Snapping Turtle - Chelydra serpentina
The Snapping Turtle shell is hard and very serrated ("saw-toothed") on the rear edge of the carapace; the plastron is relatively small. The tail is as long or longer than the carapace, with a crest of large, bony scales. The head is large, with a hooked upper jaw and two barbels on the chin. The limbs are strong, with webbed toes and robust claws. Skin and shell color is black to tan. Maximum carapace length is 50 centimeters, but usually is less than 36 centimeters; average weight of wild individuals is 16 kilograms. In mature males, the anal opening extends farther beyond the base of the tail than in females, and is usually beyond the rear margin of the carapace (under the rear edge in females). In adults, the carapace is relatively smooth, and the longitudinal ridges are not prominent. In juveniles, there are three longitudinal ridges on the carapace; in hatchlings the carapace is rough with conspicuous ridges. Eggs are moderately pliable, somewhat brittle, and average 28 by 27 millimeters.
The Snapping Turtle differs from other Montana turtles by the presence of a plastron (ventral shell) that is reduced to a cross-like structure, covering perhaps only half of the ventral surface; the presence of keeled scutes or scales on the carapace (dorsal shell); and the presence of a tail at least as long as the carapace. The shell is hard, not soft and leathery, nor is it flattened or pancake-like, as is the case with the Spiny Softshell.
Western Hemisphere Range
Summary of Observations Submitted for Montana
Number of Observations:
(Click on the following maps and charts to see full sized version)
Map Help and Descriptions
(Observations spanning multiple months or years are excluded from time charts)
No specific information for Montana is available.
Research from other locations indicates that the Snapping Turtle may migrate up to several miles between the permanent water of usual residence and nesting areas. Some may travel up to a few kilometers between summer range and winter hibernation sites; others hibernate within summer range (Brown and Brooks 1994). In Ontario, the maximum distance traveled from the nesting site was 370 to 2020 meters (mean 1053 meters); movements were greatest from spring to mid-July (Pettit et al. 1995).
Habitat use by Snapping Turtles in Montana is probably similar to elsewhere in the range, but studies are lacking and there is little qualitative information available. They have been captured or observed in backwaters along major rivers, at smaller reservoirs, and in smaller streams and creeks with permanent flowing water and sandy or muddy bottoms (Reichel 1995, Hendricks and Reichel 1996). Nesting habitat and nest sites have not been described.
Elsewhere, Snapping Turtles occur in all types of shallow freshwater habitats, such as streams, rivers, reservoirs, and ponds, especially those with a soft mud bottom and abundant aquatic vegetation or submerged brush and logs (Hammerson 1999), and in brackish water in some areas. Although found most often in shallower water, they have been reported on the bottom of lakes in water up to 10 meters deep. Temporary ponds may also be occupied. Hatchlings and juveniles tend to occupy shallower sites than mature individuals in the same water bodies. They are mostly bottom dwellers, where they spend much of their time. Although highly aquatic, they may make long movements overland if their pond or marsh dries (Baxter and Stone 1985, Ernest et al. 1994, Hammerson 1999). They hibernate singly or in groups in streams, lakes, ponds, or marshes; in bottom mud, in or under submerged logs or debris, under an overhanging bank, or in Muskrat tunnels; often in shallow water; sometimes in anoxic sites (Brown and Brooks 1994). Sometimes they bask out of water, especially younger individuals and in the northern extremes of the global range.
Nests are built in soft sand, loam, vegetation debris, or even sawdust piles, most often in open areas and often a hundred meters or more from water (Congdon et al. 1987, Ernst et al. 1994, Hammerson 1999); they also nest in Beaver and Muskrat lodges.
Ecological Systems Associated with this Species
- Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (high, medium, or low) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
- Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2001, Adams 2003, and Werner et al. 2004);
- Evaluating structural characteristics and distribution of each ecological system relative to the species’ range and habitat requirements;
- Examining the observation records for each species in the state-wide point database associated with each ecological system;
- Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of “observations versus availability of habitat”.
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were associated as using an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if
point observations were associated with that system.
High, medium, and low habitat quality was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species in the literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignments of habitat quality.
If you have any questions or comments on species associations with ecological systems, please contact Bryce Maxell at email@example.com
or (406) 444-3655.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: http://mtnhp.org/requests/default.asp
) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species’ known geographic range.
- Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
- Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
- Foresman, K.R. 2001. The wild mammals of Montana. Special Publication No. 12. Lawrence, KS: The American Society of Mammalogists. 278 p.
- Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
- Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
- Maxell, B.A. 2000. Management of Montana’s amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
- Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
Snapping Turtle diets have not been studied in Montana, but they are known to eat about anything that can be captured while foraging in the water. They eat many kinds of vertebrates (fish, amphibians, reptiles, aquatic birds, small mammals), invertebrates (insects, spiders, crustaceans, mollusks, leeches, sponges), algae and various aquatic vascular species (Ernst et al. 1994). Carrion is also consumed. Young Snapping Turtles actively forage for food, but adults often lie in ambush to seize their prey. This species is known to eat nine orders of insects (ants, beetles, and moths the most abundant), and spiders, scorpions, ticks, and mites have been reported in the diet (Hammerson 1999). Adults sometimes eat hatchling lizards.
Snapping Turtles are most active at night in the southern parts of the range, but this is not so at northern locations, where activity is more pronounced in the morning (Ernest et al. 1994). They may be active year round in the southern United States, but cold weather there will drive them into dormancy. Most Snapping Turtles in more northern populations do not emerge until April, and enter hibernation by late October. In Colorado, Snapping Turtles emerge from hibernation in March and remain active through October (Hammerson 1999); there they are active day or night. The period of activity in Montana is poorly documented, with records mostly from early July through September (Reichel 1995, Hendricks and Reichel 1996); active individuals have been seen during mid-day in small streams.
In Ontario, males occupied relatively stable, overlapping home ranges; summer ranges were 0.4 to 2.3 hectares (Galbraith et al. 1987). Also in Ontario, July to August foraging home ranges in three sites during one year were 2.3 to 18.1 hectares (means fell between 5 and 9 hectares); home range length was about 550 to 1990 meters; home range size did not vary with habitat productivity (Brown et al. 1994). In another Ontario study, home range size over a year was 1.0 to 28.4 hectares, averaging about 9 hectares for females and about 2 to 3 hectares for males (Pettit et al. 1995).
Snapping Turtles frequently incur high rates of nest predation (30 to 100% in Michigan) by various predators, especially skunks, Raccoon, foxes, bears, crows, and snakes (Congdon et al. 1987, Ernst et al. 1994, Hammerson 1999). Coyotes, Northern River Otters, bears, and often humans prey on adults; herons and bitterns, large hawks, eagles, various predatory fish, and American Bullfrogs prey on hatchlings and juveniles. Predators in Montana, other than humans have not been reported. See Iverson (1991) for a compilation of survivorship data (egg survival low, not more than 0.22; adult survival generally high, over 0.90). A population in Ontario, Canada, was characterized as stable, with adult female annual survivorship greater than 0.95; later, a great increase in adult mortality occurred, apparently due primarily to Northern River Otter predation on hibernating Snapping Turtles; there was no compensatory density-dependent response in reproduction and recruitment (Brooks et al. 1991). In Michigan, actual annual survivorship of juveniles was over 0.65 by age 2 and averaged 0.77 between ages 2 and 12 years; annual survivorship of adult females ranged from 0.88 to 0.97; population stability was most sensitive to changes in adult or juvenile survival and less sensitive to changes in age at sexual maturity, nest survival, or fecundity (Congdon et al. 1994).
In eastern Ontario 42 of 257 clutches emerged in fall. Of those overwintering, one survived. The species may be limited in range in the north by overwintering mortality (Obbard and Brooks 1981). Densities in marshes in South Dakota reached one per 2 acres. Predators destroyed 59% of nests. Emergence in undisturbed nests was less than 20% (Hammer 1969).
The youngest mature female known from Michigan was 12 years. In Ontario the average age of first nesting ranged from 17 to 19 years. Virtually no reproductive data exists specific to Montana; however, Montana populations likely exhibit traits similar to those elsewhere. Warming temperatures trigger nesting behavior in females. Obbard and Brooks (1987) developed a model to predict the onset of nesting activity from temperature data in Ontario. Females may travel several kilometers to locate a suitable nest site. Obbard and Brooks (1980) reported a round trip distance of 16 km to a nest site and back (Ernst et al. 1994). Nests are usually built in open areas a hundred meters or more from water; excavated in soft sand, loam, vegetation debris, sawdust piles, and Beaver and Muskrat lodges. Females generally dig nests from 7 to 18 cm (Congdon et al. 1987, Ernst et al. 1994, Hammerson 1999). In northern regions, eggs are generally deposited in late May to early June. Recorded clutch sizes range from 6 to 104, but typically 20 to 40 eggs are laid. Clutch size tends to increase from southerly to northerly latitudes. Eggs incubate for 55 to 125 days (usually 75 to 95) before hatching, with incubation period increasing with latitude. The sex of each hatchling is determined by each egg’s temperature during a critical developmental period (temperature dependent sex determination). Relatively cooler temperatures produce females and warmer temperatures produce males. The sex ratio of hatchlings is commonly 1:1 (Ernst et al. 1994). Nest site selection is critical for survival of hatchlings. Eggs burried in moist substrate generally support better embryonic development and larger hatchlings, than in drier environments (Morris et al. 1983). The incubation environment of eggs can later effect growth and viability of young Snapping Turtles (McKnight and Gutzke (1993). Bobyn and Brooks (1994) suggest incubation temperature and moisture limit the northern distribution of the Snapping Turtle.
