Angled Paludella Moss - Paludella squarrosa
Plants: Acrocarpous, tufted, dull-yellow green with a touch of chartreuse (Vitt 1988), often almost black below. Stems usually unbranched, frequently with rhizoids bunched in the leaf axils (Lawton 1971), 3-7 cm (FNA 2014).
Leaves: Stem leaves firm when wet or dry, 1.5-2 mm long, strongly squarrose-recurved in the apical half, obovate (FNA 2014) or ovate-lanceolate from an upright, oblong base, widest at the middle, keeled (Lawton 1971); margins serrulate apically, recurved near mid leaf; apex sharply-angled to short-acuminate; costa thin, not reaching the apex (FNA 2014).
Leaf Cells: Rounded or 6-sided, mammillose-papillose on both surfaces, thick-walled (FNA 2014); distal cells shorter than the elongate basal cells (Lawton 1971).
A calcareous fen habitat and strongly squarrose-recurved leaves are all that is required for identification (FNA 2014).
Greenland; Canada: AB, BC, MB, NL, NT, NS, NU, ON, QC, SK, YT; USA: AK, CO, IA, ME, MI, MN, MT, NY, VT; n Eurasia (FNA 2014). In Montana: Beaverhead, Carbon, Glacier, and Park Counties (Elliott 2016).
Observations in Montana Natural Heritage Program Database
Number of Observations:
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(Observations spanning multiple months or years are excluded from time charts)
Calcareous fens. Low to high elevations (FNA 2014).
Dioicous (FNA 2014). Perigonia and perichaetia terminal; perichaetial bracts lanceolate, upright-spreading, not squarrose; perigonia discoid; perigonial bracts upright with recurved tips (Lawton 1971). Seta 25-45 mm. Capsule 2-3 mm (FNA 2014), brown, smooth, bowed a little; neck creased when dry; operculum cone-shaped (Lawton 1971).
- Literature Cited AboveLegend: View Online Publication
- Elliott, J.C. and A.K. Pipp. 2018. A Checklist of Montana Mosses (1880-2018). Updated 3 January, 2020. Montana Natural Heritage Program, Helena, Montana. 73 pp.
- Flora of North America Editorial Committee, eds. 2014. Flora of North America North of Mexico. Volume 28. Bryophytes: Mosses, Part 2. Oxford University Press, Inc., NY. xxi + 702 pp.
- Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
- Additional ReferencesLegend: View Online Publication
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- Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.