Greenleaf Manzanita - Arctostaphylos patula
Arctostaphylos x media
State Rank Reason (see State Rank above)
Known from two or three seperate locations in Montana. Population sizes are very small and are susceptible to the negative effects associated with such. Additional negative impacts from timber harvesting, invasive weeds and development are possible.
Primarily a species of the Great Basin and California, and disjunct in Montana. Not known from either Idaho or Wyoming.
Green-leaf Manzanita is a shrub with reddish brown, spreading stems which are up to 1 m (3 ft) high and can propagate by layering. The alternate, yellow-green leaves have broadly elliptic blades which are 2-5 cm long with entire margins and petioles less than 1/2 as long. Foliage is sparsely covered with short hairs and yellow glands. Short-stalked, pink flowers are clustered in branched inflorescences on the stem tips. The urn-shaped corolla, which is ca. 6 mm long, has 5 short lobes, and the calyx has 5 nearly separate sepals. There are 10 stamens. The brownish, glabrous berry is 7-10 mm across.
Flowering in late May-early June, fruiting in August.
This is the only Arctostaphylos species with a bush-stature in our area; distinguished from other bush-forming genera in the Health Family by the fruit which is free of, and not enclosed by, the calyx, alternate leaves, 5-part flowers, and corolla length less than 1 cm. A. uva-ursi has trailing stems that rarely are over 0.2 m tall. Hybrids between these two species are known elsewhere in their range.
South-central WA to CA, east to AZ, NV and northwest MT. Disjunct.
Observations in Montana Natural Heritage Program Database
Number of Observations:
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(Observations spanning multiple months or years are excluded from time charts)
Rocky soil in open coniferous forests in the montane zone.
Ecological Systems Associated with this Species
The following animal species have been reported as pollinators of this plant species or its genus where their geographic ranges overlap: Bombus bifarius
, Bombus melanopygus
, Bombus insularis
, and Bombus flavidus
(Thorp et al. 1983, Williams et al. 2014).
- Literature Cited AboveLegend: View Online Publication
- Thorp, R.W., D.S. Horning, and L.L. Dunning. 1983. Bumble bees and cuckoo bumble bees of California (Hymenoptera: Apidae). Bulletin of the California Insect Survey 23:1-79.
- Williams, P., R. Thorp, L. Richardson, and S. Colla. 2014. Bumble Bees of North America. Princeton, NJ: Princeton University Press. 208 p.
- Additional ReferencesLegend: View Online Publication
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- Ball C.T., J. Keeley, H. Mooney, J. Seemann, and W. Winner. 1983. Relationship between form, function, and distribution of two Arctostaphylos species (Ericaceae) and their putative hybrids. Acta Oecologica 4:153-164.
- Diggs, G.M., D. Soltis, and P. Soltis. 1988. Hybridization and genetic variation in Arctostaphylos (Ericaceae). American Journal of Botany 75(6):169.
- Ellstrand, N.C., J.M. Lee, J.E. Keeley, and S.C Keeley. 1987. Ecological isolation and introgression: biochemical confirmation of introgression in an Arctostaphylos (Ericaceae) population. Acta Oecologica 8:299-308.
- Lesica, P. and P. V. Wells. 1986. Noteworthy collection: Montana. Madrono 33:227-228.
- Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
- Schierenbeck, K.A. 1988. Evolutionary relationships among Arctostaphylos Mewukka and associated species. American Journal of Botany 75(6):204.