Montana populations of Snapping Turtle are poorly understood, making management of them more difficult. It is likely that dams and large reservoirs on rivers (e.g. Fort Peck Dam and Reservoir) are detrimental to population continuity to some degree, judging by the apparent lack of viable populations on the Missouri River in Montana (Maxell et al. 2003), although this species can travel overland and may be able to bypass some dams. Impacts of other habitat disturbances are not clear, but this species occupies man-made water bodies throughout its range that provide necessary resources and habitat characteristics. Studies of nesting success, population structure, dispersal, and population size need to be conducted throughout the range of the Snapping Turtle in Montana. Routine surveys for Snapping Turtles in appropriate habitats could be made a standard part of the field duties of agency fishery biologists. Records should be maintained of the incidental "take" by anglers, who should be encouraged to report any captured individuals; killed animals should be examined by agency fishery or wildlife biologists if possible so that data on sex, size, and food habits can be gathered and a base of information developed on the biology of this species in Montana. It is possible that even moderate harvest of adults by anglers in most localities will result in population declines, similar to Colorado (Hammerson 1999), because the life history of this species indicates recruitment of juveniles into breeding populations is low, and population densities in western states is probably low. Identified nesting sites should be monitored and protected from disturbance by humans.
- Literature Cited AboveLegend: View Online Publication
- Baxter, G.T. and M.D. Stone. 1985. Amphibians and reptiles of Wyoming. Second edition. Wyoming Game and Fish Department. Cheyenne, WY. 137 p.
- Bobyn, M.L. and R.J. Brooks. 1994. Incubation conditions as potential factors limiting the northern distribution of snapping turtles, Chelydra serpentina. Canadian Journal of Zoology 72(1): 28-37.
- Brooks, R. J., G. P. Brown, and D. A. Galbraith. 1991. Effects of a sudden increase in natural mortality of adults on a population of the common snapping turtle (Chelydra serpentina). Canadian Journal of Zoology 69: 1314-1320.
- Brown, G. P. and R. J. Brooks. 1994. Characteristics of and fidelity to hibernacula in a northern population of snapping turtles, Chelydra serpentina. Copeia 1: 222-226.
- Brown, G.P., C.A. Bishop, and R.J. Brooks. 1994. Growth rate, reproductive output, and temperature selection of snapping turtles in habitats of different productivities. Journal of Herpetology 28(4): 405-410.
- Congdon, J .D., A. E. Dunham and R. C. Van Loben Sels. 1994. Demographics of common snapping turtles (Chelydra serpentina): implications for conservation and management of long-lived organisms. American Zoologist 34: 397-408.
- Congdon, J. D., G. L. Breitenbach, R. C. Van Loben Sels, and D. W. Tinkle. 1987. Reproduction and nesting ecology of snapping turtles (Chelydra serpentina) in southeastern Michigan (USA). Herpetologica 43(1): 39-54.
- Ernst, C. H., R. W. Barbour, and J. E. Lovich. 1994. Turtles of the United States and Canada. Smithsonian Institution Press. Washington, D.C. 578 p.
- Galbraith, D. A., M. W. Chandler, and R. J. Brooks. 1987. The fine structure of home ranges of male Chelydra serpentina: are snapping turtles territorial? Canadian Journal of Zoology 65(11): 2623-2629.
- Hammer, D. A. 1969. Parameters of a marsh snapping turtle population, Lacreek Refuge, South Dakota. Journal of Wildlife Management 33(4):995-1005.
- Hammerson, G.A. 1999. Amphibians and reptiles in Colorado. University Press of Colorado & Colorado Division of Wildlife. Denver, CO. 484 p.
- Hendricks, P. and J.D. Reichel. 1996. Preliminary amphibian and reptile survey of the Ashland District, Custer National Forest: 1995. Montana Natural Heritage Program, Helena, MT. 79 p.
- Iverson, J.B. 1991. Patterns of survivorship in turtles (order Testudines). Canadian Journal of Zoology 69: 385-391.
- Maxell, B.A., J.K. Werner, P. Hendricks, and D.L. Flath. 2003. Herpetology in Montana: a history, status summary, checklists, dichotomous keys, accounts for native, potentially native, and exotic species, and indexed bibliography. Society for Northwestern Vertebrate Biology, Northwest Fauna Number 5. Olympia, WA. 135 p.
- McKnight, C.M. and W.H.N. Gutzke. 1993. Effects of the embryonic environment and of hatchling housing conditions on growth of young snapping turtles (Chelydra serpentina). Copeia (2): 475-482.
- Morris, K.A., G.C. Packard, T.J. Boardman, G.L. Paukstis, and M.J. Packard. 1983. Effect of the hydric environment on growth of embryonic snapping turtles (Chelydra serpentina). Herpetologica 39: 272-285.
- Obbard, M.E. and R.J. Brooks. 1980. Nesting migrations of the snapping turtle (Chelydra serpentina). Herpetologica 36: 158-162.
- Obbard, M.E. and R.J. Brooks. 1981. Fate of overwintered clutches of the common snapping turtle (Chelydra serpentina) in Algonquin Park, Ontario, Canada. Canadian Field Naturalist 95:350-352.
- Obbard, M.E. and R.J. Brooks. 1987. Prediction of the onset of the annual nesting season of the common snapping turtle, Chelydra serpentina. Herpetologica 43: 324-328.
- Pettit, K. E., C. A. Bishop, and R. J. Brooks. 1995. Home range and movements of the common snapping turtle, Chelydra serpentina serpentina, in a coastal wetland of Hamilton Harbour, Lake Ontario, Canada. Canadian Field Naturalist 109(2): 192-200.
- Reichel, J.D. 1995. Preliminary amphibian and reptile survey of the Sioux District of the Custer National Forest: 1994. Montana Natural Heritage Program, Helena, MT. 75 p.
- Additional ReferencesLegend: View Online Publication
Do you know of a citation we're missing?
- [DCC] Decker Coal Company. 1998. 1997 Consolidated annual progress report. Decker Coal Company West, North and East Pits. Decker, MT.
- [OEA] Olson Elliot and Associates Research. 1985. 1983-1984 Wildlife monitoring report for the CX Ranch project. Olson Elliot and Associates Research. Helena, MT.
- [VTNWI] VTN Wyoming Incorporated. No Date. Second year's analysis of terrestrial wildlife on proposed mine access and railroad routes in southern Montana and northern Wyoming, March 1979 - February 1980. VTN Wyoming Incorporated. Sheridan, WY. 62 p.
- [WESTECH] Western Technology and Engineering Incorporated. 1998. Wildlife Monitoring Absaloka Mine Area 1997. Western Technology and Engineering, Inc., Helena, Mt.
- Abel, B. 1992. Snapping turtle attacks on trumpeter swan cygnets in Wisconsin. Passenger Pigeon 54(3): 209-213.
- Aderhold, M. 1980. The 32-pound snapper of Redwater River. Montana Outdoors 11(4): 9-10.
- Albers, P.H., L. Sileo, and B.M. Mulhem. 1986. Effects of environmental contaminants on snapping turtles of a tidal wetland. Archives of Environmental Contamination and Toxicology 15(1): 39-49.
- Anderson, J.T. and A.M. Anderson. 1996. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 27(3): 150.
- Anderson, J.T. and A.M. Anderson. 1996. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 27(3): 150.
- Applegate, R.D., R.C. Spencer, and F.M. Trasko. 1995. Extensions to the known range for three Maine reptiles. Canadian Field Naturalist 109(4): 444-446.
- Asch, R.P. 1951. A preliminary report on the size, egg number, incubation period, and hatching in the common snapping turtle, Chelydra serpentina. Virginia Academy of Science 29: 312.
- Ashley, E.P., and J.T. Robinson. 1996. Road mortality of amphibians, reptiles and other wildlife on the Long Point causeway, Lake Erie, Ontario. Canadian Field Naturalist 110: 403-412.
- Association for the Study of Reptilia and Amphibia. 1996. Revenge of the ninja turtles. Rephiberary 228:9.
- Atkinson, E.C. and M.L. Atkinson. 2004. Amphibian and reptile survey of the Ashland and Sioux of the Custer National Forest with special emphasis on the Three-Mile Stewardship Area:2002. Marmot's Edge Conservation. 22 p.
