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Beaver - Castor canadensis
Other Names:  Ksik stakii (Blackfeet), North American Beaver

Image Copyright and Usage Information
Native Species

Global Rank: G5
State Rank: S5
(see State Rank Reason below)


Agency Status
USFWS:
USFS:
BLM:


 

External Links


Listen to an Audio Sample
Copyright Jeff Rice, all rights reserved. Audio file courtesy of the Acoustic Atlas at Montana State University (www.acousticatlas.org)
State Rank Reason (see State Rank above)
Species is relatively common within suitable habitat and widely distributed across portions of the state
  • Details on Status Ranking and Review
    Beaver (Castor canadensis) Conservation Status Review
    Review Date = 05/03/2018
    Range Extent

    ScoreG - 200,000-2,500,000 km squared (about 80,000-1,000,000 square miles)

    Comment380,531 square Kilometers from Natural Heritage Program range maps

    Long-term Trend

    ScoreE - Relatively Stable (±25% change)

    CommentWhile the species has undergone significant declines due to trapping, populations have recovered across its historic range.

    Short-term Trend

    ScoreF - Increasing. Increase of >10% in population, range, area occupied, and/or number or condition of occurrences

    CommentSpecies continues to actively colonize areas from which it was extirpated

    Threats

    ScoreH - Unthreatened. Threats if any, when considered in comparison with natural fluctuation and change, are minimal or very localized, not leading to significant loss or degradation of populations or area even over a few decades’ time. (Severity, scope, and/or immediacy of threat considered Insignificant.)

    CommentSpecies is harvested/controlled, local populations suffer from outbreaks of tularemia. Species has not recolonized all areas from which it was extirpated, but is expanding in both range and population. Existing populations secure and stable.

    Intrinsic Vulnerability

    ScoreB - Moderately Vulnerable. Species exhibits moderate age of maturity, frequency of reproduction, and/or fecundity such that populations generally tend to recover from decreases in abundance over a period of several years (on the order of 5-20 years or 2-5 generations); or species has moderate dispersal capability such that extirpated populations generally become reestablished through natural recolonization (unaided by humans).

    CommentModerately Vulnerable. Species exhibits moderate age of maturity, frequency of reproduction, and/or fecundity such that populations generally tend to recover from decreases in abundance within 5-20 years or 2-5 generations. Species has good dispersal ca

    Environmental Specificity

    ScoreD - Broad. Generalist. Broad-scale or diverse (general) habitat(s) or abiotic and/or biotic factors are used or required by the species, with all key requirements common in the generalized range of the species in the area of interest. If the preferred food(s) or breeding/nonbreeding microhabitat(s) become unavailable, the species switches to an alternative with no resulting decline in numbers of individuals or number of breeding attempts.

    CommentRequires lentic or lotic waterbodies but this requirement aside, the species is found in a wide diversity of ecosystems and shows remarkable flexibly in its life history

    Raw Conservation Status Score

    Score 3.5 + 0 (geographic distribution) + 0.5 (environmental specificity) + 0.25 (short-term trend) + 1 (threats) = 5.25

