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Loggerhead Shrike - Lanius ludovicianus
Species of Concern Native Species
Global Rank :
G4
State Rank :
S3B
Agency Status
USFWS :
MBTA
USFS :
BLM :
SENSITIVE
FWP SWAP :
SGCN3
PIF :
2
External Links
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Copyright by Borror Laboratory of Bioacoustics, Department of Evolution, Ecology, and Organismal Biology, Ohio State University, Columbus, OH, all rights reserved.
General Description
Slightly smaller than the American Robin (
Turdus migratorius ), the total length of this bird averages 23 cm. Males and females are similar in appearance. This species has a stout, hooked bill that has dark upper and lower mandibles. It has a broad black mask extending above the eye and thinly across top of bill. Its head and back are covered with a bluish-gray cowl, while its underparts and rump are white or grayish-white (underparts are very faintly barred in adults). It has a black tail with white tip and large white patches on black wings. Juveniles are paler and barred overall, with brownish-gray upperparts and buffy wing patches (Miller 1931, Fraser and Luukkonen 1986). Most nests are made of coarse twigs with a lining of plant material and animal hair (Fraser and Luukkonen 1986).
For a comprehensive review of the conservation status, habitat use, and ecology of this and other Montana bird species, please see
Marks et al. 2016, Birds of Montana.
Diagnostic Characteristics
Loggerhead Shrikes differ from Northern Shrikes (Lanius excubitor ) by having the base of the lower mandible black instead of pale, unbarred or barely barred underparts (adults), a shorter and less hooked bill, a darker head and back, and a more extensive black mask. They differ from the Northern Mockingbird (Mimus polyglottos ) by having a black mask and a shorter, less curved bill.
Species Range
Montana Range
Range Descriptions
Western Hemisphere Range
Observations in Montana Natural Heritage Program Database
Number of Observations: 4234
(Click on the following maps and charts to see full sized version)
Map Help and Descriptions
Relative Density
Recency
SUMMER (Feb 16 - Dec 14)
Direct Evidence of Breeding
Indirect Evidence of Breeding
No Evidence of Breeding
WINTER (Dec 15 - Feb 15)
Regularly Observed
Not Regularly Observed
(Observations spanning multiple months or years are excluded from time charts)
Migration
In the Bozeman area, normal migration periods are April 20 to May 15 and August 10 to September 10.
Habitat
Open landscapes with short vegetation, including pastures with fence rows, mowed roadsides, agricultural fields, riparian areas, and open woodlands (Yosef 1996). In Idaho, nests are found in sagebrush (65%), bitterbush, and greasewood, and are equally successful in all three (Woods and Cade 1996).
Ecological Systems Associated with this Species
Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system,
even if point observations were associated with that system.
Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.
If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at:
mtnhp.org/requests ) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.
Literature Cited
Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
Maxell, B.A. 2000. Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
Commonly Associated with these Ecological SystemsForest and Woodland Systems
Grassland Systems
Human Land Use
Recently Disturbed or Modified
Shrubland, Steppe and Savanna Systems
Sparse and Barren Systems
Wetland and Riparian Systems
Occasionally Associated with these Ecological SystemsHuman Land Use
Recently Disturbed or Modified
Food Habits
Diet includes arthropods, amphibians, small to medium-sized reptiles, small mammals and birds. Hunts from perch and impales prey on sharp objects. (Yosef 1996)
Ecology
Highly territorial. Uniquely placed in the food chain as both a passerine and top-level predator. (Yosef 1996)
Reproductive Characteristics
Loggerhead Shrikes nest from mid-June to mid-July (Davis 1961).
Management
Declining across the continent, probably in large part due to changes in land use, spraying of biocides, and competition with species that better tolerate human-caused changes (Yosef 1996).
Stewardship Responsibility
References
Literature Cited AboveLegend: View Online Publication Davis, C.V. 1961. A distributional study of the birds of Montana. Ph.D. Dissertation. Corvallis, OR: Oregon State University. 462 p. Fraser, J. D. and D. R. Luukkonen. 1986. The loggerhead shrike. In: R. L. DiSilvestro, ed. Audubon Wildlife Report 1986. Academic Press, New York. p. 933-941. Marks, J.S., P. Hendricks, and D. Casey. 2016. Birds of Montana. Arrington, VA. Buteo Books. 659 pages. Miller, A. H. 1931. Systematic revision and natural history of the American shrikes (Lanius ). University of California Publications in Zoology 38(2):11-242. Woods, C.P. and T.J. Cade. 1996. Nesting habits of the loggerhead shrike in sagebrush. Condor 98: 75-81.