- Avery, H.W. and L.J. Vitt. 1984. How to get blood from a turtle. Copeia 1984: 209-210.
- Baker, M.R. 1986. Falcaustra sp. (Nematoda: Kathlaniidae) parasitic in turtles and frogs in Ontario (Canada). Canadian Journal of Zoology 64(1): 228-237.
- Baldwin, F.M. 1925b. The relation of body to environmental temperature in turtles, Chrysemys marginata belli (Gray) and Chelydra serpentina (Linn.). Biology Bulletin 48: 432-445.
- Bickham, J.W. and J.L. Carr. 1983. Taxonomy and phylogeny of the higher categories of cruptodiran turtles based on a cladistic analysis of chromosomal data. Copeia 1983:918-932.
- Bishop, C.A., R.J. Brooks, J.H. Carey, P. Ng, R.J. Norstrom, and D.R.S. Lean. 1991. The case for a cause-effect linkage between environmental contamination and development in eggs of the common snapping turtle (Chelydra serpentina serpentina) from Ontario, Canada. Journal of Toxicology and Environmental Health 33(4): 521-548.
- Black, J.H. and J.N. Black. 1971. Montana and its turtles. International Turtle and Tortoise Society 1971(May-July): 10-11, 34-35.
- Bleakney, S. 1957. A snapping turtle, Chelydra serpentina serpentina, containing eighty-three eggs. Copeia 1957: 143.
- BLM. 1982b. Moorhead baseline inventory - wildlife. Bureau of Land Management, Miles City District Office. Miles City, MT. 29 pp.
- Bobyn, M.L. and R.J. Brooks. 1994. Interclutch and interpopulation variation in the effects of incubation conditions on sex, survival and growth of hatchling turtles (Chelydra serpentina). Journal of Zoology 233(2): 233-257.
- Boice, R., Q.C. Boice, and R.C. Williams. 1974. Competition and possible dominance in turtles, toads and frogs. Journal of Comparative Physiology and Psychology 86:1116-1131.
- Bolek, M.G. 2001. Chelydra serpentina (common snapping turtle) and Emydoidea blandingii (Blanding's turtle) parasites. Herpetological Review 32(1): 37-38.
- Bonin, J., J.L. DesGranges, C.A. Bishop, J. Rogrigue, A. Gendron, J.E. Elliott. 1995. Comparative study of contaminants in the mudpuppy (Amphibia) and the common snapping turtle (Reptilia), St. Lawrence River, Canada. Archives of Environmental Contamination and Toxicology 28(2): 184-194.
- Boulenger, G.A. 1902. On the southern snapping turtle (C. rossignonii). Annals and Magazine of Natural History (7)9(69): 49-51.
- Bramblett, Robert G., and Alexander V. Zale. 2002. Montana Prairie Riparian Native Species Report. Montana Cooperative Fishery Research Unit, Montana State University - Bozeman.
- Brittle, D. 1991. Chelydra serpentina (snapping turtle). Catesbeiana 11(2): 39.
- Brooks, R. J., D. A Galbraith, E. G. Nancekivell, and C. A. Bishop. 1988. Developing management guidelines for snapping turtles. In: R.C. Szaro, K.E. Severson, and D.R. Patton, technical coordinators. pp. 174-179. Management of amphibians, reptiles, and small mammals in North America. General Technical Report RM-166. U.S. Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
- Brooks, R.J., D.A. Galbraith, and J.A. Layfield. 1990. Occurrence of Placobdella parasitica (Hirudinea) on snapping turtles, Chelydra serpentina, in southeastern Ontario. Journal of Parasitology 76(2): 190-195.
- Brooks, R.J., D.A. Galbraith, and m.L. Bobyn. 1989. Intraspecific variation in measures of life history in the common snapping turtle, Chelydra serpentina. In Proceedings of the First World Congfess of Herpetology. University of Kent, Canterbury, U.K., 11-19 September 1989. The Durrell Institute. Canterbury, Kent, U.K. p. 42.
- Brooks, R.J., D.A. Galbraith, E.G. Nancekivell and C.A. Bishop. 1988. Developing management guidelines for snapping turtles. Pages 174-179 in R.C. Szaro, K.E. Severson, and D.R. Patton, technical coordinators. Management of amphibians, reptiles, and small mammals in North America. General Technical Report RM-166. U.S. Forest Service, Rocky Mountain Forest and Range Experiment Station, Fort Collins, Colorado.
- Brooks, R.J., G.P. Brown and D.A. Galbraith. 1991. Effects of a sudden increase in natural mortality of adults on a population of the common snapping turtle (Chelydra serpentina). Canadian Journal of Zoology 69: 1314-1320.
- Brooks, R.J., M.L. Bobyn, D.A. Galbraith, J.A. Layfield, and E.G. Nancekivell. 1991. Maternal and environmental influences on growth and survival of embryonic and hatchling snapping turtles (Chelydra serpentina). Canadian Journal of Zoology 69: 2667-2676.
- Brown, G.P. 1992. Thermal and spatial ecology of a northern population of snapping turtles, Chelydra serpentina. M.S. Thesis, University of Guelph. Guelph, Ontario.
- Brown, G.P. and R.J. Brooks. 1991. Thermal and behavioural responses to feeding in free-ranging turtles, Chelydra serpentina. Journal of Herpetology 25(3): 273-278.
- Brown, G.P. and R.J. Brooks. 1993. Sexual and seasonal differences in activity in northern population of snapping turtles, Chelydra serpentina. Herpertologica 49(3): 311-318.
- Brown, G.P., R.J. Brooks, and J.A. Layfield. 1990. Radiotelemetry of body temperatures of free-ranging snapping turtles (Chelydra serpentina) during summer. Canadian Journal of Zoology 68(8): 1689-1663.
- Brown, G.P., R.J. Brooks, E. Siddall, and S.S. Desser. 1994. Parasites and reproductive output in the snapping turtle, Chelydra serpentina. Copeia 1994(1): 228-231.
- Brunson, R.B. 1955. Check list of the amphibians and reptiles of Montana. Proceedings of the Montana Academy of Sciences 15: 27-29.
- Budhabhatti, J., and E.O. Moll. 1990. Chelydra serpentina (common snapping turtle). Feeding behavior. Herpetological Review 21(1): 19.
- Bureau of Indian Affairs, Department of the Interior. 1981. Draft Environmental Impact Statement. Unpublished report for the Crow/Shell Coal Lease, Crow Indian Reservation, Montana.
- Burke, A.C. 1989. Development of the turtle carapace: implications for the evolution of a novel Bauplan. Journal of Morphology 199(3): .363-378.
- Burke, A.C., and P. Alberch. 1985. The development and homology of the chelonian carpus and tarsus. Journal of Morphology 186(1): 119-131.
- Burke, V.J, R.D. Nagle, M. Ostentoski, and J.D. Congdon. 1993. Common snapping turtles associated with ant mounds. Journal of Herpetology 27: 114-115.
- Burkholder, G.L. and W.W. Tanner. 1974. Life history and ecology of the Great Basin sagebrush swift, Sceloporus graciosus graciosus Baird and Girard, 1852. BYU Science Bulletin Biological Series 14(5): 1-42.
- Cagle, F.R. 1944a. A Technique for obtaining turtle eggs for study. Copeia 1944: 60.
- Cagle, F.R. 1948. The growth of turtles in Lake Glendale, Illinois. Copeia 1948: 197-203.
- Carlson, J. (Coordinator, Montana Animal Species of Concern Committee). 2003. Montana Animal Species of Concern. Helena, MT: Montana Natural Heritage Program and Montana Fish, Wildlife, and Parks. In Press. 12p.
- Carr, A.F. 1952. Handbook of turltles: the turtles of the United States, Canada, and Baja California. Cornell University Press, Ithaca, New York.
- Censky, E.J. 1986. Sceloporus graciosus Baird and Girard. sagebrush lizard. Catalogue of American Amphibians and Reptiles 386:1-4.
- Chiszar, D. and H.M. Smith. 1995. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 26(4): 208.
- Chiszar, D., R.L. Holland, C. Ristau, and H.M. Smith. 1994. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 25(4): 162.
- Chiszar, D., R.L. Holland, C. Ristau, and H.M. Smith. 1994. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 25(4): 162.
- Christiansen, J.L. and R.R. Burken. 1979. Growth and maturity of the snapping turtle (Chelydra serpentina) in Iowa. Herpetologica 35: 261-266.
- Cink, C.L. 1991. Snake predation on nestling eastern phoebes followed by turtle predation on snake. Kansas Ornithological Society Bulletin 42(3): 29.
- Coaker, M. 1993. The common snapping turtle (Chelydra serpentina) of North America. Southwestern Herpetologists Society Journal 2(3): 33-34.
- Collins, J.T. 1991. A new taxonomic arrangement for some North American amphibians and reptiles. Herpetological Review 22:42-43.
