The Wolverine is a bear-like mustelid with massive limbs and long, dense, dark brown pelage, paler on the head, with two broad yellowish stripes extending from the shoulders and joining on the rump. Variable white or yellowish markings are often present on the throat and chest. The tail is bushy. The feet are relatively large (6.5 to 11.3 centimeters total length) with robust claws. Wolverines weigh between 7 and 32 kilograms and range from 0.9 to 1.1 meters in length. Females average about 10% less than males in linear measurements and 30% less in mass (Ingles 1965, Hall 1981, Nowak 1991).
Wolverines are most similar to Fishers (Martes pennanti) but are nearly twice as large. Fishers also lack the light colored lateral markings of the Wolverine and the tail is less bushy. Badgers have shorter legs and are much lighter colored with a distinctive black and white pattern on the face.
Western Hemisphere Range
Observations in Montana Natural Heritage Program Database
Number of Observations:
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Map Help and Descriptions
(Observations spanning multiple months or years are excluded from time charts)
Wolverines in northwestern Montana and Alaska tended to occupy higher elevations in summer and lower elevations in winter (Hornocker and Hash 1981, Whitman et al. 1986). Seasonal ranges were all within a large home range; dispersal movements of more than 300 kilometers are known (Magoun 1985, Gardner et al. 1986).
Wolverines are limited to alpine tundra, and boreal and mountain forests (primarily coniferous) in the western mountains, especially large wilderness areas. However, dispersing individuals have been found far outside of usual habitats. They are usually in areas with snow on the ground in winter. Riparian areas may be important winter habitat. When inactive, Wolverines occupy dens in caves, rock crevices, under fallen trees, in thickets, or similar sites. Wolverines are primarily terrestrial but may climb trees.
In Montana, Hornocker and Hash (1981) found most Wolverine use in medium to scattered timber, while areas of dense, young timber were used least. Wolverines avoided clearcuts and burns, crossing them rapidly and directly when they were entered at all. Hash (1987) reported Wolverines in the Northern Rocky Mountain region were associated with fir, pine, and larch. Aspen stands were also used, as were cottonwoods in riparian areas. Ecotonal areas appeared to be important habitat components (Hash 1987). Hatler (1989) believed Wolverines are not dependant on any particular vegetative habitat type. Banci (1986) reported "habitat requirements appear to be large, isolated tracts of wilderness supporting a diverse prey base, rather than specific plant associations or topography." South of the boreal forest, most habitat descriptions in the literature agree with Grove's (1988) characterization of "large, mountainous, and essentially roadless areas."
Ecological Systems Associated with this Species
- Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
- Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
- Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
- Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
- Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if
point observations were associated with that system.
Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.
If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: mtnhp.org/requests
) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.
- Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
- Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
- Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
- Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
- Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
- Maxell, B.A. 2000. Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
- Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
- Commonly Associated with these Ecological Systems
Forest and Woodland Systems
Recently Disturbed or Modified
Shrubland, Steppe and Savanna Systems
Wetland and Riparian Systems
- Occasionally Associated with these Ecological Systems
Forest and Woodland Systems
Recently Disturbed or Modified
Wetland and Riparian Systems
Wolverines are opportunistic. They feed on a wide variety of roots, berries, small mammals, birds' eggs and young, fledglings, and fish (Hatler 1989). They may attack moose, caribou, and deer hampered by deep snow. Small and medium size rodents and carrion (especially ungulate carcasses) often make up a large percentage of the diet. Prey is captured by pursuit, ambush, digging out dens (Biosystems Analysis 1989), or climbing into trees. They may cache prey in the fork of tree branches or under snow.
Wolverines are generally solitary and wide-ranging. They occur at relatively low densities (e.g., 1 per 65 square kilometers in northwestern Montana) (Hornocker and Hash 1981). Home ranges of males are larger than those of females, with home ranges of up to several hundred square kilometers. The mean annual home range of males was 535 square kilometers in Alaska, and 422 square kilometers in Montana. Female home ranges were 105 square kilometers in Alaska and 388 square kilometers in Montana (Hornocker and Hash 1981). Males in some areas apparently are territorial, but in Montana there was extensive overlap of the ranges of both the same and opposite sexes (Hornocker and Hash 1981). Apparently territory/range size depends on availability of denning sites and food supply (Wilson 1982). Some individuals travel regularly over the same route (Wilson 1982). Available evidence indicates that juveniles disperse usually around 30 to 100 kilometers from their natal range, though dispersal movements of more than 300 kilometers are known (Magoun 1985, Gardner et al. 1986). There are no important predators other than humans.