Additional ReferencesLegend: View Online Publication Do you know of a citation we're missing? American Ornithologists’ Union [AOU]. 1998. Check-list of North American birds, 7th edition. American Ornithologists’ Union, Washington, D.C. 829 p. Anderson, D.E., O.J. Rongstad, and W.R. Mytton. 1985. Line transect analysis of raptor abundance along roads. Wildl. Soc. Bull. 13(4):533-539. Bird Conservancy of the Rockies. 2017. Pocket Guide to Northern Prairie Birds. Brighton, CO: Bird Conservancy of the Rockies. 98 p. Bleho, B., K. Ellison, D.P. Hill and L.K. Gould. 2015. Chestnut-collared Longspur (Calcarius ornatus ), The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology; Retrieved from the Birds of North America Online: http://bna.birds.cornell.edu/bna/species/288 (Accessed 18 March 2016) Bramblett, R.G., and A.V. Zale. 2002. Montana Prairie Riparian Native Species Report. Montana Cooperative Fishery Research Unit, Montana State University - Bozeman. Cameron, E. S. 1908. The birds of Custer and Dawson counties, Montana. Auk 25:39-56. Casey, D. 2000. Partners in Flight Draft Bird Conservation Plan Montana. Version 1.0. 287 pp. Chalfoun, A. 2005. Habitat use and quality for non-game shrub-steppe birds, Final performance report Confluence Consulting Inc. 2013. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT. Dechant, J. A., M. L. Sondreal, D. H. Johnson, L. D. Igl, C. M. Goldade, M. P. Nenneman, A. L. Zimmerman, and B. R. Euliss. 2003e. Effects of management practices on grassland birds: Loggerhead Shrike. Northern Prairie Wildlife Research Center, Jamestown, ND. Northern Prairie Wildlife Research Center Dobkin, D. S. 1992. Neotropical migrant landbirds in the Northern Rockies and Great Plains. U.S.D.A. For. Serv. N. Region Publ. R1-93-34. Missoula, Mont. Dobkin, D.S. 1994. Conservation and management of neotropical migrant landbirds in the Northern Rockies and Great Plains. Univ. Idaho Press, Moscow, Idaho. 220 pp. Dood, A.R. 1980. Terry Badlands nongame survey and inventory final report. Montana Department of Fish, Wildlife, and Parks and Bureau of Land Management, Helena, MT. 70 pp. DuBois, K.L. 1979. An inventory of the avifauna in the Long Pines of Southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 113 p. Ecological Consulting Service, Helena, MT., 1975, Peabody Coal Company Big Sky Mine: Wildlife environmental indicators monitoring survey. Progress report. Project 58-23-A. June 16, 1975. ECON, Inc. (Ecological Consulting Service), Helena, MT., 1975, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1975. Proj. 71-23-A. December 31, 1975. ECON, Inc. (Ecological Consulting Service), Helena, MT., 1976, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1976. Proj. 135-85-A. December 31, 1976. ECON, Inc. (Ecological Consulting Service), Helena, MT., 1977, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1977. Proj. 164-85-A. December 31, 1977. ECON, Inc. (Ecological Consulting Service), Helena, MT., 1979, Annual wildllife report of the Colstrip Area for 1978. Proj. 195-85-A. April 6, 1979. ECON, Inc. (Ecological Consulting Service), Helena, MT., 1979, Annual wildllife report of the Colstrip Area for 1979, including a special raptor research study. Proj. 216-85-A. March 1, 1980. ECON, Inc. (Ecological Consulting Service), Helena, MT., 1979, Area B four-section wildlife report. August 3, 1979. Econ, Inc. 1988. Wildlife monitoring report, 1987 field season, Big Sky Mine. March 1988. In Peabody Mining and Reclamation Plan Big Sky Mine Area B. Vol. 8, cont., Tab 10 - Wildlife Resources. Appendix 10-1, 1987 Annual Wildlife Report. Econ, Inc., Helena, MT., 1991, Wildlife monitoring report: 1989 field season, Big Sky Mine. March 1991. Ehrlich, P., D. Dobkin, and D. Wheye. 