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		Alkali Sagebrush - Artemisia arbuscula ssp. longiloba
		
			
				Other Names:  
				
				Artemisia longiloba
			
		
		
		
		
Native Species
			Global Rank: 
G5T4
			State Rank: 
S3S4
			(see State Rank Reason below)
				C-value: 
			
			
			
			Agency Status
			USFWS: 
			USFS: 
			BLM: 
		
			
			
				
			
			
				External Links
				
			
		 
	 
	
			
            State Rank Reason (see State Rank above)
            Artemisia arbuscula subspecies longiloba is known from about 17 locations in the very southern portions of Beaverhead, Madison, and Park Counties. While the range of subspecies longiloba appears limited within Montana, its presence does not suggest rarity. Current information on this plant's locations, population sizes, and threats are greatly needed.
			
               
			
				- Details on Status Ranking and Review
                
					
					    
                    	
                    	    Range ExtentScoreE - 5,000-20,000 sq km (~2,000-8,000 sq mi) Area of OccupancyScoreD - 6-25 4-km2 grid cells Number of PopulationsScoreB - 6 - 20 Number of Occurrences or Percent Area with Good Viability / Ecological IntegrityScoreC - Few (4-12) occurrences with excellent or good viability or ecological integrity Environmental SpecificityScoreC - Moderate. Generalist or community with some key requirements scarce ThreatsScoreD - Low CommentNo known threats. 
                    	        
                    	     
 
 
		
	 
	
	 
		General Description
		PLANTS: Aromatic shrubs of 10–40 cm height (Lesica 2012) that sprout from roots (FNA 2006). Stems are gray-green to brown, brittle, and glabrate (FNA 2006).  Plants are glabrate to tomentose, and sometimes with obscure resin dots beneath the tomentum (FNA 2006; Lesica 2012). 
LEAVES: Aromatic, gray-green, wedge-shaped (cuneate), 5–25 mm long, and tipped with 3 lobes (FNA 2006).  Upper leaves are densely hairy (not sticky) (FNA 2006).  Leaves on flowering stems are deciduous while those on non-flowering stems are persistent (FNA 2006).
INFLORESCENCE: Spiciform with linear bracts (Lesica 2012). 
In Montana we have subspecies arbuscula and longiloba (refer to Habitat, Phenology & Diagnostic Characteristics).
		
	
		Phenology
		Subspecies arbuscula flowers from mid- to late summer (FNA 2006), which in Montana is mid-August to September (Lesica 2012).  
Subspecies longiloba flowers from early to late spring (FNA 2006), which in Montana is from July to mid-August (Lesica 2012). Its ability to begin blooming as snow melts in the spring is a unique characteristic for the subgenus Tridentatae (FNA 2006).
		
	
		Diagnostic Characteristics
		Refer also to Phenology.
Subspecies arbuscula has leaves with acute lobes and stiffly erect to ascending stems (Lesica 2012). Leaves are broadly cuneate (lobes less than half the blade length, 1-3 mm wide, and rounded) (FNA 2006). The involucres are 3.5-4 mm tall and 2-4.5 mm in diameter (FNA 2006). Plants are found on stony calcareous soil (Lesica 2012)
Subspecies longiloba has leaves with rounded lobes and flexuous, prostrate-spreading stems (Lesica 2012). Leaves are broadly cuneate (4-10 mm by 2-5 mm, often irregularly lobed, with rounded lobes, and middle lobes overlapping lateral lobes) (FNA 2006). The involucres are 2-3 mm tall and 1.5-2.5 mm in diameter (FNA 2006). Plants are found on fine-textured soil of stream terraces (Lesica 2012).  Ecologically it is distinguished from other subspecies by growing at low elevations in fine-grained clay soils (FNA 2006).
		
	
    		Range Comments
			In Montana subspecies arbuscula has been found in Beaverhead, Gallatin, Madison, and Park Counties (https://www.pnwherbaria.org). Its range includes California, Colorado, Idaho, Montana, Nevada, Utah, Washington, and Wyoming.
In Montana subspecies longiloba has been found in Beaverhead, Madison, and Park Counties (MTNHP 2018 botany database). Its range includes California, Colorado, Idaho, Montana, Oregon, Utah, and Wyoming.
			
