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		American Springfly - Skwala americana
		
		
		
		
	 
	
	 
		General Description
		The Skwala is a colorful stonefly nymph that hatches in early spring when not many other insects are hatching, making it the first large, post-winter meal for trout, and good fishing for fisherman. The Skwala hatch begins as river waters begin to warm, usually the end of February, which provides a pre-runoff fishing opportunity. The Bitterroot River has become a favorite river for the skwala hatch, but many other large Montana rivers such as the Clark Fork River, Blackfoot, Smith, Yellowstone and Madison Rivers contains these insects in large enough numbers to make fishing the hatch worthwhile.
		
	
		Phenology
		The Skwala hatch begins as river waters begin to warm, usually the end of February and continues through April in dark canyon reaches that remain colder.
		
	
	
	Species Range
	
		
			Montana Range
		Range Descriptions
			 
			
			
					
						 Native
						Native
					 
				
				
					
			 
			
		 
		
	 
	
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 196
		
		
(Click on the following maps and charts to see full sized version)
		Map Help and Descriptions
		
		
			
				Relative Density
				 
			 
		
			
				Recency
				 
			 
		
		 
		
			
				 
			
				 
			
					 (Observations spanning multiple months or years are excluded from time charts)
				
			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Food Habits
		Merritt and Cummins (1996) report that members of the  families, Perlidae and Perlodidae, are largely predators eating other aquatic invertrebrates, especially Diptera (Chironomidae and Simuliidae, midges and blackflies) and Mayflies.
		
	
		Ecology
		The Skwala hatch begins as river waters begin to warm, usually the end of February,
		
	
		Stewardship Responsibility
		
		
	
	References
	
		
			- Literature Cited AboveLegend:   View Online Publication View Online Publication Cummins, K.W. and R.W. Merritt. 1996. Ecology and distribution of aquatic insects. Chapter 6, pages 74-86 in R.W. Merritt and K.W. Cummins (eds.) An Introduction to the Aquatic Insects of North America. Third Edition. Kendall/Hunt Publishing Company, Dubuque, Iowa. 862 pp. Cummins, K.W. and R.W. Merritt. 1996. Ecology and distribution of aquatic insects. Chapter 6, pages 74-86 in R.W. Merritt and K.W. Cummins (eds.) An Introduction to the Aquatic Insects of North America. Third Edition. Kendall/Hunt Publishing Company, Dubuque, Iowa. 862 pp.
 
- Additional ReferencesLegend:   View Online Publication View Online Publication
 Do you know of a citation we're missing? Fraley, J.J. 1978. Effects of elevated summer water temperatures below Ennis Reservoir on the microinvertebrates of the Madison River, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 120 p. Fraley, J.J. 1978. Effects of elevated summer water temperatures below Ennis Reservoir on the microinvertebrates of the Madison River, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 120 p.
 Gangemi, John Thomas. 1991. Comparative effects of wildfire and timber harvest on periphyton and zoobenthos in two northern Rocky Mountain streams. M.S. Thesis. University of Montana. Missoula, MT. Gangemi, John Thomas. 1991. Comparative effects of wildfire and timber harvest on periphyton and zoobenthos in two northern Rocky Mountain streams. M.S. Thesis. University of Montana. Missoula, MT.
 Glorvigen, T.H. 1972. The responses of insect communities in the East Gallatin River, Montana, to sewage effluents. M.Sc. Thesis. Bozeman, Montana: Montana State University. 24 p. Glorvigen, T.H. 1972. The responses of insect communities in the East Gallatin River, Montana, to sewage effluents. M.Sc. Thesis. Bozeman, Montana: Montana State University. 24 p.
 Gustafson, D. L. 1990. Ecology of aquatic insects in the Gallatin River drainage. Ph.D. Dissertation. Bozeman, MT: Montana State University. 194 p. Gustafson, D. L. 1990. Ecology of aquatic insects in the Gallatin River drainage. Ph.D. Dissertation. Bozeman, MT: Montana State University. 194 p.
 Hansen, Barry Steven. 1988. Changes in the benthic community of Lake Creek, Montana, resulting from mine tailings contamination. M.S. Thesis. University of Montana. Missoula, MT. Hansen, Barry Steven. 1988. Changes in the benthic community of Lake Creek, Montana, resulting from mine tailings contamination. M.S. Thesis. University of Montana. Missoula, MT.
 Stadnyk, L. 1971. Factors affecting the distribution of Stoneflies in the Yellowstone River, Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 36 p. Stadnyk, L. 1971. Factors affecting the distribution of Stoneflies in the Yellowstone River, Montana. M.Sc. Thesis. Bozeman, Montana: Montana State University. 36 p.
 
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