Although the Wolverine usually breeds in summer, the event may occur from April to October. Implantation is delayed and does not occur until winter. Gestation lasts 7 to 9 months; active gestation is 30 to 40 days. One to six (usually 2 to 4) young are born January through April, mainly in February or March, and reportedly April through June in the Pacific states (Ingles 1965), in a den among rocks or tree roots, in a hollow log, under a fallen tree, or in dense vegetation, including sites under snow. Young are weaned beginning at about 7 or 8 weeks, and separate from the mother in the fall. They are sexually mature generally in the second or third year. Males sexually mature sometimes as yearlings (Alaska and Yukon); males over three years old were sexually mature in British Columbia. Some females mature at 12 to 15 months and produce their first litter when two years old (Wilson 1982). In some areas, females may produce litters only every 2 or 3 years. In British Columbia, most mature females were reproductively active. The Wolverine lives to an age of up to about 10, or sometimes 15 to 18, years.
On October 8, 2020 the US Fish and Wildlife Service determined that the best available science show that factors affecting wolverine populations were not as significant as previously believed when the species was proposed for listing under the Endangered Species Act in 2013. The service cites new research and analysis that demonstrate stability in populations within the US and connectivity with those in Canada as the reason for withdrawing the proposal to list the species as threatened. Additional information on the species' management can be found on the U.S. Fish and Wildlife Service's Species Account
Wolverines were nearly extinct in Montana during the early 1900's and have been increasing in numbers and range since. Recovery originated in northwestern Montana and subsequently spread to its current range (Newby and Wright 1955, Newby and McDougal 1964).
Wolverines are classified as a furbearer in Montana. However, the trapping season is currently suspended with a statewide quota of zero.
- Literature Cited AboveLegend: View Online Publication
- Banci, V. 1986. The wolverine in the Yukon: myths and management. Discovery 15(4):134-137.
- Biosystems Analysis, Inc. 1989. Endangered species alert program manual: species accounts and procedures. Southern California Edison Environmental Affairs Division.
- Gardner, C.L., W.B. Ballard and R.H. Jessup. 1986. Long distance movements by an adult wolverine. Journal of Mammalogy 67(3):603.
- Groves, C.R. 1988. Distribution of the wolverine in Idaho as determined by mail questionnaire. Northwest Science 62(4):181-185.
- Hall, E.R. 1981. The mammals of North America, volumes I and II. John Wiley & Sons, New York, NY. 1181 pp.
- Hash, H.S. 1987. Wolverine. In: M. Novak, J. Baker, M. Obbard and B. Malloch (eds). Wild furbearer management and conservation in North America. pp. 574-585. Ontario, CA: Ontario Trappers Association. 1150 p.
- Hatler, D.F. 1989. A wolverine management strategy for British Columbia. British Columbia Ministry of Environment, Victoria. Wildlife Bulletin B-60. 124 p.
- Hornocker, M.G. and H.S. Hash. 1981. Ecology of the wolverine in northwestern Montana. Canadian Journal of Zoology 59(7):1286-1301.
- Ingles, L.G. 1965. Mammals of the Pacific states. Stanford University Press, Stanford, California. 506 pp.
- Magoun, A.J. 1985. Population characteristics, ecology, and management of wolverines in northwestern Alaska. Ph.D. Dissertation. Fairbanks, AK: University of Alaska. 197 p.
- Nowak, R.M. 1991. Walker's mammals of the world. Fifth edition. Volumes I and II. Johns Hopkins University Press, Baltimore. 1629 pp.
- Wilson, D. E. 1982. Wolverine, Gulo gulo. In: J. A. Chapman and G. A. Feldhamer, eds. Wild mammals of North America: biology, management, and economics. pp. 644-652. Johns Hopkins University Press, Baltimore. 1147 p.
- Additional ReferencesLegend: View Online Publication
Do you know of a citation we're missing?
- Adelman, E.B. 1979. A survey of the nongame mammals in the Upper Rattlesnake Creek drainage of western Montana. M.S. thesis. University of Montana, Missoula. 129 pp.
- Bachman, D., G. Gadwa and C. Groves. 1990. A winter survey for wolverines (Gulo gulo) on the Sawtooth and Challis National Forests, Idaho. Natural Heritage Section, Idaho Department of Fish and Game. 30 p.
