. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Arnica Creek to Little Thumb Creek. 1 August, 1996. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 35 p.
    • Patla, D.A. and C.R. Peterson. 1997. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Mammoth Hot Springs to Norris Junction. 1 February, 1997. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 60 p.
    • Petersen, C.R. and S.J. Arnold. 1986. Individual variation in the thermoregulatory behavior of free-ranging garter snakes, Thamnophis elegans. American Zoologist 26(4): 112A.
    • Peterson, C.R. 1978. Constriction in the wandering garter snake. American Zoologist 18(3): 649.
    • Peterson, C.R. 1979. Body temperature variation in free-living wandering garter snakes. American Zoologist 19(3): 975.
    • Peterson, C.R. 1981. Body temperature and energy balance in wandering garter snakes. American Zoologist 21(4): 957.
    • Peterson, C.R. 1983. Body temperature variation in free-living garter snakes (Thamnophis elegans vagrans). Ph.D. Dissertation, Washington State University; 186p. 1982.
    • Peterson, C.R. 1987. Daily variation in the body temperatures of free-ranging garter snakes. Ecology 68(1): 160-169.
    • Peterson, C.R. and H.J. Fabian. 1984. Thamnophis elegans vagrans (wandering garter snake). Coloration. Herpetological Review 15(4): 113.
    • Peterson, C.R. and J.P. Shive. 2002. Herpetological survey of southcentral Idaho. Idaho Bureau of Land Management Technical Bulletin 02-3:1-97.
    • Peterson, C.R., A.J. Vitale, and V.A. Cobb. 1989. Measuring the acitivity patterns of free-ranging animals with radio-telemetry. American Zoologist 29(4): 43A.
    • Peterson, C.R., C.J. Askey, and D.A. Patla. 1993. Amphibians and reptiles along the Grand Loop and Fountain Freight Roads between Madison Junction and Biscuit Basin in Yellowstone National Park. 26 July, 1993. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 45 p.
    • Peterson, C.R., D.A. Patla, and S.R. Sullivan. 1995. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Madison Junction to Norris Campground. 7 July, 1995. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 64 p.
    • Pilliod, D.S. and T.C. Esque. 2023. Amphibians and reptiles. pp. 861-895. In: L.B. McNew, D.K. Dahlgren, and J.L. Beck (eds). Rangeland wildlife ecology and conservation. Cham, Switzerland: Springer. 1023 p.
    • Queiroz, A. and R.R. Groen. 2001. The inconsistent and inefficient constricting behavior of Colorado western terrestrial garter snakes, Thamnophis elegans. Journal of Herpetology 35(3): 450-460.
    • Rauscher, R.L. 1998. Amphibian and reptile survey on selected Montana Bureau of Reclamation impoundments. Montana Department of Fish, Wildlife and Parks, Nongame Program. Bozeman, MT. 24 pp.
    • Rauscher, R.L. 2000. Tiger salamander axolotls in southwest Montana, final report. Bozeman, MT: Montana Department of Fish, Wildlife and Parks. 28p.
    • Reaser, J.K. and R.E. Dexter. 1996a. Rana pretiosa (spotted frog) predation. Herpetological Review 27(2): 75.
    • Reichel, J. and D. Flath. 1995. Identification of Montana's amphibians and reptiles. Montana Outdoors 26(3):15-34.
    • Reichel, J. D. In prep. Amphibian and reptile survey in southwest Montana: 1996. Unpublished report.
    • Reichel, J.D. 1995a. Preliminary amphibian and reptile survey of the Lewis & Clark National Forest: 1994. Montana Natural Heritage Program. Helena, MT. 92 p.
    • Reichel, J.D. 1996. Preliminary amphibian and reptile survey of the Helena National Forest: 1995. Montana Natural Heritage Program. Helena, MT. 87 pp.
    • Reichel, J.D. 1997a. Amphibian, reptile and northern bog lemming survey on the Rocky Mountain Front: 1996. Montana Natural Heritage Program, Helena, MT. 81 p.
    • Ridenhour, B.J., E.D. Brodie, Jr., and E.D. Brodie III. 1999. Effects of repeated injection of tetrodotoxin on growth and resistance to tetrodotoxin in the garter snake Thamnophis sirtalis. Copeia 1999: 531-535.