		
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 20
		
		
(Click on the following maps and charts to see full sized version)
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				 (Observations spanning multiple months or years are excluded from time charts)
			
			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Habitat
		Subspecies arbuscula plants are found in rocky sedimentary soils in high valleys and mountain slopes at elevations from 1500 to 3800 meters (FNA 2006).  In Montana plants occur on stony calcareous soil (Lesica 2012).  
Subspecies longiloba plants are found on clay soils of alkaline basins and valleys, occasionally on outwash plains of mountains, at elevations from 1500 to 2500 meters (FNA 2006). In Montana plants occur on fine-textured soil of stream terraces (Lesica 2012).
		
		
	
		Ecology
		Artemisia arbuscula ssp. longiloba may be heavily browsed in winter where there is little snow cover (Boyle and Reeder 2005). This taxon is tolerant of saturated soils in spring and dry periods in summer and fall (Boyle and Reeder 2005).
		
	
		Reproductive Characteristics
		Artemisia arbuscula flowers (heads):  The involucre is campanulate, about 2–5 mm high with green, tomentose phyllaries (Lesica 2012). The receptacle is glabrous. Heads consist of 4-8 perfect (contains both male and female structures) disk flowers that are glabrous or glandular; corolla is about 2 mm long (Lesica 2012). Achenes are resinous and less than 1 mm long (Lesica 2012). 
The involucres for subspecies arbuscula are 3.5-4 mm tall and 2-4.5 mm in diameter (FNA 2006). 
The involucres for subspecies longiloba are 2-3 mm tall and 1.5-2.5 mm in diameter (FNA 2006).
		
	
		Stewardship Responsibility
		
		
	
	References
	
		
			- Literature Cited AboveLegend:   View Online Publication View Online Publication Boyle, S. A. and D. R. Reeder. 2005. Colorado sagebrush: a conservation assessment and strategy. Grand Junction: Colorado Division of Wildlife. Boyle, S. A. and D. R. Reeder. 2005. Colorado sagebrush: a conservation assessment and strategy. Grand Junction: Colorado Division of Wildlife.
 eFloras. 2017. Flora of North America. Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA. eFloras. 2017. Flora of North America. Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
 Flora of North America Editorial Committee. 2006. Flora of North America North of Mexico. Vol. 19. Magnoliophyta: Asteridae, part 6: Asteraceae, part 1. Oxford Univ. Press, New York. xxiv + 579 pp. Flora of North America Editorial Committee. 2006. Flora of North America North of Mexico. Vol. 19. Magnoliophyta: Asteridae, part 6: Asteraceae, part 1. Oxford Univ. Press, New York. xxiv + 579 pp.
 Flora of North America Editorial Committee. 2006. Flora of North America North of Mexico. Vol. 20. Magnoliophyta: Asteridae, part 7: Asteraceae, part 2. Oxford Univ. Press, New York. xxii + 666 pp. Flora of North America Editorial Committee. 2006. Flora of North America North of Mexico. Vol. 20. Magnoliophyta: Asteridae, part 7: Asteraceae, part 2. Oxford Univ. Press, New York. xxii + 666 pp.
 Kartesz, J.T. 2015. The Biota of North America Program (BONAP) North American Plant Atlas. Chapel Hill, N.C. [maps generated from Kartesz, J.T. 2015. Floristic Synthesis of North America, Version 1.0. Biota of North America Program (BONAP). (in press)]. Kartesz, J.T. 2015. The Biota of North America Program (BONAP) North American Plant Atlas. Chapel Hill, N.C. [maps generated from Kartesz, J.T. 2015. Floristic Synthesis of North America, Version 1.0. Biota of North America Program (BONAP). (in press)].
 Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
 
- Additional ReferencesLegend:   View Online Publication View Online Publication
 Do you know of a citation we're missing? Dillard, S.L. 2019. Restoring semi-arid lands with microtopography. M.Sc. Thesis. Bpzeman, MT: Montana State University. 97 p. Dillard, S.L. 2019. Restoring semi-arid lands with microtopography. M.Sc. Thesis. Bpzeman, MT: Montana State University. 97 p.
 Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p.
 Zamora, B., and P.T. Tueller. 1973. Artemisia arbuscula, A. longiloba, and A. nova habitat types in northern Nevada. Great Basin Naturalist 33(4):225-242. Zamora, B., and P.T. Tueller. 1973. Artemisia arbuscula, A. longiloba, and A. nova habitat types in northern Nevada. Great Basin Naturalist 33(4):225-242.
 
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