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		California Butterwort - Pinguicula macroceras
		
			
				Other Names:  
				
				Pinguicula vulgaris var. macroceras
			
		
		
		
		
	 
	
			
            State Rank Reason (see State Rank above)
            In Montana Pinguicula macroceras is found only in Glacier National Park (GNP).  Although plants are relatively protected in GNP, populations are not that common, and many have small numbers. Information on about half of the known locations was collected prior to 1970. There is a potential for impacts where plants occur near trails and roads. Plants have been observed by some botanists as has having low fruit set. Currently data on locations, population sizes, threats, reproduction, as well as, better mapping are needed.
			
               
			
				- Details on Status Ranking and Review
                
					
					    
                    	
                    	    Range ExtentScoreD - 1,000-5,000 sq km (~400-2,000 sq mi) Comment1,038 square kilometers. Area of OccupancyScoreD - 6-25 4-km2 grid cells CommentMontana can be divided into 30,390 4x4 square kilometer cells.  For this species plant observations occur in 14 of these 4x4 square kilometer cells. Number of PopulationsScoreB - 6 - 20 Comment30 observations representing about 21 discrete locations. Environmental SpecificityScoreA - Very narrow. Specialist or community with key requirements scarce CommentCoefficient of Conservation value is 9 indicating it exhibits a high fedelity to a narrow range of ecological tolerance and does not tolerate disturbance. ThreatsScoreD - Low CommentPotential for impacts occur where plants grow near trails in Glacier National Park. Populations are often small. 
                    	        
                    	     Intrinsic VulnerabilityScoreB - Moderately vulnerable CommentBumblebeers visit flowers, but successful fruit set rates appear to low. 
 
 
		
	 
	
	 
		General Description
		PLANTS: Carnivorous, perennial forbs that grow from a fibrous root. Stems are single, leafless (scapose), and terminate into a single flower.  Source: Lesica et al. 2012
LEAVES: Simple, succulent, basal leaves with short petioles. Blades are 1–4 cm long, oblong with enrolled, entire margins.  Upper surface is sticky from glands (viscid).  Source: Lesica et al. 2012
INFLORESCENCE: A solitary purple flower that terminates a bractless peduncle that is 3–12 cm long. Sepals are 3–5 mm long, the 5 lobes longer than the tube. Petals are purple, funnelform, somewhat bilabiate, and 15–25 mm long with a spur of 7–11 mm long. Fruit is a capsule.  Source: Lesica et al. 2012
In Montana our plants are variety macroceras (Link.) Herder (Lesica et al. 2012).
		
	
	
	Species Range
	
		
			Montana Range
		Range Descriptions
			 
			
			
					
						 Native
						Native
					 
				
				
					
			 
			
		 
		
	 
	
    		Range Comments
			Circumboreal south to California, Montana, Michigan and New York (Lesica et al. 2012).
			
		
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 32
		
		
(Click on the following maps and charts to see full sized version)
		Map Help and Descriptions
		
		
			
				Relative Density
				 
			 
		
			
				Recency
				 
			 
		
		 
		
			
				 
			
				 
			
					 (Observations spanning multiple months or years are excluded from time charts)
				
			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Habitat
		Wet soil of fens, ledges, and stream banks in the subalpine zone or rarely in the alpine zone in Montana (Lescia et al. 2012).
		
			
		
		
	
		Ecology
		PLANT-ANIMAL INTERACTIONS
Plants are carnivorous. Insects are caught on the sticky leaves.
		
	
		Reproductive Characteristics
		Plants reproduce from seed.
FRUITS
Capsule: globose, ca. 4 mm high (Lesica et al. 2012).
		
	
		Stewardship Responsibility
		
		
	
		Threats or Limiting Factors
		STATE THREAT SCORE REASON
Threat impact not assigned because threats are not documented. (MTNHP Threat Assessment 2021).
		
	
	References
	
		
			- Literature Cited AboveLegend:   View Online Publication View Online Publication Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
 MTNHP Threat Assessment. 2021. State Threat Score Assignment and Assessment of Reported Threats from 2006 to 2021 for State-listed Vascular Plants. Botany Program, Montana Natural Heritage Program, Helena, Montana. MTNHP Threat Assessment. 2021. State Threat Score Assignment and Assessment of Reported Threats from 2006 to 2021 for State-listed Vascular Plants. Botany Program, Montana Natural Heritage Program, Helena, Montana.
 
- Additional ReferencesLegend:   View Online Publication View Online Publication
 Do you know of a citation we're missing? Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p.
 
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