- Collins, J.T. 1997. Standard common and current scientific names for North American amphibians and reptiles. Fourth edition. Society for the Study of Amphibians and Reptiles. Herpetological Circular No. 25. 40 pp.
- Congdon, J.D. and J.W. Gibbons. 1990. The evolution of turtle life histories. Pages 45-54 in Life history and ecology of the slider turtle. Edited by J.W. Gibbons. Smithsonian Institution Press, Washington, D.C., U.S.A.
- Congdon, J.D., S.W. Gotte, S.W., and R.W. Mcdiarmid. 1992. Ontogenetic changes in habitat use by juvenile turtles, Chelydra serpentina and Chrysemys picta. Canadian Field Naturalist 106(2): 241-248.
- Cope, E.D. 1875. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. U.S. Natioanl Museum Bulletin 1: 1-104.
- Costanzo, J.P., J.B. Iverson, M.F. Wright, and R.E. Lee, Jr. 1995. Cold hardiness and overwintering strategies of hatchlings in an assemblage of northern turtles. Ecology 76(6): 1772-1785.
- Coues, E. and H. Yarrow. 1878. Notes on the herpetology of Dakota and Montana. Bulletin of the U.S. Geological Geographic Survey of the Territories 4: 259-291.
- Crother, B.I. (ed.) 2008. Scientific and standard English names of amphibians and reptiles of North America north of Mexico. SSAR Herpetological Circular No. 37:1-84.
- Cunnington, D.C. and R.J. Brooks. 1996. Bet-hedging theory and eigenelasticity: a comparison of the life histories of loggerhead sea turtles (Caretta caretta) and snapping turtles (Chelydra serpentina). Canadian Journal of Zoology 74(2): 291-296.
- Curtis, S. 1994. The big sleep. Montana Outdoors 25(6): 2-7.
- Daigle, C., A. Desrosiers, and J. Bonin. 1994. Distribution and abundance of common map Turtles, Graptemys geographica, in the Ottawa River, Quebec. Canadian Field Naturalist 108(1): 84-86.
- Darrow, T.D. 1961. Food habits of western painted and snapping turtles in southeastern South Dakota and eastern Nebraska. Unpubl. MS Thesis, University of South Dakota, Vermillion.
- Day, D. 1989. Montco Terrestrial Wildlife Monitoring Report. Unpublished report for Montco, Billings, Montana.
- Day, D., P.J. Farmer, and C.E. Farmer. 1989. Montco terrestrial wildlife monitoring report December, 1987 - July, 1989. Montco, Billings, MT, and Western Technology and Engineering, Inc. Helena, MT.
- de Santis, B., B. Sheafor, H.M. Smith, and D. Chiszar. 1995. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 26(3): 154.
- Decker, J.D. 1967. Motility of the turtle embryo, Chelydra serpentina (Linne.). Science 157: 952-954.
- Degenhardt, W. G., C. W. Painter, and A. H. Price. 1996. Amphibians and reptiles of New Mexico. University of New Mexico Press, Albuquerque.
- Dinkelacker, S.A., J.P. Costanzo, and R.E. Lee Jr. 2005. Anoxia tolerance and freeze tolerance in hatchling turtles. Journal of Comparative Physiology B Biochemical Systemic and Environmental Physiology 175(3):209-217.
- Distel, C. 2005. Chelydra serpentina serpentina (Common Snapping Turtle). Flatulence. Herpetological Review 36:310.
- Dixon, J.R., O.W. Thornton, Jr. 1996. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 27(1): 31.
- Dodd, C.K., Jr. and E.D. Brodie, Jr. 1975. Notes on the defensive behavior of the snapping turtle, Chelydra serpentina. Herpetologica 31: 286-288.
- Dunham, A.E. and K.L. Overall. 1994. Population responses to environmental change: Life history variation, individual-based models, and the population dynamics of short-lived organisms. American Zoologist 34(3): 382-396.
- Dunson, W.A. 1986. Estuarine populations of the snapping turtle (Chelydra) as a model for the evolution of marine adaptations in reptiles. Copeia 1986:741-756.
- ECON, Inc. (Ecological Consulting Service), Helena, MT., 1976, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1976. Proj. 135-85-A. December 31, 1976.
- Ewert, M.A., J.W. Lang, and C.E. Nelson. 2005. Geographic variation in the pattern of temperature-dependent sex determination in the American snapping turtle (Chelydra serpentina). Journal of Zoology (London) 265(1):81-95.
- Ewert, M.E. 1979. The embryo and its egg: development and natural history. In Turtles: perspecgtives and research. Edited by M. Harless and H. Morlock. John Wiley and Sons, Inc., New York. pp. 333-413.
- Ewert, M.E. and C.E. Nelson. 1991. Sex determination in turtles: diverse patterns and some possible adaptive values. Copeia 1991: 50-69.
- Farmer, P. 1980. Terrestrial wildlife monitoring study, Pearl area, Montana June, 1978 - May, 1980. Western Technology and Engineering, Inc. Helena, MT.
- Farmer, P. J. 1980. Terrestrial Wildlife Monitoring Study, Pearl Area, Montana, June, 1978 - May, 1980. Tech. Rep. by WESTECH for Shell Oil Co.
- Feinberg, J.A. and K. Hoffmann. 2004. Chelydra serpentina (Snapping Turtle) and Scaphiopus holbrookii (Eastern Spadefoot). Diet. Herpetological Review 35(4):380.
- Fergus, C. 1985. Out of their element, turtles bend the regulations. Oceans 18(1): 66-67.
- Feuer, R.C. 1966. Variation in snapping turtles. Chelydra serpentina Linnaeus: a study in quantitative systematics. Ph.D. Dissertation. University of Utah, Salt Lake City.
- Feuer, R.C. 1971a. Ecological factors in success and dispersal of the snapping turtle Chelydra serpentina (Linnaeus). Bulletin of the Philadelphia Herpetological Society 19: 3-14.
- Feuer, R.C. 1971b. Intergradation of the snapping turtles Chelydra serpentina serpentina (Linnaeus, 1758) and Chelydra serpentina osceola Stejneger, 1918. Herpetologica 27(4): 379-384.
- Finkler, M.S. and D.L. Claussen. 1997. Use of the tail in terrestrial locomotor activities of juvenile Chelydra serpentina. Copeia 1997: 884-887.
- Finkler, M.S. and D.L. Knickerbocker. 2000. Influence of hydric conditions during incubation and population on overland movement of neonatal snapping turtles. Journal of Herpetology 34(3): 452-455.
- Finneran, L.C. 1947. A large clutch of eggs of Chelydra serpentina serpentina (linnaeus). Herpetologica 3: 182.
- Flath, Dennis L., 1979, Nongame species of special interest or concern: Mammals, birds, reptiles, amphibians, fishes. January 1979.
- Frair, W. 1972. Taxonomic relations among chelydrid and kinosternid turtles elucidated by serological tests. Copeia 1972:97-108.
- Fraser, G. 1994. Possible predation of a Forster's tern chick by a snapping turtle. Prairie Naturalist 26(1): 33-35.
- Frazer, N.B., J.W. Gibbons, and T.J. Owens. 1990. Turtle trapping: preliminary tests of conventional wisdom. Copeia 1990(4): 1150-1152.
- Froese, A. D. and G. M. Burghardt. 1975. A dense natural population of the common snapping turtle (Chelydra serpentina). Herpetologica 31:204-208.
- Froese, A.D. 1974. Aspects of space use in the common snapping turtle, Chelydra s. serpentina. Unpubl.Ph. D. Thesis, University of Tennessee, Knoxville.
- Froese, A.D. 1978. Habitat preferences of the common snapping turtle, Chelydra s. serpentina (Reptilia, Testudines, Chelydridae). Journal of Herpetology 12: 53-58.
- Froese, A.D. and G.M. Burghardt. 1974. Food competition in captive juvenile snapping turtles, Chelydra serpentina. Animal Behavior 22: 735-740.
- Gaffney, E.S. 1975b. Phylogeny of the chelydrid turtles: a study of shared derived characters of the skull. Fieldiana, Geology 33: 157-178.
- Gaffney, E.S. 1984a. Historical analysis of theories of chelonian relationship. Systematic Zoology 33:283-301.
- Gaffney, E.S. 1984b. Progress towards a natural hierarchy of turtles. Studia Geologica Salmanticensia (Studia Palaeocheloniologica I) 1:125-131.
- Galbraith, D.A. 1994. Ecology research on snapping turtles (Chelydra serpentina) in Ontario, Canada. Asra Journal 1994: 23-49.
- Galbraith, D.A. and R.J. Brooks. 1987a. Addition of annual lines in adult snapping turtles Chelydra serpentina. Journal of Herpetology 21: 359-363.
- Galbraith, D.A. and R.J. Brooks. 1987c. Survivorship of adult females in a northern population of common snapping turtles, Chelydra serpentina. Canadian Journal of Zoology 65(7): 1581-1586.
