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Heim's Hennediella Moss - Hennediella heimii
Other Names:
Desmatodon heimii
General Description
Plants: Acrocarpous, the tufts green with brown below, growing in large patches (turfs) or cushions. Stems approximately 5-10 mm, enclosing a central strand; axillary hairs consisting of about 5 cells, the lower ones occasionally with wide walls (FNA 2007).
Leaves: Leaves flat against the stem when dry, spreading when wet, broadly lanceolate to ovate, tubular, firm; the base sheathing the stem slightly; apex acute; the costa single (FNA 2007).
Leaf Cells: Proximal cells rectangular, (2-)4-6:1, the walls thin; distal medial cells quadrate to short-rectangular, hexagonal, 1-2:1; distal margins differentiated with elongate cells, occasionally thick-walled, 1-stratose; ventral cells of costa short-rectangular or quadrate, in 4-5 rows, the transverse section elliptic or rounded, lacking an adaxial stereid band, guide cells 2-4 per layer in 1-2 layers; dorsal side of costa including a stereid band of substereid cells, elliptic or rounded in X-section (FNA 2007).
Hennediella heimii var. heimii: Present in Montana. Leaves are tubular, but not concave or hood-shaped; margins toothed; apex acute; costa terminating at apex (percurrent) (FNA 2007).
Hennediella heimii var. arctica: Leaves are concave and frequently cucullate; margins commonly smooth; apex generally obtuse; costa frequently terminating before the apex. Leaves 0.5-0.8 mm and usually considerably smaller than var. heimii (FNA 2007).
Phenology
Variety heimii: Capsules ripen June-July (FNA 2007).
Variety arctica: Capsules ripen in August (late summer) (FNA 2007).
Diagnostic Characteristics
Extensive variation occurs in the leaf border and dentition of the margin (FNA 2007).
Tortula obtusifolia is similar but has no hint of marginal teeth (FNA 2007).
Range Comments
Variety heimii: Greenland; Canada: AB, BC, MB, NL, NU, ON, QC, SK, YT; USA: AK, CA, CO, ID, MT, NE, NM, OR, UT, WA, WY; s South America; Europe; Asia; Pacific Islands (New Zealand); Australia; Antarctica (FNA 2007). In Montana: Cascade, Flathead, Gallatin, Glacier, Ravalli, and Sweet Grass Counties (Elliott 2016).
Variety arctica: Greenland; Canada: NU, NT, ON; USA: AK; Europe; n Asia (FNA 2007).
Observations in Montana Natural Heritage Program Database
Number of Observations: 18
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Map Help and Descriptions
Relative Density
Recency
(Observations spanning multiple months or years are excluded from time charts)
Habitat
Variety heimii: Damp soil of banks and near streams, often alkaline (FNA 2007), including soil derived from volcanic ash (Elliott 2016).
Variety arctica: Open silt and soil (FNA 2007).
Reproductive Characteristics
Monoicous, either autoicous or synoicous. The capsule urn is wide-mouthed and 1-2.4(-3.5) mm in length; the peristome is lacking (FNA 2007).
References
- Literature Cited AboveLegend:
View Online Publication- Elliott, J.C. and A.K. Pipp. 2018. A Checklist of Montana Mosses (1880-2018). Updated 3 January, 2020. Montana Natural Heritage Program, Helena, Montana. 73 pp.
- Flora of North America Editorial Committee, eds. 2007. Flora of North America North of Mexico. Volume 27. Bryophytes: Mosses, Part 1. Oxford University Press, Inc., NY. xxi + 713 pp.
- Additional ReferencesLegend: View Online Publication
Do you know of a citation we're missing?- Elliot, J. C. 1993. Second checklist of Montana mosses. Unpublished report. U.S. Forest Service, Region 1. Missoula, MT. 45 pp.
Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.- Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
- Malcolm, W. M., and Nancy Malcolm. 2000. Mosses and Other Bryophytes: An Illustrated Glossary. Nelson, New Zealand: Micro-Optics Press.
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