    • Rodgers, T. L. and W. L. Jellison. 1942. A collection of amphibians and reptiles from western Montana. Copeia (1):10-13.
    • Roedel, M.D. and P. Hendricks. 1998a. Amphibian and reptile survey on the Bureau of Land Management Lewistown District: 1995-1998. Montana Natural Heritage Program, Helena, MT. 53 p.
    • Roedel, M.D. and P. Hendricks. 1998b. Amphibian and reptile inventory on the Headwaters and Dillon Resource Areas in conjunction with Red Rocks Lakes National Wildlife Refuge: 1996-1998. Montana Natural Heritage Program, Helena, MT. 46 p.
    • Rosenberg, H.I. 1973. Functional anatomy of pulmonary ventilation in the garter snake, Thamnophis elegans. Journal of Morphology 140(2): 171-184.
    • Rossman, D.A. 1963a. Relationships of the elegans complex of the garter snakes genus Thamnophis. Yearbook of the American Philosophical Society. 1963: 347-348.
    • Rossman, D.A. 1979. Morphological evidence for taxonomic partitioning of the Thamnophis elegans complex (serpentes, colubridae). Occasional Papers of the Museum of Zoology, Louisiana State University 55: 12p.
    • Rossman, D.A., N.B. Ford, and R.A. Seigel. 1996. The garter snakes: evolution and ecology. Norman: University of Oklahoma Press.
    • Ruthven, A.G. 1908. Variations and genetic relationships of the garter snakes. Bulletin of the U.S. National Museum 61: xii + 201pp.
    • Scott, J.R. 1978. Thermal biology of the wandering garter snake. Ph.D. Dissertation, Colorado State University, Fort Collins, Colorado.
    • Scott, J.R. and D. Pettus. 1979. Effects of seasonal acclimation on the preferred body temperature of (thamnophis elegans vagrans). Journal of Thermal Biology 4(4): 307-309.
    • Scott, J.R., C.R. Tracy, and D. Pettus. 1982. A biophysical analysis of daily and seasonal utilization of climate space by a montane snake. Ecology 63(2): 482-493.
    • Scott, N.J., Jr. and A.L. Aquino-Shuster. 1989. The effects of freezing on formalin preservation of specimens of frogs and snakes. Collections Forum 5(2): 41-46.
    • Seidel, M.E. and R.G. Lindeborg. 1973. Lags in metabolic response to temperature of two garter snakes, Thamnophis elegans and Thamnophis radix. Herpetological 29(4): 358-360.
    • Sestrich, Clint. 2006. 2006 Hebgen Reservoir Amphibian Survey, USDA Forest Service Annual Progress Report to PPL Montana. Hebgen Lake Ranger District. Gallatin National Forest. West Yellowstone Montana.
    • Seyler, C. and S.J. Arnold. 2005. Selection on coloration in the garter snake Thamnophis elegans. Abstract. Northwestern Naturalist 86:115.
    • Skinner, M.P. 1924. The Yellowstone Nature Book. A.C. McClurg Company, Chicago, IL. 221 p.
    • Smith, H.M., G.L. Smith, and D. Chiszar. A new record and review of partially scaleless snakes. Bulletin of the Maryland Herpetological Society 32(4): 107-112.
    • Soderquist, T.R. and N.R. Middlebrook. 1984. New herpetological records from Arizona north of the Grand Canyon. Herpetological Review 15(4): 115.
    • St. John, A.D. 2002. Reptiles of the northwest: California to Alaska, Rockies to the coast. Lone Pine Publishing, Renton, WA. 272 p.
    • Stebbins, R. C. 2003. A field guide to western reptiles and amphibians. 3rd Edition. Houghton Mifflin Company, Boston and New York. 533 p.
    • Stevenson, R.D., C.R. Peterson and J. Tsuji. 1982. Locomotion, digestion and tongue flicking of the garter snake, Thamnophis elegans, at different body temperatures. America Zoologist 22(4): 916.
    • Stevenson, R.D., C.R. Peterson, and J.S. Tsuji. 1985. The thermal dependence of locomotion, tongue flicking, digestion and oxygen consumption in the wandering garter snake Thamnophis elegans. Physiological Zoology 58(1): 46-57.