- Galbraith, D.A. and R.J. Brooks. 1989. Age estimates for snapping turtles. Journal of Wildlife Management 53(2): 502-508.
- Galbraith, D.A., B.N. White, R.J. Brooks, and P.T. Boag. 1993. Multiple paternity in clutches of snapping turtles (Chelydra serpentina) detected using DNA fingerprints. Canadian Journal of Zoology 71(2): 318-324.
- Galbraith, D.A., C.A. Bishop, R.J. Brooks, W.L. Simser, and K.P. Lampman. 1988. Factors affecting the density of populations of common snapping turtles (Chelydra serpentina serpentina). Canadian Journal of Zoology 66(5): 1233-1240.
- Galbraith, D.A., C.J. Graesser, and R.J. Brooks. 1988. Egg retention by a snapping turtle, Chelydra serpentina, in central Ontario (Canada). Canadian Field Naturalist 102(4): 734.
- Galbraith, D.A., R.J. Brooks, and M.E. Obbard. 1989. The influence of growth rate on age and body size at maturity in female snapping turtles (Chelydra serpentina). Copeia (4): 896-904.
- Garbin, C.P. 1992. Performance of hatchling snapping turtles (Chelydra serpentina) on single-food diets: a comparison of three commercial foods. Bulletin of the Chicago Herpetological Society 27(12): 249-251.
- Gates, M.T. 2005. Amphibian and reptile baseline survey: CX field study area Bighorn County, Montana. Report to Billings and Miles City Field Offices of Bureau of Land Management. Maxim Technologies, Billings, MT. 28pp + Appendices.
- Gatten, R.E. 1980. Aerial and aquatic oxygen uptake by freely-diving snapping turtles (Chelydra serpentina). Oecologia 46: 266-271.
- Gaunt, A.S. and C. Gans. 1969. Mechanics of respiration in the snapping turtle, Chelydra serpentina (Linne.). Journal of Morphology 128: 195-228.
- Gerholdt, J.E. and B. Oldfield. 1987. Life history notes. Chelydra serpentina serpentina (common snapping turtle). Size. Herpetological Review 18(4): 73.
- Gettinger, R.D., G.L. Paukstis, and G.C. Packard. 1984. Influence of hydric environment on oxygen consumption by embryonic turtles Chelydra serpentina and Trionyx spiniferus. Physiological Zoology 57: 468-473.
- Gibbons, J.W. 1968b. Growth rates of the common snapping turtle, Chelydra serpentina, in a polluted river. Herpetologica 24: 266-267.
- Gibbons, J.W. and D.H. Nelson. 1978. The evolutionary significance of delayed emergence from the nest by hatchling turtles. Evolution 32: 297-303.
- Gibbons, J.W., S.S. Novak, and C.H. Ernst. 1988. Chelydra serpentina (Linnaeus). Snapping turtle. Catalogue of American Amphibians and Reptiles 420: 1-4.
- Graff, H. and H. Graff. 1991. Snapping turtle (Chelydra serpentina). Newsletter of the Australian Society of Herpetologists Inc. 34(6): 131-133.
- Graham, T.E. and R.W. Perkins. 1976. Growth of the common snapping turtle, Chelydra serpentina, in a polluted marsh. Maryland Herpetological Society Bulletin 12: 123-125.
- Green, G.A., K.B. Livezey, and R.L. Morgan. 2001. Habitat selection by northern sagebrush lizards (Sceloporus graciosus graciosus) in the Columbia Basin, Oregon. Northwestern Naturalist 82:111-115.
- Gutzke, W.H.N. 1984. Modification of the hydric environment by eggs of snapping turtles (Chelydra serpentina). Canadian Journal of Zoology 62(12): 2401-2403.
- Gutzke, W.H.N. and D. Crews. 1988. Embyonic temperature determines adult sexuality in a reptile. Nature 332: 832-834.
- Haas, A. 1985. Year-long outdoor behavior of the snapping turtle Chelydra serpentina serpentina in southern Germany (Testudines, Chelydridae). Salamandra 21(1): 1-9.
- Hamilton, A.M. and A.H. Freedman. 2002. Effects of deer feeders, habitat and sensory cues on predation rates on artificial turtle nests. American Midland Naturalist 147:123-134.
- Hamilton, W.J., Jr. 1940. Observations on the reproductive behavior of the snapping turtle. Copeia 1940: 124-126.
- Hammer, D.A. 1971a. Reproductive behavior of the common snapping turtle. Canadian Herpetological Society 1: 9-13.
- Hammer, D.A. 1971b. The durable snapping turtle. Natural History 80: 59-65.
- Hanson, J. No Date. Unpublished report on Youngs Creek. U.S. Fish and Wildlife Service. Hardin, MT. 14 p.
- Haxton, T. 2000. Road mortality of snapping turtles, Chelydra serpentina, in central Ontario during their nesting period. The Canadian Field Naturalist 114(1): 106.
- Hebert, C.E., V. Glooshenko, G.D. Haffner, and R. Lazar. 1993. Organic contaminants in snapping turtle (Chelydra serpentina) populations from southern Ontario, Canada. Archives of Environmental Contamination and Toxicology 24(1): 35-43.
- Hemphill, N. and W. Worthey. 1994. Do predaceous turtles affect stream fishes? International Vereinigung Fuer Theoretische Und Angewandte Limnologie Verhandlungen 25(4): 2105-2107.
- Hendricks, P. 1999. Amphibian and reptile survey of the Bureau of Land Management Miles City District, Montana. Montana Natural Heritage Program, Helena, MT. 80 p.
- Hendricks, P. and L.N. Hendricks. 2002. Predatory attack by green-tailed towhee on sagebrush lizard. Northwestern Naturalist 83:57-59.
- Hogg, D.M. 1975. The snapping turtles of Wye Marsh. Ontario Fish and Wildlife Review 14: 16-20.
- Holman, A.J. 1988. The status of Michigan's Pleistocene herpetofauna. Michigan Academician 20(2): 125-132.
- Holman, J.A. and J.N. Mcdonald. 1986. A late quaternary herpetofauna from Saltville, Virginia (USA). Brimleyana 0(12): 85-100.
- Hotaling, E.C., D.C. Wilhoft, and S.B. McDowell. 1985. Egg position and weight of hatchling snapping turtles, Chelydra serpentina, in natural nests. Journal of Herpetology 19(4): 534-536.
- Hoyt, J.S.Y. 1941. The incubation period of the snapping turtle. Copeia 1941: 180.
- Humphrey, R., D. Chiszar, and H.M. Smith. 1995. Chelydra serpentina (snapping turtle). Herpetological Review 26(2): 106.
- Iverson, J.B. 1992a. A revised checklist with distribution maps of the turtles of the world. Paust Printing, Richmond, Indiana.
- Iverson, J.B., H. Higgins, A. Sirulnik, and C. Griffiths. 1997. Local and geographic variation in the reproductive biology of the snapping turtle (Chelydra serpentina). Herpetologica 53(1): 96-117.
- Janzen, F.J. 1990. Egg size and hatching success of snapping turtle eggs: evaluation of natural selection. American Zoologist 30: 55A.
- Janzen, F.J. 1992. Heritable variation for sex ratio under environmental sex determination in the common snapping turtle (Chelydra serpentina). Genetics 131(1): 155-161.
- Janzen, F.J. 1993. An experimental analysis of natural selection on body size of hatchling turtles. Ecology 74(2): 332-341.
- Janzen, F.J. 1994b. Temperature-dependent sex determination influences the mortality of hatchling snapping turtles in nature. American Zoologist 34(5): 9A.
- Janzen, F.J. 1995. Experimental evidence for the evolutionary significance of temperature dependent sex determination. Evolution 49(5): 864-873.
- Janzen, F.J. and S. O’Steen. 1990. An instance of male combat in the common snapping turtle (Chelydra serpentina). Bulletin of the Chicago Herpetological Society 25(1): 11.
- Kinneary, J.J. 1992. The effect of water salinity on growth and oxygen consumption of snapping turtle (Chelydra serpentina) hatchlings from an estuarine habitat. Journal of Herpetology 26(4): 461-467.
- Kinneary, J.J. 1993. Salinity relations of Chelydra serpentina in a Long Island estuary. Journal of Herpetology 27(4): 441-446.
- Kiviat, E. 1980. A Hudson River tidemarsh snapping turtle population. Transactions Northeastern Section Wildlife Society 158-168.
- Kiviat, E. 1994. Consider the snapping turtle... Conservationist 48(5-6): 36-41.
- Knight, T.W., J.A. Layfield, and R.J. Brooks. 1990. Nutritional status and mean selected temperature of hatchling snapping turtles (Chelydra serpentina): is there a thermophilic response to feeding? Copeia 1990: 1067-1072.
- Knowlton, G.F. and J.S. Stanford. 1942. Reptiles eaten by birds. Copeia 1942: 186.