- Banci, V. 1987. Ecology and behaviour of Wolverine in Yukon. M.Sc. Thesis. Burnaby, BC: Simon Fraser University. 178 p.
- Banci, V. 1994. Wolverine. Pp. 99-127. IN: Ruggiero, L. F., K. B. Aubry, S. W. Buskirk, L. J. Lyon, amd W. J. Zielinski (eds.), The scientific basis for conserving forest carnivores, American marten, fisher, lynx, and wolverine in the western United States. U.S.D.A, Forest Serv. Rocky Mountain Forest and Range Exper. Stat. Gen. Tech. Report RM-254. 184 pp.
- Banci, V. and A. Harestad. 1988. Reproduction and natality of wolverine (Gulo gulo) in Yukon. Annales Zoologici Fennici 25:265-270.
- Banci, V. and A.S. Harestad. 1990. Home range and habitat use of wolverines in Yukon, Canada. Holarctic Ecology 13:195-200.
- Butts, T.W. 1992. Wolverine (Gulo gulo) biology and management: a literature review and annotated bibliography. USDA Forest Service Northern Region: Threatened, Endangered, and Sensitive Program. 106 p. + appendix.
- Carroll, K.A. 2019. Wolverine habitat quality, connectivity, and prioritization at the landscape scale. Ph.D. Dissertation. Bozeman, MT: Montana State University. 138 p.
- Casebeer, R. L., M. J. Rognrud and S. M. Brandberg. 1950. Rocky Mountain goats in Montana. Montana Fish and Game Comm., Wildl. Rest. Div. Bull. No.5. 107 pp.
- Chadwick, D.H. 1973. Mountain goat ecology--logging relationships in the Bunker Creek drainage of western Montana. M.S. thesis. University of Montana, Missoula. 262 pp.
- Cross, James, and Richard P. Weckwerth, 1975, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-6, Job No. II-1, July 1, 1974 - June 30, 1975.
- Cross, James, and Richard P. Weckwerth, 1976, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-7, Job No. II-1, July 1, 1975 - June 30, 1976.
- Cross, James, and Richard P. Weckwerth, 1977, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-8, Job No. II-1, July 1, 1976 - June 30, 1977.
- Cross, James, and Richard P. Weckwerth, 1978, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-9, Job No. II-1, July 1, 1977 - June 30, 1978.
- Cross, James, and Richard P. Weckwerth, 1983, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-14, Job No. II-1, July 1, 1982 - June 30, 1983.
- Deems, E.F., Jr. and D. Pursley (eds). 1978. North American furbearers: their management, research and harvest status in 1976. Int. Assoc. Fish and Wildlife Agencies and University of Maryland. 171 p.
- Flath, D.L. 1979. Nongame species of special interest or concern: Mammals, birds, reptiles, amphibians, fishes. Wildlife Division, Montana Department of Fish and Game. Helena, MT.
- Flathead National Forest. U.S. Forest Service., 1993, Wildlife landscape evaluation, Swan Valley. Draft Report.
- Foresman, K.R. 2001. The wild mammals of Montana. American Society of Mammalogists, Special Publication Number 12. Lawrence, KS. 278 pp.
- Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
- Gardner, C.L. 1985. The ecology of wolverines in southcentral Alaska. M.Sc. Thesis. Fairbanks, AK: University of Alaska. 82 p.
- Gipson, P.S. 1979. Ecology of wolverines in an Arctic ecosystem. National Geographic Society Research Report 1979 Grant 2110:253-263.
- Groves, C. 1987. Distribution of the wolverine (Gulo gulo) in Idaho, 1960-1987. Idaho Natural Heritage Program, Idaho Department of Fish and Game. 24 p.
- Groves, C. and G. Gadwa. 1989. Status survey for wolverines (Gulo gulo) on the Sawtooth National Forest and adjacent areas. Boise, ID: Idaho Department of Fish and Game. Cooperative Challenge Cost Share Project. 49 p.
- Haglund, B. 1966. Winter habits of lynx (Lynx Lynx L.) and wolverine (Gulo gulo L.) as revealed by tracking in the snow. Viltrevy 4:245-285.
- Hash, H.S. and M.G. Hornocker. 1980. Immobilizing wolverines with ketamine hydrochloride. Journal of Wildlife Management 44(3):713-715.
- Hoak, J.H., J.L. Weaver and T.W. Clark. 1980. Wolverines in western Wyoming. Northwest Science 56(3):159-161.