1988. The birder’s handbook: a field guide to the natural history of North American birds. Simon and Schuster Inc. New York. 785 pp. Faanes, C.A. 1983. Breeding birds of wooded draws in western North Dakota. Prairie Nat. 15(4): 173-187. Farmer, Patrick J., and Thomas W. Butts, Western Technology & Eng., Inc., Helena, MT., 1994, McDonald Project Terrestrial Wildlife Study, November 1989 - November 1993. April 1994. In McDonald Gold Project: Wildlife & Fisheries. [#18]. Seven-up Pete Joint Venture, Lincoln, MT. Unpub. No date. Feigley, H. P. 1997. Colonial nesting bird survey on the Bureau of Land Management Lewistown District: 1996. Unpublished report, U.S. Bureau of Land Management, Lewistown, Montana. Fink, D., K.V. Rosenberg, F.A. La Sorte, M.J. Iliff, C. Wood, and S. Kelling. 2013. Species distribution modeling of priority bird species on Bureau of Land Management lands to determine stewardship responsibility for conservation planning. Ithaca, NY: Cornell Lab of Ornithology. 224 p. Fjell, Alan K., and Brian R. Mahan, compilers., 1984, Peabody Coal Company Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1983 field season. February 1984. Fjell, Alan K., and Brian R. Mahan., 1985, Peabody Coal Company Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1984 field season. February 1985. Fjell, Alan K., and Brian R. Mahan., 1987, Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1986 field season. April 1987. Gillihan, SW. and T. VerCauteren. 2015. Pocket Guide to Prairie Birds. Brighton, CO: Bird Conservancy of the Rockies. 91 p. Gniadek, S. 1983. Southwest Glendive Wildlife Baseline Inventory. Miles City, Mont: Bureau of Land Management, Miles City District Office. 56 pp with appendices. Hays, R., R.L. Eng, and C.V. Davis (preparers). 1984. A list of Montana birds. Helena, MT: MT Dept. of Fish, Wildlife & Parks. Hejl, S.J., R.L. Hutto, C.R. Preston, and D.M. Finch. 1995. The effects of silvicultural treatments on forest birds in the Rocky Mountains. pp. 220-244 In: T.E. Martin and D.M. Finch (eds). Ecology and Management of Neotropical Migratory Birds. New York, NY: Oxford Univ. Press. 489 p. Hendricks, P. 2000. Roadside bird counts on BLM lands in Petroleum and Fergus Counties, Montana. Montana Natural Heritage Program, Helena, MT. 57pp. Hendricks, P., and K.H. Dueholm. 1995. Cliff-nesting raptor survey of the Sioux District, Custer National Forest: 1994. Unpubl. report to U.S.D.A. For. Serv., Custer N.F., Billings. 20 pp. Hendricks, P., G.M. Kudray, S. Lenard, and B.A. Maxell. 2007. A Multi-Scale Analysis Linking Prairie Breeding Birds to Site and Landscape Factors Including USGS GAP Data. Helena, Mont: Montana Natural Heritage Program. Hendricks, P., S. Lenard, C. Currier, and J. Carlson. 2007. Grassland bird surveys in north Valley County, Montana: 2001-2006. Report to the Bureau of Land Management, Glasgow Field Office. Montana Natural Heritage Program, Helena, Montana. 19 pp. plus appendices. Hendricks, P., S. Lenard, D.M. Stagliano, and B.A. Maxell. 2013. Baseline nongame wildlife surveys on the Fort Peck Indian Reservation. Report to the Assiniboine and Sioux Tribes of the Fort Peck Indian Reservation. Montana Natural Heritage Program, Helena, MT. 83 p. Herbert, J.T. 1977. An inventory of the bird population within the Sarpy Creek drainage, southeastern Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 81 p. Humphris, Michael., 1990, Wildlife Monitoring Report. Spring Creek Coal Company 1990 Mining Annual Report. Appendix I. April 11, 1990. Humphris, Michael., 1993, Wildlife Monitoring Report. Spring Creek Coal Company 1993 Mining Annual Report. Appendix I. April 11, 1993. Humphris, Michael., 1994, Wildlife Monitoring Report. Spring Creek Coal Company 1994 Mining Annual Report. Appendix I. April 1994. Jewett, S.G., W.P. Taylor, W.T. Shaw, and J.W. Aldrich. 1953. Birds of Washington state. Univ. Washington Press, Seattle. 767 pp. Johnsgard, P.A. 1979. Birds of the Great Plains: breeding species and their distribution. University of Nebraska Press, Lincoln. 539 pp. Johnsgard, P.A. 1992. Birds of the Rocky Mountains with particular reference to national parks in the northern Rocky Mountain region. Lincoln: University of Nebraska Press. xi + 504 pp. Joslin, Gayle, and Heidi B. Youmans. 