A Ptychostomum Moss - Ptychostomum lonchocaulon
Bryum lonchocaulon, Bryum cirrhatum
MNPS Threat Rank
Plants: Acrocarpous (Vitt 1988).Growing in large, loose to crowded patches (FNA 2014) or clumps (Flowers 1973) of erect shoots, green, or green with yellow tones. Stems mostly 1-2 cm tall (FNA 2014), shortly-branched and red (Flowers 1973), with leaves dense at the top of the stem (comose), and generally with rhizoids present (FNA 2014).
Leaves: Green, scarcely larger at the distal end of the stem, curved and twisted when dry, mostly 2-3 mm in length, egg- to lance-shaped, plane or faintly cupped, acuminate; margins rolled tightly back and downward proximally to half or more of the leaf length, the border strong (FNA 2014); costa somewhat yellow or brown, sometimes red proximally, extending well beyond the apex to form an awn, the awn long (sometimes more than half the blade length), spinulose (Flowers 1973) to somewhat smooth (FNA 2014).
Leaf Cells: Middle and upper laminal cells diamond-shaped (FNA 2014), frequently becoming nearly linear in the apex (Flowers 1973), 3-4:1, the walls fine or slightly thicker (firm), as wide as or narrower than the proximal laminal cells; proximal cells quadrangular, 3-4:1; border (limbidium) usually consisting of 3-4 rows of cells that are not yellowish, unistratose (FNA 2014).
Fruit ripens in spring to late summer or early fall (FNA 2014).
Bryum lisae var. cuspidatum resembles this species, but has a yellowish, weaker border (usually 2-3 rows of cells rather than 3-4) and faithfully synoicous reproduction (FNA 2014).
North American Range
Canada: BC, AB, and QC; USA: AK, WA s to CA and eastward to MT, WY, CO and NM, also MN and ME (FNA 2014). Known in Montana from Deer Lodge, Flathead, Lake, and Ravalli Counties (Elliott 2016).
Mostly in shaded areas on moist soil or stones, frequently on somewhat dry mountain sides (Flowers 1973). Elevation: 0-11,380 feet (FNA 2014).
Mixed synoicous, polygamous, or producing only antheridia (FNA 2014). Seta reddish (Flowers 1973), mostly 10-20 mm tall. Capsule pear-shaped, 2-4 mm in length, brown, the opening yellow (FNA 2014), the neck somewhat thick and usually not quite half the length of the spore sac region (Flowers 1973); peristome fully developed; exostome teeth yellow below, transparent above; exostome segments with widely ovate openings, the cilia long and transversely ridged (FNA 2014).
No specialized vegetative reproduction present (FNA 2014).
- Literature Cited AboveLegend: View Online Publication
- Elliott, J.C. and A.K. Pipp. 2018. A Checklist of Montana Mosses (1880-2018). Updated 3 January, 2020. Montana Natural Heritage Program, Helena, Montana. 73 pp.
- Flora of North America Editorial Committee, eds. 2014. Flora of North America North of Mexico. Volume 28. Bryophytes: Mosses, Part 2. Oxford University Press, Inc., NY. xxi + 702 pp.
- Flowers, S. 1973. Mosses: Utah and the West. Brigham Young University, Provo, Utah. 567 p.
- Vitt, D. J. Marsh, and R. Bovey. 1988. Mosses, Lichens & Ferns of Northwest North America. Seattle, WA: University of Washington Press. 296 p.
- Additional ReferencesLegend: View Online Publication
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- Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.
- Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
- Malcolm, B., N. Malcolm, J. Shevock, and D. Norris. 2009. California Mosses. Nelson, New Zealand: Micro-Optics Press. 430 pp.