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		A Syntrichia Moss - Syntrichia princeps
		
			
				Other Names:  
				
				Tortula princeps
			
		
		
		
		
	 
	
	 
		General Description
		Plants: Acrocarpous (Vitt 1988). Growing in upright, crowded clumps or patches of upright shoots, green distally, russet below (Smith 1980). Stems 1-4 cm tall (occasionally taller), frequently forked, sometimes with a central strand present (Lawton 1971). 
Leaves: Incurved, slightly contorted and spiraled around the stem to varying degrees when dry, curved outwardly to broadly spreading when wet, 2-4 mm in length, 1-1.5 mm in width, cupped, spatula-shaped; margins tightly rolled out and downward proximally up to 3/4 of the leaf length; leaf tips acute or sometimes cut off abruptly; costa extending into a long, saw-toothed awn, the awn mostly transparent except for the red base (FNA 2007).
Leaf Cells: Basal cells large, transparent, smooth, and fine-walled, nearly square or a little rectangular near the margins, longer interiorly, well-demarcated from the cells above; upper cells nearly square to 6-sided, somewhat swollen, each with 4-6 papillae (FNA 2007); costa in X-section with 2 guide cells, 2 adaxial cell layers, and a strong abaxial stereid band (Lawton 1971).
		
	
		Diagnostic Characteristics
		T. princeps differs from T. ruralis, T. papillosissima, and T. norvegia by its synoicous state (if present); otherwise by broader basal cells, and the presence of a central strand (if present) (FNA 2007).
		
	
    		Range Comments
			North American Range
Canada: BC; USA: states west of and including MT s to AZ; Mexico (FNA 2007). In Montana, known from Flathead, Lake, Lincoln, Missoula, Park, and Ravalli Counties (Elliott 2016).
			
		
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 16
		
		
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			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Habitat
		Tree bark, soil (FNA 2007) and basic rock, on bluffs (Smith 1980). Elevation: low to medium (FNA 2007).
		
		
	
		Reproductive Characteristics
		Archegonia and antheridia usually in the same inflorescence, but apparently on separate plants occasionally. Seta 1-1.8 cm tall, red. Capsule 3-4 mm in length, russet, a little bowed, with a conspicuous neck, the peristome’s 32 reddish and papillose thread-like divisions spiraled ca 2 revolutions (FNA 2007).
Specialized vegetative propagula unknown (FNA 2007).
		
	
		Stewardship Responsibility
		
		
	
	References
	
		
			- Literature Cited AboveLegend:   View Online Publication View Online Publication Elliott, J.C. and A.K. Pipp. 2018. A Checklist of Montana Mosses (1880-2018). Updated 3 January, 2020. Montana Natural Heritage Program, Helena, Montana. 73 pp. Elliott, J.C. and A.K. Pipp. 2018. A Checklist of Montana Mosses (1880-2018). Updated 3 January, 2020. Montana Natural Heritage Program, Helena, Montana. 73 pp.
 Flora of North America Editorial Committee, eds. 2007. Flora of North America North of Mexico. Volume 27. Bryophytes: Mosses, Part 1. Oxford University Press, Inc., NY. xxi + 713 pp. Flora of North America Editorial Committee, eds. 2007. Flora of North America North of Mexico. Volume 27. Bryophytes: Mosses, Part 1. Oxford University Press, Inc., NY. xxi + 713 pp.
 Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices. Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
 Smith, A.J.E. 1980. The Moss Flora of Britain and Ireland. Cambridge University Press, Cambridge. 705 pp. Smith, A.J.E. 1980. The Moss Flora of Britain and Ireland. Cambridge University Press, Cambridge. 705 pp.
 Vitt, D. J. Marsh, and R. Bovey. 1988. Mosses, Lichens & Ferns of Northwest North America. Seattle, WA: University of Washington Press. 296 p. Vitt, D. J. Marsh, and R. Bovey. 1988. Mosses, Lichens & Ferns of Northwest North America. Seattle, WA: University of Washington Press. 296 p.
 
- Additional ReferencesLegend:   View Online Publication View Online Publication
 Do you know of a citation we're missing? Elliot, J. C. 1993. Second checklist of Montana mosses. Unpublished report. U.S. Forest Service, Region 1. Missoula, MT. 45 pp. Elliot, J. C. 1993. Second checklist of Montana mosses. Unpublished report. U.S. Forest Service, Region 1. Missoula, MT. 45 pp.
 Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp. Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.
 
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