- Koch, E.D. and C.R. Peterson. 1995. Amphibians and reptiles of Yellowstone and Grand Teton national parks. University of Utah Press, Salt Lake City, UT. 188 p.
- Ladd, C.G. and G.G. Galbraith. 1989. Chelydra s. serpentina (common snapping turtle). Herpetological Review 20(1): 14.
- Lagler, K.F. and V.C. Applegate. 1943. Relationship between the length and the weight in the snapping turtle Chelydra serpentina Linnaeus. American Naturalist 77: 476-478.
- Lawniczak, C.J. and M.A. Teece. Spatial mobilization of calcium and magnesium from the eggshell of the snapping turtle, Chyledra serpentina. Journal of Herpetology 39(4):659-664.
- Layfield, J.A., D.A. Galbraith, and R.J. Brooks. 1988. A simple method to mark hatchling turtles. Herpetological Review 19(4): 78-79.
- Linnaeus, C. 1758. Syst. Nat., 10th ed., Vol. 1, p. 199.
- Loncke, D.J. and M.E. Obbard. 1977. Tag success, dimensions, clutch size and nesting site fidelity for the snapping turtle, Chelydra serpentina, (Reptilia, Testudines, Chelydridae) in Algongquin Park, Ontario, Canada. Journal of Herpetology 11: 243-244.
- Long, C.A., J.E. Long, and C.A. Long. 1987. A listing of known amphibians and reptiles from Washington Island, Lake Michigan, with new records of the milk snake and red-bellied snake. Jack-Pine Warbler 65(3-4): 39.
- Lovich, J.E. and J.W. Gibbons. 1996. Conservation of covert species: protecting species we don't even know. Proceedings: conservation restoration and managemen of turtles and tortoises, and international conference. State University of New York, Purchase.
- M'Closkey, R.T., S.J. Hecnar, D.R. Chalcraft, J.E. Cotter, J. Johnston, and R. Poulin. 1997. Colonization and saturation of habitats by lizards. Oikos 78:283-290.
- Mahmoud, I.Y. and R.V. Cyrus. 1992. The testicular cycle of the common snapping turtle, Chelydra serpentina, in Wisconsin. Herpetologica 48(2): 193-201.
- Manlius, N. 1996. Pectoral girdle and forelimb structure of the leatherback turtle Dermochelys coriacea Vandelli, 1761 (Chelonii, Dermochelyidae) compared with four other chelonian species. Bulletin de la Societe Herpetologique de France 1996: 1-18.
- Martin, P.R. 1980a. Terrestrial wildlife habitat inventory in southeastern Montana. Montana Department of Fish, Wildlife and Parks and Bureau of Land Management, Helena MT. 114 p.
- Martin, P.R. 1980b. Terrestrial wildlife inventory in selected coal areas of Montana. Montana Department of Fish, Wildlife and Parks and Bureau of Land Management, Helena, MT. 84 p.
- Martin, P.R., K. Dubois and H.B. Youmans. 1981. Terrestrial wildlife inventory in selected coal areas, Powder River resources area final report. Montana Department of Fish, Wildlife and Parks and Bureau of Land Management, Helena, MT. No. YA-553-CTO- 24. 288 p.
- Matthews, W.L. 1979. Wibaux-Beach wildlife baseline study - nongame species. Bureau of Land Management, Miles City, MT. 93 p.
- Mayeaux, M.H. 1996. A possible constraint to high-density culture of male common snapping turtles Chelydra serpentina at consistently warm temperatures. Journal of the World Aquaculture Society 27(1): 137-139.
- McAllister, C.T. 1988. New county records for amphibians and reptiles of north central Texas. Herpetological Review 19(3): 61-62.
- McAllister, C.T., S.J. Upton, and S.E. Trauth. 1990. Coccidian parasites (Apicomplexa: Eimeriidae) of Chelydra serpentina (Testudines: Chelydridae) from Arkansas and Texas, U.S.A., with descriptions of Isospora chelydrae sp. nov. and Eimeria serpentina sp. nov. Canadian Journal of Zoology 68(5): 865-868.
- McAlpine, D.F. and G. Godin. 1986. New records of snapping turtles, Chelydra serpentina, and painted turtles, Chrysemys picta, from New Brunswick. Canadian Field Naturalist 100(1):63-68.
- McEneaney, T. and J. Jensen. 1974. The reptiles and amphibians of the Charles M. Russell National Wildlife Range - 1974. Charles M. Russell National Wildlife Refuge. Lewistown, MT. 3 p.
- Medem, F. 1977. Contribucion al conocimiento sobre la taxonomia, distribucion geografica y ecologia de la tortuga "bache" (Chelydra serpentina acutirostris). Caldasia 12(56):41-98.
- Meeks, R.L. and G.R. Ultsch. 1990. Overwintering behavior of snapping turtles. Copeia 1990(3): 880-884.
- Meylan, P.A., C.A. Stevens, M.E. Barnwell, and E.D. Dohm. 1992. Observations on the turtle community of Rainbow Run, Marion Co., Florida. Florida Scientist 55(4): 219-228.
- Miller, K. 1993. The improved performance of snapping turtles (Chelydra serpentina) hatched from eggs incubated on a wet substrate persists through the neonatal period. Journal of Herpetology 27(2): 228-233.
- Miller, K. and G.C. Packard. 1992. The influence of substrate water potential during incubation on the metabolism of embryonic snapping turtles (Chelydra serpentina). Physiological Zoology 65(1): 172-187.
- Miller, K., G.C. Packard, and M.J. Packard. 1987. Hydric conditions during incubation influence locomotor perrformance of hatchling snapping turtles. Journal of Experimental Biology 127: 401-412.
- Mitchell, J.C. 1995. Abnormal coloration in a common snapping turtle (Chelydra serpentina serpentina) from Virginia. Banisteria 6 1995:32-33.
- Mitchell, J.C. and R.O. De Sa. 1994. Terrapene carolina carolina (Eastern Box Turtle) reproduction. Herpetological Review 25(2): 64.
- Mitchell, J.C., M.C. Odom. 1995. Record clutch size for Chelydra serpentina (Testudines: Chelydridae) in Virginia. Brimleyana No. 22:73-75. 1995.
- Mitsukuri, K. 1895. How many times does a snapping turtle lay eggs in one season? Zoological Magazine of Tokyo 6: 143-147.
- Moll, D. and F.J. Dodd, Jr. 1985. New records for Chelydra serpentina rossignoni in Belize. Bulletin of the Maryland Herpetological Society 21(1): 34-37.
- Moravec, F. and M.D. Little. 2004. Redescription of Dracunculus globocephalus Mackin, 1927 (Nematoda: Dracunculidae), a parasite of the snapping turtle, Chelydra serpentina. Folia Parasitologica (Ceske Budejovice) 51(4):339-345.
- Mosimann, J.E. and J.R. Bider. 1960. Variation, sexual dimorphism, and maturity in a Quebec population of the common snapping turtle, Chelydra serpentina. Canadian Journal of Zoology 38: 19-38.
- Murphy, G.G. and J.F. Sharber, Jr. 1973. Preliminary analysis of activity of the common snapping turtle, Chelydra serpentina, as determined by biotelemetry. Journal of the Tennessee Academy of Science 48: 56.
- Netten, H. and F. Zuurmond. 1985. Offspring of the common snapping turtle Chelydra serpentina in the reptile zoo Iguana. Lacerta 44(3): 42-43.
- Norris-Elye, L.T.S. 1949. The common snapping turtle (Chelydra serpentina) in Manitoba. Canadian Field Naturalist 63: 145-157.
- Nussbaum, R.A., E.D. Brodie, Jr. and R.M. Storm. 1983. Amphibians and reptiles of the Pacific Northwest. University of Idaho Press. Moscow, ID. 332 pp.
- Obbard, M.E. 1983. Population ecology of the common snapping turtle, Chelydra serpentina, in north-central Ontario. Ph.D. Thesis, University of Guelph, Guelph, Ontario.
- Obbard, M.E. 1985. Can we safely harvest snapping turtles in Ontario? Canadian Amphibian and Reptile Conservation Society Bulletin 23(2): 1-4.
- Obbard, M.E. and R.J. Brooks. 1979. Factors affecting basking in a northern population of the common snapping turtle, Chelydra serpentina. Canadian Journal of Zoology 57: 435-440.
- Obbard, M.E. and R.J. Brooks. 1981a. A radio-telemetry and mark-recapture study of activity in the common snapping turtle, Chelydra serpentina. Copeia 1981: 630-637.
- OEA Research. 1985. Wildlife Inventory:Monitoring Report for the CX Ranch Project. 1983-1984. Unpublished report for Consolidation Coal Company, Pittsburgh, Pennsylvania.
- Overmann, S.R., and J.J. Krajicek. Snapping turtles (Chelydra serpentina) as biomonitors of lead contamination of the big river in Missouri's old lead belt. Environmental Toxicological Chemistry 14(4): 689-695.