    • Storey, K.B. 1996. Metabolic adaptations supporting anoxia tolerance in reptiles: Recent advances. Comparative Biochemisty and Physiology B 113(1): 23-35.
    • Stuart, J.N. 1987. Serpentes. Thamnophis elegans vagrans (wandering garter snake). Herpetological Review 18(1): 21.
    • Sullivan, S.R. and C.R. Peterson. 1996. Amphibians and reptiles along the highway in Yellowstone National Park: Tower Junction to the Northeast Entrance. 25 February, 1996. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 60 p.
    • Szaro, R.C., S.C. Belfit, J.K. Aitkin, and J.N. Rinne. 1985. Impact of grazing on a riparian garter snake. Pages 359-363 in R.R. Johnson, C.D. Ziebell, D.r. Patton, P.F. Ffolliott and F.H. Hamre, technical coordinators. Riparian ecosystems and their management: reconciling conflicting uses. General Technical Report RM-120. U.S. Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
    • Szaro, R.C., S.C. Belfit, J.K. Aitkin, and R.D. Babb. 1988. The use of timed fixed-area plots and a mark-recapture technique in assessing riparian garter snake populations. U.S. Forerst Service General Technical Report RM-166: 239-246.
    • Tanner, W.W. 1949. Food of the wandering garter snake, Thamnophis elegans vagrans (Baird & Girard), in Utah. Herpetologica 5: 85-86.
    • Tanner, W.W. 1950. Variation in scale and color pattern of the wander garter snake, in Utah and southern Idaho. Herpetologica 6: 194-196.
    • Tanner, W.W. and C.H. Lowe. 1989. Variations in Thamnophis elegans with descriptions of new subspecies. Great Basin Naturalist 49(4): 511-516.
    • Test, F.C. 1893. Annotated list of reptiles and batrachians collected. In B.W. Evermann. A reconnaisance of the streams and lakes of western Montana and northwestern Wyoming. Bulletin of United States Fish Commission 11(1891): 57-59.
    • Thompson, L. S. 1985. Montana’s explorers: The pioneer naturalists, 1805-1864. Montana Geographic Series No. 9. Montana Magazine, Helena.
    • Timken, R. No Date. Amphibians and reptiles of the Beaverhead National Forest. Western Montana College, Dillon, MT. 16 p.
    • Todd, F. 1965. Scale variation among garter snakes with particular emphasis on the upper and lower labials. Unpublished research project for Dr. R.B. Brunson. 27 pp.
    • Turner, F.B. 1951. A checklist of the reptiles and amphibians of Yellowstone National Park with incidental notes. Yellowstone Nature Notes 25(3): 25-29.
    • Turner, F.B. 1955. Reptiles and amphibians of Yellowstone National Park. Yellowstone Interpretive Series No. 5. Yellowstone Library and Museum Association. Yellowstone National Park, WY. 40 p.
    • Turner, F.B. 1957. The ecology and morphology of Rana pretiosa pretiosa in Yellowstone Park, Wyoming. Ph.D. dissertation, University of California. Berkeley, CA. 252 pp.
    • Van Leuven, J. 2008. Survey of melanistic garter snakes by traditional and geometric morphometrics. Undergraduate Thesis. Department of Natural Sciences, Carroll College, Helena, MT. 22pgs.
    • Velhagen, W.A. and A.H. Savitzky. 1998. Evolution of embryonic growth in thamnophiine snakes. Copeia 1998: 549-558.
    • Vest, D.K. 1981a. Envenomation following the bite of a wandering garter snake (Thamnophis elegans vagrans). Clinical Toxicology 18(5): 537-579.
    • Vest, D.K. 1981b. The toxic Duvernoy's secretion of the wandering garter snake, Thamnophis elegans vagrans. Toxicon 19(6): 831-839.
    • Vitt, L.J., J.P. Caldwell, and D.B. Shepard. 2005. Inventory of amphibians and reptiles in the Billings Field Office Region, Montana. Sam Noble Oklahoma Museum of Natural History and Department of Zoology, University of Oklahoma, Norman, OK. 33 pp.
    • VTN Colorado, Inc. Decker Coal Company., 1975, Draft environmental impact assessment for the proposed North Extension of the West Decker Mine.