- Hoffmann, R.S. and D.L. Pattie. 1968. A guide to Montana mammals: identification, habitat, distribution, and abundance. Missoula, MT: University of Montana. 133 p.
- Holbrook, J.D., J.R. Squires, L.E. Olson, N.J. DeCesare, and R.L. Lawrence. 2017. Understanding and predicting habitat for wildlife conservation: the case of Canada Lynx at the range periphery. Ecosphere 8(9):e01939.
- Jackson, H.H. 1961. Genus Gulo pallas, Wolverines. pp. 357-362. In: Mammals of Wisconsin. Madison, WI: University of Wisconsin Press. 520 p.
- Johnson, R. E. 1977. An historical analysis of wolverine abundance and distribution in Washington. Murrelet 58:13-16.
- Joslin, Gayle, and Heidi B. Youmans. 1999. Effects of recreation on Rocky Mountain wildlife: a review for Montana. [Montana]: Montana Chapter of the Wildlife Society.
- Koehler, G.M. and J.D. Brittell. 1990. Managing spruce-fir habitat for lynx and snowshoe hares. Journal of Forestry 88:10-14.
- Koehler, G.M., M.G. Hornocker and H.S. Hash. 1980. Wolverine marking behavior. The Canadian Field Naturalist 94:339-341.
- Koehler, G.M., M.G. Hornocker, and H.S. Hash. 1979. Lynx movements and habitat use in Montana. Canadian Field-Naturalist 93:441-442.
- Kosterman, M.K., J.R. Squires, J.D. Holbrook, D.H. Pletscher, and M. Hebblewhite. 2018. Forest structure provides the income for reproductive success in a southern population of Canada Lynx. Ecological Applications 28(4):1032-1043.
- Magoun, A. J. 1987. Summer and winter diets of wolverines, Gulo gulo, in arctic Alaska. Canadian Field-Naturalist 101:392-397.
- Maj, M., and E. O. Garten. 1994. Fisher, lynx, wolverine: Summary of distribution information. Pp. 169-175. IN: Ruggiero, L. F., K. B. Aubry, S. W. Buskirk, L. J. Lyon, amd W. J. Zielinski (eds.), The scientific basis for conserving forest carnivores, American marten, fisher, lynx, and wolverine in the western United States. U.S.D.A, Forest Serv. Rocky Mountain Forest and Range Exper. Stat. Gen. Tech. Report RM-254. 184 pp.
- Matthews, W.L. and J.E. Swenson. 1982. The mammals of east-central Montana. Proc. Mont. Acad. Sci. 39: 1-13.
- McKay, R. 1991. Biological assessment and inventory plan for the wolverine (Gulo gulo) in the Uinta Mountains. Salt Lake City, UT: Utah Department of Natural Resources. cooperative Challenge Cost Share Project. 58 p.
- Mead, E.M. and J.I. Mead. 1989. Snake Creek burial cave and a review of the Quaternary mustelids of the Great Basin. Great Basin Naturalist 49(2):143-154.
- Newby, F. E. 1952. Distribution and status of the wolverine. Montana Fish and Game Dept.. P-R Quarterly Rep., Oct.-Dec.:76-80.
- Newby, F.E. and J.J. McDougal. 1964. Range extension of the wolverine in Montana. Journal of Mammalogy 45(3):485-487.
- Newby, F.E. and P.L. Wright. 1955. Distribution and status of the wolverine in Montana. Journal of Mammalogy 36(2):248-253.
- Newby, Fletcher E. 1953. Distribution and Status of the Wolverine. Project No. W-49-R-2, Job No. II-F.
- Obbard, M.E., et al. 1987. Furbearer harvests in North America, wolverine pp.1022-1023 In: M. Novak, et al. (eds). Wild furbearer management and conservation in North America, technology, techniques, and harvests. Ontario Ministry of Natural Resources and The Ontario Trappers Association. 1150 p.
- Oechsli, L.M. 2000. Ex-urban development in the Rocky Mountain West: consequences for native vegetation, wildlife diversity, and land-use planning in Big Sky, Montana. M.Sc. Thesis. Montana State University, Bozeman. 73 p.
- Pasitschniak-Arts, M. and S. Lariviere. 1995. Gulo gulo. American Society of Mammalogists, Lawrence, KS. Mammalian Species No. 499:1-10.