 
General Description
The largest rodent in North America north of Panama. On land is a large, clumsy, hump-backed animal. In the water, becomes sleek and torpedo-shaped. Propels itself with powerful webbed hind feet. Beavers use their large dorsally flattened, scale-covered tail to maneuver in water. Slapping the tail on water surface is used as a signal of alarm. Split nail on the second hind toe is used for grooming. Incisors are large and continually growing. Fur is rich brown with black to reddish guard hairs. Under-fur is soft and extremely dense with excellent insulating qualities. Both sexes have a pair of anal glands and castor sacs located ventrally. Beavers emit anal gland secretions year round. Total length: 34 to 40 inches. Weight: 30 to 60 pounds. Builds stick and mud dams across streams. Mostly nocturnal. May build large conical houses at the edge of a lake or burrow into the bank for a den along rivers. Beaver life is based on a family unit consisting of a pair of adults, yearlings, and kits. The information in next section is from Hill (1982), Miller (1983), and Novak (1987a), unless otherwise indicated. The beaver is the largest North American rodent; most adults weigh 16 to 23 kg (35 to 50 pounds), rarely 32 to 38 kg (70 to 85 pounds) or more. Kits weigh about 0.5 kg or a little less (1 pound) at birth Hill (1982), when they are about 38 cm (15 inches) long Schwartz and Schwartz (1981). The relationship between beaver age and mass is roughly as follows: less than 0.5 years old, less than 5 kg (11 pounds); 0.5 to 1.5 years old, 6 to 11 kg (13 to 24 pounds); 1.5 to 2.5 years old, 10 to 13 kg (22 to 29 pounds); greater than 2.5 years old, greater than 14 kg (31 pounds). The preceding weight-age data are often accurate, but it must be realized that weight is influenced by various factors other than age. The total length of a beaver varies with age as follows: yearlings are 26 to 34 inches; adults of 2 to 3 years, 35 to 40 inches; older adults, 47+ inches (maximum about 4.5 feet) in the warmer months and generally ends at age 4 to 5 years. Pelage color varies geographically from yellowish-brown to black (this range of color may be found in a single watershed). There is one annual molt. The pelt is prime from late fall to early spring. The sexes are difficult to distinguish externally. Hodgen (1978) provided information on sexing beavers by their behavior in a live trap. See Novak (1987a) for additional references on sexing methods.

Diagnostic Characteristics
River Otter - tail covered with fur. Muskrat - smaller, tail slender. An inexperienced observer could mistake a Muskrat (Ondatra zibethicus), Round-tailed Muskrat (Neofiber alleni) or a Nutria (Myocastor coypus) for a Beaver, but these other rodents do not have a broad flattened tail.

Species Range
Montana Range Range Descriptions

Native

Western Hemisphere Range

 


Observations in Montana Natural Heritage Program Database
Number of Observations: 10368

(Click on the following maps and charts to see full sized version) Map Help and Descriptions
Relative Density

Recency

 

(Observations spanning multiple months or years are excluded from time charts)



Habitat
Occupies a wide variety of habitats in North America. Water and associated woody vegetation are the most essential components of beaver habitat. Ideal beaver living sites include ponds, small lakes, meandering streams, and rivers.

Ecological Systems Associated with this Species
  • Details on Creation and Suggested Uses and Limitations
    How Associations Were Made
    We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
    1. Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
    2. Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
    3. Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
    4. Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
    Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.  In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.  However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if point observations were associated with that system.  Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.  The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.  If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.

    Suggested Uses and Limitations
    Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.  These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: mtnhp.org/requests) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.  Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.  Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.  Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).  Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.

    Literature Cited
    • Adams, R.A.  2003.  Bats of the Rocky Mountain West; natural history, ecology, and conservation.  Boulder, CO: University Press of Colorado.  289 p.
    • Dobkin, D. S.  1992.  Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34.  Missoula, MT.
    • Foresman, K.R.  2012.  Mammals of Montana.  Second edition.  Mountain Press Publishing, Missoula, Montana.  429 pp.
    • Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998.  Montana atlas of terrestrial vertebrates.  Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT.  1302 p.
    • Hutto, R.L. and J.S. Young.  1999.  Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32.  72 p.
    • Maxell, B.A.  2000.  Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species.  Report to U.S. Forest Service Region 1.  Missoula, MT: Wildlife Biology Program, University of Montana.  161 p.
    • Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath.  2004.  Amphibians and reptiles of Montana.  Missoula, MT: Mountain Press Publishing Company. 262 p.

Food Habits
Beavers are strict herbivores. Beavers eat a variety of woody and herbaceous species. Willows, mountain alder, and aspen are important foods. Will also consume herbaceous vegetation during summer. Much of the woody vegetation beavers cut is not used for food. Caches woody vegetation near shore for winter food.

Reproductive Characteristics
Breeds from January through March. Two to four young are born after a 105- to 107-day gestation period. One litter per year is produced. Two-year-olds are eventually driven from or leave the parental home.

Management
Beaver management and the role of beaver in riparian restoration in Montana is extensively addressed by Ritter et al. (2023), including sections on conflict management, land management changes needed to address beave restoration and management, beaver mimicry, encouraging natural colonization, and beaver transplantation.


