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Montana Field Guide

Montana Field Guides

Bristly Bentleaf Moss - Campylophyllum hispidulum
Other Names:  Campylium hispidulum

Native Species

Global Rank: G5
State Rank: SNR

Agency Status
MNPS Threat Rank:

External Links

General Description
Plants: Pleurocarpous. Growing in open mats (Lawton 1971), green, sometimes with yellow tinges (FNA 2014), eventually turning brown. Stems creeping (Lawton 1971), freely forked to pinnate; central strand weak or lacking; hyalodermis not present; paraphyllia wanting (FNA 2014) or occasionally present, filiform or foliose (Lawton 1971); axillary hairs with up to 3 distal cells; rhizoids or their initials occurring on the outer surface of the costa attachment or on the stem, seldom producing woolly mats, branched (FNA 2014).

Leaves: Stem leaves crowded to lax, a little more upright when dry than when moist, heart- shaped to widely or narrowly egg-shaped (FNA 2014), strongly cupped (Crum & Anderson et al. 1981), 0.4-0.9 mm in length, 0.2-0.6 mm in width; margins flat or seldom curved back and down at the base, typically both smooth and finely saw-toothed above, but rarely smooth or saw-toothed throughout (Lawton 1971); acumen slightly more or less than half the length of the leaf, grooved; base upright to nearly spreading; heart-shaped to widely or narrowly egg-shaped, (FNA 2014).

Leaf Cells: Basal alar cells quadrangular, sometimes short, the distal alar cells quadrangular, elongated crosswise, sometimes short or square, the alar area extending 1/2 to ca 2/3 of the distance from the leaf edge to the costa attachment, the boundary not well-defined (FNA 2014); laminal cells incrassate (thick-walled), occasionally with tiny papillae dorsally from protruding cell ends (Lawton 1971); medial laminal cells narrowly 6-sided to very thin; marginal cells in 1 layer (FNA 2014).

Range Comments
North American Range

Canada: BC to NL; USA: AK, ID and MT, also central and eastern US (ND to TX, east from there to the coast) (FNA 2014). Known in Montana from Fallon, Flathead, Gallatin, and Glacier Counties (Elliott 2016).

Observations in Montana Natural Heritage Program Database
Number of Observations: 2

(Click on the following maps and charts to see full sized version) Map Help and Descriptions
Relative Density



(Observations spanning multiple months or years are excluded from time charts)

Bark crevices (Crum & Anderson et al. 1981), soil, bottoms of trees, and rotting wood in forests and covered habitats (FNA 2014).

Reproductive Characteristics
Autoicous. Seta 9-17 mm tall. Capsule bowed, level (FNA 2014), contracted under the opening when dry (Crum & Anderson et al. 1981); theca 0.7-2 mm in length (Lawton 1971); exostome edges toothed or with a few teeth above; endostome cilia knobby (FNA 2014) and without perforations (Crum & Anderson et al. 1981).

  • Literature Cited AboveLegend:   View Online Publication
    • Crum, H.A. and L.E. Anderson. 1981. Mosses of Eastern North America. 2 volumes. Columbia University Press, New York. 1328 pp.
    • Elliott, J.C. and A.K. Pipp. 2018. A Checklist of Montana Mosses (1880-2018). Updated 3 January, 2020. Montana Natural Heritage Program, Helena, Montana. 73 pp.
    • Flora of North America Editorial Committee, eds. 2014. Flora of North America North of Mexico. Volume 28. Bryophytes: Mosses, Part 2. Oxford University Press, Inc., NY. xxi + 702 pp.
    • Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
  • Additional ReferencesLegend:   View Online Publication
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    • Flowers, S. 1973. Mosses: Utah and the West. Brigham Young University, Provo, Utah. 567 p.
    • Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.
    • Malcolm, B., N. Malcolm, J. Shevock, and D. Norris. 2009. California Mosses. Nelson, New Zealand: Micro-Optics Press. 430 pp.
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Citation for data on this website:
Bristly Bentleaf Moss — Campylophyllum hispidulum.  Montana Field Guide.  .  Retrieved on , from