- Packard, G.C. and M.J. Packard. 1990b. Patterns of survival at subzero temperatures by hatchling painted turtles and snapping turtles. Journal of Experimental Zoology 254(2): 223-236.
- Packard, G.C., G.L. Paukstis, T.J. Boardman, and W.H.N. Gutzke. 1985. Daily and seasonal variation in hydric conditions and temperature inside nests of common snapping turtles (Chelydra serpentina). Canadian Journal of Zoology 63(10): 2422-2429.
- Packard, G.C., K. Miller, and M.J. Packard. 1992. A protocol for measuring water potential in subterranean nests of reptiles. Herpetologica 48(2): 202-209.
- Packard, G.C., K. Miller, and M.J. Packard. 1993. Environmentally induced variation in body size of turtles hatching in natural nests. Oecologica 93(3): 445-448.
- Packard, G.C., K. Miller, M.J. Packard, and G.F. Birchard. 1999. Environmentally induced variation in body size and condition in hatchling snapping turtles (Chelydra serpentina). Canadian Journal of Zoology 77: 278-289.
- Packard, G.C., K.A. Ruble, M.J. Packard. 1993. Hatchling snapping turtles overwintering in natural nests are inoculated by ice in frozen soil. Journal of Thermal Biology 18(4): 185-188.
- Packard, G.C., M.J. Packard, and K. Miller. 1990. Chelydra serpentina (common snapping turtle). Fecundity. Herpetological Review 21(4): 92.
- Packard, G.C., M.J. Packard, K. Miller, and T.J. Boardman. 1985. Influence of water potential and temperature on eggs of snapping turtles (Chelydra serpentina) incubating on different types of substrates. American Zoology 25: 134A.
- Packard, G.C., M.J. Packard, K. Miller, and T.J. Boardman. 1987. Influence of moisture, temperature, and substrate on snapping turtle eggs and embryos. Ecology 68 (4): 983-993.
- Packard, G.C., T.L. Taigen, M.J. Packard, and T.J. Boardman. 1980. Water relations of pliable-shelled eggs of common snapping turtles (Chelydra serpentina). Canadian Journal of Zoology 58: 1404-1411.
- Packard, M.J. 1980. Ultrastructural morphology of the shell and shell membrane of eggs of common snapping turtles (Chelydra serpentina). Journal of Morphology 165: 187-204.
- Paterson, W.B. and S.S. Desser. 1976. Observations on Haemogregarina balli sp.n. from the common snapping turtle, Chelydra serpentina. Journal of Photozoology 23: 294-301.
- Pell, S.M. 1940. Notes on the food habits of the common snapping turtle. Copeia 1940: 131.
- Pell, S.M. 1941. Notes on the habits of the common snapping turtle, Chelydra serpentina (Linn.) in central New York. Unpubl. MS Thesis, Cornell University, Ithaca, New York.
- Petokas, P.J. and M.M. Alexander. 1980. The nesting of Chelydra serpentina in northern New York. Journal of Herpetology 14:239-244.
- Phillips, C.A., W.W. Dimmick, and J.L. Carr. 1996. Conservation genetics of the common snapping turtle (Chelydra serpentina). Conservation Biology 10(2): 397-405.
- Pisani, G.R. 2004. Chelydra serpentina (common snapping turtle) mating behavior. Journal of Kansas Herpetology 12:17.
- Platt, T.R. 1988. Hapalorhynchus brooksi, new species (Trematoda: Spirorchiidae) from the snapping turtle (Chelydra serpentina), with notes on Hapalorhynchus gracilis and Hapalorhynchus stunkardi. Proceedings of the Helminthological society of Washington 55(2): 317-323.
- Pritchard, P.C.H. 1979a. Encyclopedia of turtles. T.F.H. Publications, Hong Kong.
- Pryor, G.S. 1996. Observations of shorebird predation by snapping turtles in eastern Lake Ontario. Wilson Bulletin 108(1): 190-192.
- Punzo, F. 1975. Studies on the feeding behavior, diet, nesting habits and temperature relationships of Chelydra serpentina osceola (Chelonia: Chelydridae). Journal of Herpetology 9: 207-210.
- Raney, E.C. and R.A. Josephson. 1954. Record of combat in the snapping turtle, Chelydra serpentina. Copeia 1954: 228.
- Redmer, M. 1986. Herpetofauna observed by a CHS member in northern Minnesota. Chicago Herpetological Society Newsletter 1986:1
- Reichel, J. and D. Flath. 1995. Identification of Montana's amphibians and reptiles. Montana Outdoors 26(3):15-34.
- Reichel, J.D. 1995. Montana Species of Special Concern. Montana Natural Heritage Program, Helena, Montana. 10 pp.
- Rhen, T. and J.W. Lang. 1994. Temperature-dependent sex determination in the snapping turtle: manipulation of the embryonic sex steroid environment. General and Comparative Endocrinology 96(2): 243-254.
- Rhen, T. and J.W. Lang. 1995. Phenotypic plasticity for growth in the common snapping turtle: effects of incubation temperature, clutch, and their interaction. American Naturalist 146(5): 726-747.
- Richmond, N.D. 1958. The status of the Florida snapping turtle, Chelydra osceola stejneger. Copeia 1958:41-43.
- Robinson, C. and J.R. Bider. 1988. Nesting synchrony - a strategy to decrease predation of snapping turtle (Chelydra serpentina) nests. Journal of Herpetology 22(4): 470-473.
- Roff, D.A. 1994. Evolution of dimorphic traits: Effect of directional selection on heritability. Heredity 72(1): 36-41.
- Roh, B.R. 1994a. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 25(4): 162-163.
- Russell, A. P. and A. M. Bauer. 1993. The amphibians and reptiles of Alberta. University of Calgary Press. Calgary, Alberta. 264 p.
- Saba, V.S. and J.R. Spotila. 2003. Survival and behavior of freshwater turtles after rehabilitation from an oil spill. Environmental Pollution 126(2):213-223.
- Saumure, R.A. 1996. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 27(3): 150.
- Schladweiler, Philip, and John P. Weigand., 1983, Relationships of endrin and other chlorinated hydrocarbon compounds to wildlife in Montana, 1981-1982. September 1983.
- Schuett, G.W. and R.E. Gatten. 1980. Thermal preference in snapping turtles (Chelydra serpentina). Copeia 1980: 149-152.
- Schwab, D. 1988a. Field notes. Chelydra serpentina serpentina (common snapping turtle). Catesbeiana 8(2): 33.
- Schwab, D. 1991. Chelydra serpentina serpentina (common snapping turtle). Catesbeiana 11(1): 19.
- Scott, A.F. and E. Twombly. 1994. Pond-dwelling amphibians of Land Between The Lakes (Kentucky and Tennessee): a quantitative survey. Journal of the Tennessee Academy of Science 69(2): 52-58.
- Shipman, P.A., D.R. Edds, and D. Blex. 1994. Macroclemys temminckii (alligator snapping turtle) and Chelydra serpentina (common snapping turtle). Agonistic behavior. Herpetological Review 25(1): 24-25.
- Siddall, M.E. and E.S. Gaffney. 2004. Observations on the leech Placobdella ornata feeding from bony tissue of turtles. Journal of Parasitology 90(5):1186-1188.
- Sims, P.A., G.C. Packard, and P.L. Chapman. 2001. The adaptive strategy for overwintering by hatchling snapping turtles (Chelydra serpentina). Journal of Herpetology 35(3): 514-517.
- Sites, J. W., Jr., J.W. Archie, C.J. Cole, V.O. Flores. 1992. A review of phylogenetic hypotheses for lizards of the genus (Phrynosomatidae): implications for ecological and evolutionary studies. Bulletin of the American Museum of Natural History (213): 1-110.
- Sperger, R.H., R. Humphrey, D. Chiszar, and H.M. Smith. 1995a. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 26(1): 43.
- St. John, A.D. 2002. Reptiles of the northwest: California to Alaska, Rockies to the coast. Lone Pine Publishing, Renton, WA. 272 p.
- Stebbins, R. C. 2003. A field guide to western reptiles and amphibians. 3rd Edition. Houghton Mifflin Company, Boston and New York. 533 p.
- Stebbins, R.C. 1985. A field guide to western reptiles and amphibians. Houghton Mifflin Company, Boston. 336 pp.
- Stevenson, D. and D. Crowe. 1992a. Chelydra serpentina osceola (Florida snapping turtle). Herpetological Review 23(3): 87-88.
- Steyermark, A.C. and J.R. Spoila. 2001. Effects of maternal identity and incubation temperature on hatching and hatchling morphology in snapping turtles, Chelydra serpentina. Copeia 2001(1): 129-135.
- Storey, K.B. 1990. Life in a frozen state: adaptive strategies for natural freeze tolerance in amphibians and reptiles. American Journal of Physiology 258: R559-R568.