References
  • Literature Cited AboveLegend:   View Online Publication
    • Hill, E.P. 1982. Beaver. Pages 256-281 in J.A. Chapman and G.A. Feldhamer, editors. Wild Mammals of North America: biology, management, and economics. Johns Hopkins University Press, Baltimore. 1147pp.
    • Hodgdon, H.E. 1978. Social dynamics and behavior within an unexploited beaver (Castor canadensis) population. Ph.D. dissertation. University of Massachusetts, Amherst. 292 p.
    • Miller, J.E. 1983. Beavers. Prevention and control of wildlife damage. Cooperative Extension Service, Institute of Agricultural and Natural Resources, University of Nebraska, Lincoln.
    • Novak, M. 1987. Beaver. Pp. 282-312 in: M. Novak, J.A. Baker, M.E. Obbard, and B. Malloch, eds. Wild furbearer management and conservation in North America. Ministry Nat. Res., Ontario. 1150 pp.
    • Ritter, T., M. McGree, D. Schmetterling, C. Gower, V. Boccadori. 2023. Beavers and their role in riparian restoration in Montana. Montana Fish, Wildlife & Parks White Paper, Version 1.0. 165 p.
    • Schwartz, C.W. and E.R. Schwartz. 1981. The wild mammals of Missouri. Revised edition. University of Missouri Press and Department of Conservation. Columbia and London. 356 pp.
  • Additional ReferencesLegend:   View Online Publication
    Do you know of a citation we're missing?
    • Adams, A.K. 1953. Some psycho-chemical effects of beaver dams upon Michigan trout streams in the Watersmeet area. Ph.D. dissertation, University of Michigan, Ann Arbor.
    • Adelman, E.B. 1979. A survey of the nongame mammals in the Upper Rattlesnake Creek drainage of western Montana. M.S. thesis. University of Montana, Missoula. 129 pp.
    • Aldous, M. C. 1955. A tool for beaver dam removal. J. WildI. Manage. 19: 153-154.
    • Aldous, S.E. 1938. Beaver food utilization studies. Journal of Wildlife Management 2: 215-222.
    • Aleksiuk, M. 1968. Scent-mound communication, territoriality, and population regulation in beaver (Castor canadensis Kuhl). Journal of Mammalogy 49(4): 759-762.
    • Aleksiuk, M. 1970. The seasonal food regime of arctic beavers. Ecology 51: 264-270.
    • Aleksiuk, M. and I.M. Cowan. 1969. Aspects of energy expenditure in the beaver (Castor canadensis Kuhl) at the northern limit of its distribution. Canadian Journal of Zoology 47(4): 471-481.
    • Aleksiuk, M. and I.M. Cowan. 1969. The winter metabolic depression in arctic beavers (Castor canadensis Kuhl) with comparisons to California beavers. Canadian Journal of Zoology 47(5): 965-979.
    • Allen, A.W. 1983. Habitat suitability index models: beaver. U.S. Fish and Wildlife Service, FWS/OBS-82/10.30. 20pp.
    • Allen, T.D.H. 2011. Wildlife guards for deer and Black Bear. M.Sc. Thesis. Bozeman, Montana: Montana State University. 71 p.
    • Allred, M. 1986. Beaver behavior. Happy Camp, California: Naturegraph Publishers, Inc. 110p.
    • Alt, Kurt, 1984, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-15, Job No. 2, July 1, 1983 - June 30, 1984.
    • Alt, Kurt, John E. Firebaugh, Lyn S. Nielsen, and Robert Henderson, 1985, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-16, Job No. 2, July 1, 1984 - June 30, 1985.
    • Alt, Kurt, John E. Firebaugh, Lyn S. Nielsen, and Robert Henderson, 1987, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-18, Job No. 2, July 1, 1986 - June 30, 1987.
    • Alt, Kurt., John E. Firebaugh, Lyn S. Nielsen, and Robert Henderson, 1986, Statewide Wildlife Survey and Inventory. II Game Bird and Fur (Small Game) Survey and Inventory. Region Two Survey and Inventory. W-130-R-17, Job No. 2, July 1, 1985 - June 30, 1986.