- Peterson, Joel, and Michael R. Frisina, 1985, II Upland Game Bird Survey and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-16, Job No. II-3, July 1, 1984 - June 30, 1985.
- Peterson, Joel, and Michael R. Frisina, 1986, II Upland Game Bird Survey and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-17, Job No. II-3, July 1, 1985 - June 30, 1986.
- Peterson, Joel, and Michael R. Frisina, 1987, II Upland Game Bird and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-18, Job No. II-3, July 1, 1986 - June 30, 1987.
- Peterson, R. L. 1966. The mammals of eastern Canada. Oxford University Press. Toronto, Canada. 465 pp.
- Rausch, R.A. and A.M. Pearson. 1972. Notes on the wolverine in Alaska and the Yukon Territory. Journal of Wildlife Management 36(2):249-268.
- Reichel, J.D., D.L. Genter and E. Atkinson. 1992. Sensitive animal species in the Elkhorn and Big Belt Mountains of the Helena National Forest. Unpublished report to the Helena National Forest. Montana Natural Heritage Program, Helena. 158 p.
- Reid, F. 2006. Peterson Field Guide to Mammals of North America, 4th Edition. Houghton Mifflin Company: Boston and New York, 608 pp.
- Roybal, J.P. 1998. Winter mortality in the Northern Yellowstone Elk herd 1988-1990. M.Sc. Thesis. Bozeman, MT: Montana State University. 191 p.
- Ruggiero, L.F., K.B. Aubry, S.W. Buskirk, L.J. Lyon, and W.J. Zielinski. 1994. The scientific basis for conserving forest carnivores: American marten, fisher, lynx, and wolverine in the western United States. Fort Collins, CO: General Technical Report RM-254. U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 184 p.
- Rust, H. J. 1946. Mammals of northern Idaho. J. Mammal. 27(4): 308-327.
- Thompson, L.S. 1982. Distribution of Montana amphibians, reptiles, and mammals. Bozeman: Montana Audubon Council. 24 pp.
- Thompson, Richard W., Western Resource Dev. Corp., Boulder, CO., 1996, Wildlife baseline report for the Montana [Montanore] Project, Lincoln and Sanders counties, Montana. In Application for a Hard Rock Operating Permit and Proposed Plan of Operation, Montanore Project, Lincoln and Sanders Counties, Montana. Vol. 5. Stroiazzo, John. Noranda Minerals Corp., Libby, MT. Revised September 1996.
- Tweten, R.G. 1984. Baseline survey of furbearing mammals within the South Fork drainage Sun River, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 63 p.
- USDI National Park Service., 2000, Bison Management for the State of Montana and Yellowstone National Park. Final Environmental Impact Statement for the Interagency Bison Management Plan for the State of Montana and Yellowstone National Park. Vol. I. August 2000.
- van Zyll de Jong, C.G. 1975. The distribution and abundance of the wolverine (Gulo gulo) in Canada. Can. Field-Nat. 89(4):431-437.
- Vinkey, R.S. 2003. An evaluation of fisher (Martes pennanti) introductions in Montana. M.S. Thesis. University of Montana. Missoula, MT. 97 pp.
- Weckwerth, R. P. 1957. The relationship between the marten population and the abundance of small mammals in Glacier National Park. M.S. thesis. University of Montana, Missoula. 76 pp.
- Whitman, J.S., W.B. Ballard, and C.L. Gardner. 1986. Home range and habitat use by wolverines in southcentral Alaska. Journal of Wildlife Management 50(3):460-463.
- Wolverine (Gulo gulo) pp. 153-156 In: Deems, E.F. and D. Pursley (eds). 1983. North American furbearers: a contemporary reference. International Association of Fish and Wildlife Agencies with the MD Dept. of Natural Resources-Wildlife Administration. 217 p.
- Wright, P.L. and R. Rausch. 1955. Reproduction in the wolverine, Gulo gulo. Journal of Mammalogy 36(3):346-355.
- Youngman, P.M. 1975. Mammals of the Yukon Territory. Ottawa, Ontario: Publications in Zoology, No. 10. National Museums of Canada. 192 p.
- Zielinski, W.J. and T.E. Kucera. 1995. American marten, fisher, lynx, and wolverine: survey methods for their detection. Berkeley, CA: USDA Forest Service Pacific SW Research Station, Gen. Tech. Rep. PSW-GTR-157. 163 p.
- Additional Sources of Information Related to "Mammals"