    • Waye, H.L. 1999. Size and age structure of a population of Western Terrestrial Garter Snakes (Thamnophis elegans). Copeia 1999(3):819-823.
    • Waye, H.L. and P.T. Gregory. 1993. Choices of neonate Thamnophis elegans vagrans between conspecific, congeneric, and heterogeneric odors. Journal of Herpetology 27(4): 435-441.
    • Waye, H.L. and P.T. Gregory. 1998. Determining the age of garter snakes (Thamnophis spp.) by means of skeletochronology. Canadian Journal of Zoology 76: 288-294.
    • Weaver, R.E. 2004b. Thamnophis elegans (Western Terrestrial Garter Snake). Herpetological Review 35(3):278.
    • Weaver, R.E. and B.A. Bauer. 2005. Diet and foraging behavior of the wandering garter snake (Thamnophis elegans vagrans) at Umtanum Creek, Kittitas County, Washington. Abstract. Northwestern Naturalist 86:121.
    • Webb, R.G. 1976. A review of the garter snake (Thamnophis elegans) in Mexico. Natural History Museum of Los Angeles County Contributions to Science No. 284. 13 p.
    • Weisel, G.F. 1952. Animal names, anatomical terms, and some ethnozoology of the Flathead Indians. Journal of the Washington Academy of Sciences 42(11): 345-355.
    • Weitzel, N.H. and H.R. Panik. 1993. Long-term fluctuations of an isolated population of the Pacific chorus frog (Pseudacris regilla) in northwestern Nevada. Great Basin Naturalist 53(4): 379-384.
    • Weldon, P.J. 1982. Responses to ophiophagous snakes by snakes of the genus Thamnophis. Copeia 1982(4): 788-794.
    • Werner, J.K. and J.D. Reichel. 1994. Amphibian and reptile survey of the Kootenai National Forest: 1994. Montana Natural Heritage Program. Helena, MT. 104 p.
    • Werner, J.K. and J.D. Reichel. 1996. Amphibian and reptile monitoring/survey of the Kootenai National Forest: 1995. Montana Natural Heritage Program. Helena, MT. 115 pp.
    • Werner, J.K. and T. Plummer. 1995a. Amphibian and reptile survey of the Flathead Indian Reservation 1993-1994. Salish Kootenai College, Pablo, MT. 55 pp.
    • Werner, J.K. and T. Plummer. 1995b. Amphibian monitoring program on the Flathead Indian Reservation 1995. Salish Kootenai College, Pablo, MT. 46 p.
    • Werner, J.K., B.A. Maxell, P. Hendricks and D.L. Flath. 2004. Amphibians and Reptiles of Montana. Mountain Press Publishing Company: Missoula, MT. 262 pp.
    • Werner, J.K., T. Plummer, and J. Weaslehead. 1998a. Amphibians and reptiles of the Flathead Indian Reservation. Intermountain Journal of Sciences 4(1-2): 33-49.
    • White, M. and J.A. Kolb. 1974. A preliminary study of Thamnophis near Sagehen Creek, California. Copeia 1974(1): 126-136.
    • Wirsing, A.J., J.D. Roth, and D.L. Murray. 2005. Can prey use dietary cues to distinguish predators? A test involving three terrestrial amphibians. Herpetologica 61(2):104-110.
    • Wright, D.L.., K.V. Kardong, and D.L. Bentley. 1979. The functional anatomy of the teeth of the western terrestrial garter snake, Thamnophis elegans. Herpetologica 35(3): 223-228.
    • Yarrow, H.C. 1882. Check list of North American reptilia and batrachia, with catalogue of specimens in the U.S. National Museum. United States National Museum Bulletin 24. 249 p.
    • Yeager, D.C. 1929. Reptiles of Yellowstone Park with notes on the Amphibia. Yellowstone Nature Notes. Yellowstone National Park Library, Mammoth, WY. 12 p.
    • Zisook, R., K. Almond, and B. Sharpe. 1996. Amphibian survey of the Birch Creek drainage, Beaverhead County. Wildland Studies Project. San Francisco State University, San Francisco, CA. 9 p.
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Citation for data on this website:
Terrestrial Gartersnake — Thamnophis elegans.  Montana Field Guide.  .  Retrieved on , from