    • Anaconda Minerals Company, and Camp, Dresser & McKee. 1981. Anaconda Stillwater Project 6-month environmental baseline report. CDM Project No. 3139. Vol. I Appendix. Jan. 15, 1981.
    • Anonymous. 1946. Trapping and transplanting beaver. Montana Fish and Game Comm. Biennial Report.
    • Anonymous. 1989. Improved drain for beaver ponds. The Habitat Express, U.S. Forest Service, Ogden, Utah.
    • Apple, L.L. 1983. The use of beavers in riparian/aquatic habitat restoration in a cold desert, gully cut stream system: a case history. Proceedings of the 18th Annual Meeting American Fisheries Society, Laramie, Wyoming: 29-35.
    • Apple, L.L. 1984. Riparian habitat restoration in cold desert, gully-cut stream systems: an innovative, cost effective, ecological approach. In: Transactions of the 49th North American wildlife and natural resources conference; 1984 March 23-28; Boston, MA. Washington, DC: Wildlife Management Institute (Unpaged). Poster Session.
    • Apple, L.L. 1985. Riparian habitat restoration and beaver. In: Riparian ecosystems and their management: reconciling conflicting uses. Rocky Mountain Forest and Range Experiment Station, USDA Forest Service, General Technician Report RM-120.
    • Apple, L.L., B.H. Smith, J.D. Dunder, and B.W. Baker. 1984. The use of beavers for riparian/aquatic habitat restoration of cold desert, gully-cut stream systems in southwestern Wyoming. In: Proceedings, American Fisheries Society/Wildlife Society joint chapter meeting; 1984 February 8-10; Logan, UT. (Publisher and city unknown): 123-130.
    • Arner, D.H. 1963. Production of duck food in beaver ponds. Journal of Wildlife Management 27: 76-81.
    • Arner, D.H. and G.R. Hepp. 1989. Beaver pond wetlands: a southern perspective. In: L.M. Smith et al., editors. Habitat management for migrating waterfowl in North America. Texas Tech University Press, Lubbock: 117-128.
    • Arner, D.H. and J.S. Dubose. 1980. The impact of the beaver on the environment and economics in the southeastern United States. Proceedings of International Wildlife Conference 14: 241-247.
    • Arner, D.H., J. Baker, D. Wesley, and B. Herring. 1969. An inventory and study of beaver impounded water in Mississippi. Proc. Annu. Conf. Southeastern Assoc. Game and Fish Commission 23: 110-128.
    • Atwater, M. M. 1939. South Fork (Montana) beaver survey. J. Wildl. Manage. 4: 100-103.
    • Atwater, M.M. 1940. South Fork (Montana) beaver survey: 1939. Journal of Wildlife Management 4: 100-103.
    • Avery, E.L. 1983. A bibliography of beaver, trout, and wildlife, and forest relationships. Wisconsin Department of Natural Resources Technical Bulletin No. 137. 23pp.
    • Avery, E.L. 1992. Effects of removing beaver dams upon a northern Wisconsin brook trout stream. Wisconsin Department of Natural Resources, Bureau of research, Fish Research Section, Study 406, Madison.
    • Bailey, R.W. and R.F. Stephens. 1951. Effects of beavers on fish. West Virginia Conservation, September 1951: 1-7.
    • Barnes, D. M. and A. U. Mallik. 2001. Effects of Beaver, Castor canadensis, Herbivory on Streamside Vegetation in a Northern Ontario Watershed. The Canadian Field Naturalist 115: 9-19.
    • Barnes, D.M. and A.U. Mallik. 1996. Use of woody plants in the construction of beaver dams in northern Ontario. Canadian Journal of Zoology 74: 1781-1786.
    • Barnes, D.M. and A.U. Mallik. 1997. Habitat factors influencing beaver dam establishment in a northern Ontario watershed. Journal of Wildlife Management 61: 1371-1377.
    • Barnes, W.J. and E. Dibble. 1986. The effects of beaver in riverbank froest succession. Canadian Journal of Botany 66: 40-44.