1999. Effects of recreation on Rocky Mountain wildlife: a review for Montana . [Montana]: Montana Chapter of the Wildlife Society. Knowles, C. J. and P. R. Knowles. 1995. A survey of biological resources at the proposed Malta, MT airport site. Unpublished report. FaunaWest Wildlife Consultants, Boulder, MT. 6 pp. Lenard, S. 2006. Birds of Blaine County, Riparian Point Count Surveys 2005. Report to the Bureau of LandManagement, Havre Field Station, Havre, Montana. Montana Natural Heritage Program, Helena, MT. 16pp.plus appendices. Lenard, S., Compiler. 2005. Surveys for Animal Species of Concern in Sage and Grassland Landscapes in Montana. An unpublished report to the Montana Department of Fish, Wildlife & Parks, State Wildlife Grants Program. Montana Natural Heritage Program, Helena, Montana. 63pp. Lenard, S., J. Carlson, J. Ellis, C. Jones, and C. Tilly. 2003. P. D. Skaar's Montana bird distribution, 6th edition. Montana Audubon, Helena, MT. 144 pp. Lyons, T.P., J.R. Miller, D.M. Debinski, and D.M. Engle. 2014. Predator identity influences the effect of habitat management on nest predation. Ecological Applications 25(6): 1596-1605. Matthews, W.L. 1980a. Wibaux-Beach comparison study: Sydney, Glendive and Plevna Study Areas. Bureau of Land Management, Miles City, MT. 50 p. Matthews, W.L. 1981. Broadus-Pumpkin Creek baseline inventory - wildlife. Bureau of Land Management, Miles City, MT. 83 p. Maxim Technologies, Inc., 2002, Western Energy Company Rosebud Mine, Colstrip, Montana: 2002 Annual Wildlife Monitoring Report; December 1, 2001 - November 30, 2002. Febr. 24, 2002. Maxim Technologies, Inc., Helena, MT., 1999, Wetlands Study Report Dry Prairie Rural Water. November 1999. Milburn, T. 1981. Status and distribution of the Loggerhead Shrike, (Lanius ludovicianus in the Northeastern U.S.. Unpublished Report. Submitted to the Usdi/fws Office of Endangered Species. 77 pp. Montana Bird Distribution Committee. 2012. P.D. Skaar's Montana bird distribution. 7th Edition. Montana Audubon, Helena, Montana. 208 pp. + foldout map. MT Fish, Wildlife & Parks. No date. Blackfoot-Clearwater Wildlife Management Area checklist. Oechsli, L.M. 2000. Ex-urban development in the Rocky Mountain West: consequences for native vegetation, wildlife diversity, and land-use planning in Big Sky, Montana. M.Sc. Thesis. Montana State University, Bozeman. 73 p. Pitkin, P. and L. Quattrini. 2017. Pocket Guide to Sagebrush Birds. Bird Conservancy of the Rockies and Point Blue Conservation Science. 68 p. Porter, D.K. and R.A. Ryder. 1972. Avian density and productivity studies and analyses on the Pawnee Site in 1972. Fort Collins, CO: Colorado State University. Unpublished report, Department of Fishery and Wildlife Biology. 77 p. Prescott, D.R.C. and D.M. Collister. 1993. Characteristics of occupied and unoccupied loggerhead shrike territories in southeastern Alberta. J. Wildl. Manage. 57: 346-352. Rauscher, R. L. 1999. Loggerhead Shrikes in Montana. Montana Department of Fish, Wildlife and Parks, Bozeman. Reichel, J. D. 1996. Preliminary colonial nesting bird survey on the Bureau of Land Management Lewistown District: 1995. Unpublished report, U.S. Bureau of Land Management, Lewistown, Montana. Reichel, J.D. 1997. Amphibian and reptile survey in northcentral Montana on the Lewistown District, BLM. Unpublished report. Robbins, C.S., D. Bystrak, and P.H. Geissler. 1986. The breeding bird survey: its first fifteen years, 1965-1979. U. S. Fish Wildl. Serv. Resour. Publ. Rogers, R.R and A. Rogers. 1995. A survey of nesting ferruginous hawks (Buteo regalis) on public lands in Petroleum and Fergus counties, Montana. Prepared for the USDI Bureau of Land Management, Judith Resource Area. 17 p. Rundquist, V.M. 1973. Avian ecology on stock ponds in two vegetational types in north-central Montana. Ph.D. Dissertation. Bozeman, MT: Montana State University. 