- Street, H.M. 1989. Semipalmated sandpiper captured by turtle. Ontario Birds 7(2): 70.
- Struger, J., J.E. Elliot, C.A. Bishop, M.E. Obbard, R.J. Norstrom, D.V. Weseloh, M. Simon, and P. Ng. 1993. Environmental contaminants in eggs of the common snapping turtle (Chelydra serpentina serpentina) from the Great Lakes-St. Lawrence River Basin of Ontario, Canada (1981,1984). Journal of Great Lakes Research 19(4): 681-694.
- Stuart, J.N. 1995. Notes on aquatic turtles of the Rio Grande drainage, New Mexico. Bulletin of the Maryland Herpetological Society 31(3): 147-157.
- Stuart, J.N. and C.S. Clark. 1991. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 22(4): 134.
- Stuart, J.N. and C.W. Painter. 1988. Geographic distribution. Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 19(1): 21.
- Temple, S.A. 1987. Predation on turtle nests increases near ecological edges. Copeia 1987(1): 250-252.
- Thorp, T.J. and L.S. Clark. 1994. Common snapping turtle eats duck eggs. Wilson Bulletin 106(2): 416.
- Tinkle, D.W., A.E. Dunham, and J.D. Congdon. 1993. Life history and demographic variation in the lizard Sceloporus graciosus: a long-term study. Ecology 74(8): 2413-2429.
- Tobey, F.J. 1989a. Chelydra serpentina serpentina (common snapping turtle). Catesbeiana 9(2): 35.
- Tuberville, T.D. and V.J. Burke. 1994. Do flag markers attract turtle nest predators? Journal of Herpetology 28(4): 514-516.
- Tuberville, T.D., J.W. Gibbons, and J.L. Greene. Invasion of new aquatic habitats by male freshwater turtles. Copeia 3:713-715.
- Tucker, J.K. 1994a. An "easy" method to remove common snapping turtles (Chelydra serpentina) from Legler hoop traps. Herpetological Review 25(1): 13.
- Tucker, J.K. and F.J. Janzen. 1997. Incidence of twinning in turtles. Copeia 1997(1): 166-173.
- Tucker, J.K. and J.T. Lamer. 2004. Another challenge in snapping turtle (Chelydra serpentina) conservation. Turtle and Tortoise Newsletter 8:10-11.
- Ultsch, G.R. and D. Lee. 1983. Radiotelemetric observations of watering snapping turtles (Chelydra serpentes) in Rhode Island. Journal of the Alabama Academy of Science 54: 200-206.
- Upton, S.J., D.K. Odell, and M.T. Walsh. 1990. Eimeria caretta, new species (Apicomplexa: Eimeriidae) from the loggerhead sea turtle, Caretta caretta (Testudines). Canadian Journal of Zoology 68(6): 1268-1269.
- Vitt, L.J., J.P. Caldwell, and D.B. Shepard. 2005. Inventory of amphibians and reptiles in the Billings Field Office Region, Montana. Sam Noble Oklahoma Museum of Natural History and Department of Zoology, University of Oklahoma, Norman, OK. 33 pp.
- Vogt, R.C. and O. Flores-Villela. 1992. Effects of incubation temperature on sex determination in a community of Neotropical freshwater turtles in southern Mexico. Herpetologica 48(3): 265-270.
- VTN Colorado, Inc. Decker Coal Company., 1975, Draft environmental impact assessment for the proposed North Extension of the West Decker Mine.
- Waage, B.C. 1998. Western Energy Company Rosebud Mine 1997 annual wildlife monitoring report December 1, 1996 to November 30, 1997 survey period. Western Energy Company, Colstrip, MT.
- Walker, D., P.E. Moler, K.A. Buhlmann, and J.C. Avise. 1998. Phylogeographic uniformity in mitochondrial DNA of the snapping turtle (Chelydra serpentina). Animal Conservation 1: 55-60.
- Walley, H.D. 1993. Chelydra serpentina (snapping turtle): Predation. Herpetological Review 24(4): 148-149.
- Watermolen, D.J. 1990a. Chelydra serpentina (snapping turtle). Herpetological Review 21(1): 24.
- Watermolen, D.J. 1990b. Chelydra serpentina (snapping turtle). Herpetological Review 21(2): 38.
- Weidl, D.A. 1985. Wildlife observations along Saskatchewan’s Souris River 1978-1979. Blue Jay 43(4): 248-252.
- Wermuth, H. and R. Martens. 1977. Liste der rezenten amphibien und reptilien: testudines, crocodilia, rhynchocephalia. Das Tierreich 100.
- Werner, J.K., B.A. Maxell, P. Hendricks and D.L. Flath. 2004. Amphibians and Reptiles of Montana. Mountain Press Publishing Company: Missoula, MT. 262 pp.
- Westmoreland Resources, Inc., Hardin, MT., 1981, 1981 Wildlife Report. April 1982.
- White, J.B. and G.G. Murphy. 1973. The reproductive cycle and sexual dimorphism of the common snapping turtle, Chelydra serpentina serpentina. Herpetologica 29: 240-246.
- Whiting, M.J. and A.H. Price. 1994. Not the last picture show: New collection records from Paris, Texas (and other places). Herpetological Review 25(3): 130.
- Wiens, J.J. and T.W. Reeder. 1997. Phylogeny of the spiny lizards (Sceloporus) based on molecular and morphological evidence. Herpetological Monographs 11:1-101.
- Wilhoft, D.C. 1986. Eggs and hatchling components of the snapping turtle (Chelydra serpentina). Comparative Biochemistry and Physiology A Comparative Physiology 84(3): 483-486.
- Wilhoft, D.C., E. Hotaling and P. Franks. 1983. Effects of temperature on sex determination in the embryos of snapping turtles, Chelydra serpentina. Journal of Herpetology 17: 38-42
- Wilhoft, D.C., M.G. Del Baglivo, and M.D. Del Baglivo. 1979. Observations on mammalian predation of snapping turtle nests. Journal of Herpetology 13: 435-438.
- Will, B.D., D. Chiszar, and H.M. Smith. 1995. Testudines: Chelydra serpentina serpentina (common snapping turtle). Herpetological Review 26(4): 208.
- Willey, J.S. and R.W. Blob. 2004. Tail kinematics of juvenile common snapping turtles during aquatic walking. Journal of Herpetology 38(3):360-369.
- Williams, D. and A. Norden. 1992. An albino snapping turtle (Chelydra serpentina) from Calvert County, Maryland. Maryland Naturalist 36(1-2): 21-22.
- Williamson, L.U., J.R. Spotila, and E.A. Standora. 1989. Growth, selected temperature, and CTM of young snapping turtles, Chelydra serpentina. Journal of Thermal Biology 14: 33-39.
- Wilson, A.K. 1994. Chelydra serpentina (snapping turtle). Herpetological Review 25(1): 32.
- Wilson, R. and A. Wilson. 1992. Adult mallard killed by snapping turtle. Passenger Pigeon 54(1):95.
- Wood, J.T. 1953. Protective behavior and photic orientation in hatchling snapping turtles, Chelydra serpentina (Linne), in an aquatic environment. J. Elisha Mitchell Science Society 68: 54-59.
- World Conservation Union. 1991. Tortoises and freshwater turtles: an action plan for their conservation. IUCN/SSC Tortoise and Freshwater Turtle Specialist Group, Gland, Switzerland.
- Yntema, C.L. 1960. Effects of various temperatures on the embryonic development of Chelydra serpentina. Anatomical Record 136: 305-306.
- Yntema, C.L. 1968. A series of stages in the embryonic development of Chelydra serpentina. Journal of Morphology 125: 219-251.
- Yntema, C.L. 1970a. Observations on females and eggs of the common snapping turtle, Chelydra serpentina. American Midland Naturalist 84: 69-76.
- Yntema, C.L. 1970b. Temperature levels and periods of sex determination during incubation of eggs of Chelydra serpentina. Journal of Morphology 159: 17-28
- Yntema, C.L. 1970c. Twinning in the common snapping turtle, Chelydra serpentina. Anatomical Record 166: 491-498.
- Yntema, C.L. 1971. Incidence and survival of twin embryos of the common snapping turtle, Chelydra serpentina. Copeia 1971: 755-758.
- Yntema, C.L. 1976. Effects of incubation temperatures on sexual differentiation in the turtle, Chelydra serpentina. Journal of Morphology 150: 453-462
- Yntema, C.L. 1978. Incubation times for eggs of the turtle Chelydra serpentina (Testudines: Chelydridae) at various temperatures. Herpetologica 34: 274-277.
- Yntema, C.L. 1979. Temperature levels and periods of sex determination during incubation of eggs of Chelydra serpentina. Journal of Morphology 159: 17-28.
- Yntema, C.L. 1981. Characteristics of gonads and oviducts in hatchlings and young of Chelydra serpentina resulting from three incubation temperatures. Journal of Morphology 167: 297-304.
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