    • Basey, J. M. and S. H. Jenkins. 1995. Influences of predation risk and energy maximization on food selection by beavers (Castor Canadensis). Canadian Journal of Zoology 73: 2197-2208.
    • Basey, J.M., S.H. Jenkins, and G.C. Miller. 1990. Food selection by beavers in relation to inducible defenses of Populus tremuloides. Oikos 59: 57.
    • Basey, J.M., S.H. Jenkins, and P.E. Busher. 1988. Optimal central place foraging by beavers: tree-size selection in relation to defensive chemicals of quaking aspen. Oecologia 6: 278.
    • Bates, J.W. 1963. The effects of beaver on stream flow. Departmental Information Bulletin No. 63-13, Federal Aid Completion Report. Utah State Department of Fish and Game.
    • Baxter, R.M. 1977. Environmental effects of dams and impoundments. Ann. Rev. Ecol. Syst. 8: 255-283.
    • Beak Consultants, Inc. 1983. Wildlife. January 1983. In Stillwater Project Environmental Studies. Addendum A, Wildlife. Vol. I. Tech. Report No. 7. 1982.
    • Beard, E.B. 1953. The importance of beaver in waterfowl management at the Seney National Wildlife Refuge. Journal of Wildlife Management 17: 398-436.
    • Beaver Advisory Committee. 2017 (revised). Utah beaver management plan. Salt Lake City, UT: Utah Division of Wildlife Resources, DWR Publication 17-20. 29 p.
    • Beer, J.R. 1955. Movements of tagged beaver. Journal of Mammalogy 19: 492-493.
    • Beier, P. and R.H. Barrett. 1987. Beaver habitat use and impact in Truckee River basin, California. Journal of Wildlife Management 51: 794-799.
    • Bell, J. F., C. R. Owen, W. L. Jellison, G. J. Moore, and E. O. Baker. 1962. Epizootic tularemia in pen-raised beavers, and field trials of vaccines. Am. J. Vet. Res. 23(95): 884-888.
    • Belovsky, G.E. 1984. Summer diet optimization by beaver. American Midland Naturalist 11: 209.
    • Bergerund, A.T. and D.R. Miller. 1977. Population dynamics of Newfoundland beaver. Canadian Journal of Zoology 55: 1480-1492.
    • Bergstrom, D. 1985. Beavers: biologists "rediscover" a natural resource. Forestry Research West (Fort Collins, CO): USDA Forest Service; October: 1-5.
    • Bhat, M., R. Huffaker, et al. 1993. Controlling forest damage by dispersive beaver populations: Centralized optimal management strategy. Ecological Applications 3(3): 518-530.
    • Bilby, R.E. 1981. Role of organic debris dams in regulating the export of dissolved and particulate matter from a forested watershed. Ecology 62: 1234.
    • Bilby, R.E. 1984. Removal of woody debris may affect stream channel stability. Journal of Forestry 82: 609-612.
    • Bilby, R.E. and G.E. Likens. 1980. Importance of organic debris dams in the structure and function of stream ecosystems. Ecology 61(5): 1107-1113.
    • Bishir, J., R.A. Lancia, and H.E. Hodgdon. 1983. Beaver family organization: its implications for colony size. Pages 105-113 in Investigations on Beavers. Edited by G. Pilleri. Institute of Brain Anatomy, Berne, Switzerland.
    • Boddicker, M.L. (undated). Beaver management - a key in western riparianhabitat excellence. Department of Fishery and Wildlife Biology, Colorado State University, Ft. Collins.
    • Bown, R. 1988. Beaver habitat along rivers and resevoirs in central Montana. University of Montana, Missoula, Montana.
    • Boyce, M. 1974. Beaver population ecology in interior Alaska. M.S. Thesis, University of Alaska, Fairbanks. 161pp.
    • Boyce, M. 1981. Beaver life history responses to exploitation. Journal of Applied Ecology 18(3): 749-753.
    • Boyce, M. S. 1981. Habitat ecology of an unexploited population of beavers in interior Alaska. pp. 155-186. In: Chapman, J.A. and D. Pursley (eds). Worldwide Furbearer Conference Proceedings. Volume 1.