112 p. Ryder, R.A. 1972. Avian population studies on the Pawnee site, 1968-1971. Fort Collins, CO: Colorado State University. Grassland Biome, U.S. International Biological Program, Tech. Rep. 57 p. Saab, V. and C. Groves. 1992. Idaho's migratory landbirds, description, habitats & conservation. Nongame Wildlife Leaflet #10. Idaho Department of Fish and Game, UBLM, USFS, USFWS, National Fish and Wildlife Foundation. 16 p. Sater, S. 2022. The insects of Sevenmile Creek, a pictorial guide to their diversity and ecology. Undergraduate Thesis. Helena, MT: Carroll College. 242 p. Saunders, A.A. 1914. The birds of Teton and northern Lewis & Clark counties, Montana. Condor 16:124-144. Sibley, D. 2014. The Sibley guide to birds. Alfred A. Knopf, New York, NY. 598 pp. Skaar, P. D., D. L. Flath, and L. S. Thompson. 1985. Montana bird distribution. Montana Academy of Sciences Monograph 3(44): ii-69. Skaar, P.D. 1969. Birds of the Bozeman latilong: a compilation of data concerning the birds which occur between 45 and 46 N. latitude and 111 and 112 W. longitude, with current lists for Idaho, Montana, Wyoming, impinging Montana counties and Yellowstone National Park. Bozeman, MT. 132 p. Stearns-Roger Inc., 1975, Environmental baseline information of the Mount Vernon Region, Montana. January 31, 1975. Stewart, R.E. 1975. Breeding birds of North Dakota. Tri-College Center for Environmental Studies, Fargo, North Dakota. 295 pp. Stewart, R.E. and H.A. Kantrud. 1972. Population estimates of breeding birds in North Dakota. The Auk 89(4):766-788. Telfer, E.S. 1993. Habitat change as a factor in the decline of the western Canadian loggerhead shrike, Lanius ludovicianus, population. Can. Field-Nat. 106: 321-326. Thompson, L.S. 1981. Circle West wildlife monitoring study: Third annual report. Technical report No. 8. Montana Department of Natural Resources and Conservation. Helena, Montana. Thompson, Richard W., Western Resource Dev. Corp., Boulder, CO., 1996, Wildlife baseline report for the Montana [Montanore] Project, Lincoln and Sanders counties, Montana. In Application for a Hard Rock Operating Permit and Proposed Plan of Operation, Montanore Project, Lincoln and Sanders Counties, Montana. Vol. 5. Stroiazzo, John. Noranda Minerals Corp., Libby, MT. Revised September 1996. U.S. Fish and Wildlife Service, Office of Migratory Bird Management. 1995. Migratory nongame birds of management concern in the United States: the 1995 list. U.S. Government Printing Office: 1996-404-911/44014. 22 pp. U.S. Forest Service. 1991. Forest and rangeland birds of the United States: Natural history and habitat use. U.S. Department of Agriculture, Forest Service Agricultural Handbook 688. 625 pages. Van Lanen, N.J., A.P. Monroe, and C.L. Aldridge. 2023. Living on the edge: Predicting songbird response to management and environmental changes across an ecotone. Ecology and Evolution 13:e10648. https://doi.org/10.1002/ece3.10648 Verner, J., and L.V. Ritter. 1985. A comparison of transects and point counts in oak-pine woodlands of California. The Condor 87:47-68. Vold, S.T. 2018. Effects of livestock grazing management on the ecology of grassland birds and their predators in a northern mixed-grass prairie ecosystem. M.Sc. Thesis. Bozeman, MT: Montana State University. 176 p. 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Ecological Monographs 43:237-270. Woods, C.P. 1993. Variation in loggerhead shrike nest composition between two shrub species in southwest Idaho. J. Field Ornithol. 64: 352-357. Yosef, R. and T.C. Grubb. 1994. Resource dependence and territory size in loggerhead shrikes. Auk 111: 465-469. Yosef, Reuven. 1996. Loggerhead Shrike (Lanius ludovicianus ). Species Account Number 231. The Birds of North America Online (A. Poole, Ed.). Ithaca, NY: Cornell Laboratory of Ornithology; Retrieved 3/25/2008 from The Birds of North America Online database
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