    • Bradt, G.W. 1938. A study of beaver colonies in Michigan. Journal of Mammalogy 19: 139-162.
    • Bradt, G.W. 1947. Michigan beaver management. Michigan Department of Conservation, Lansing. 56pp.
    • Brady, C.A. and G.E. Svendsen. 1981. Social behavior in a family of beaver, Castor canadensis. Biology of Behavior 6: 99-114.
    • Brayton, D.S. 1984. The beaver and the stream. Journal of Soil and Conservation 39(2): 108-109.
    • Brazell, R.E. and G.W. Workman. 1977. A preliminary survey on beaver (Castor canadensis) in Canyonlands National Park, Utah. Encyclia. 54(1): 25-27.
    • Brenner, F.J. 1962. Food consumed by beavers in Crawford County, Pennsylvania. Journal of Wildlife Management 26: 104-107.
    • Brooks, R.P. 1977. Induced sterility of the adult female beaver (Castor canadensis) and colony fecundity. M.S thesis, University of Massachusetts, Amherst. 90pp.
    • Brooks, R.P., M.W. Fleming, and J.J. Kennelly. 1980. Beaver colony response to fertility control: evaluating a concept. Journal of Wildlife Management 44: 568-575.
    • Broschart, M.R., C.A. Johnston, and R.J. Naiman. 1989. Predicting beaver colony density in boreal landscapes. Journal of Wildlife Management 53: 929-934.
    • Brown, D.J., W.A. Hubert, and S.H. Anderson. 1996. Beaver ponds create wetland habitat for birds in mountains of southeastern Wyoming. Wetlands 16: 127-133.
    • Brown, J.H. 1995. Organisms as engineers: a useful framework for studying effects on ecosystems. Trends Ecol. Evol. 10: 51-52.
    • Brown, M.K. and G.R. Parsons. 1979. Waterfowl production on beaver flowages in a part of northern New York. New York Fish and Game Journal 26: 142-153.
    • Brown, M.K. and G.R. Parsons. 1982. Reliability of fall aerial censuses for locating active beaver colonies in northern New York. New York Fish and Game Journal 29: 204-206.
    • Brown, R.L. 1960. Fur resource surveys and investigations: beaver population and distribution trends-aerial survey. Montana Fish and Game Department, Project # MONT. W-049-R-09/W.K.P.L.03/JOB B. 112p.
    • Bryant, M.D. 1984. The role of beaver dams as coho salmon habitat in southeastern Alaska streams. Pages 183-192. In: Walton, J.M. and D.D. Houston, editors. Proceedings of the Olympic wild fish conference. Peninsula College, Fisheries Technology Program, Port Angeles, Washington.
    • Buck, C.L. 1939. Pattern correlation of mammalian teeth as a means of identification. M.Sc. Thesis. Bozeman, Montana: Montana State University. 55 p.
    • Buckley, J.L. 1950. The ecology and economics of the beaver (Castor canadensis Kuhl) with a plan for its management on the Huntington Wildlife Forest Station. Ph.D. thesis, State University New York, Syracuse. 251pp.
    • Buckley, J.L. and W.L. Libby. 1955. Growth rates and age determination in Alakan beaver. Transactions of the 20th North American Wildlife Conference, Washington. Pages 495-507.
    • Buech, R. R. 1983. Modification of the bailey live trap for beaver. Wildlife Society Bulletin 11: 66-68.
    • Buech, R.R. 1984. Ontogeny and diurnal cycle of fecal reigestion in the North American beaver (Castor canadensis). Journal of Mammalogy 65: 347-350.
    • Buech, R.R. 1995. Sex differences in behavior of beavers living in near-boreal lake habitat. Canadian Journal of Zoology 73: 2133-2143.
    • Buech, R.R., D.J. Rugg, and N.L. Miller. 1989. Temperature in beaver lodges and bank dens in a near-boreal environment. Canadian Journal of Zoology 67: 1061.
    • Bullock, J.F. and D.H. Arner. 1985. Beaver damage to nonimpounded timber in Mississippi. Southern Journal of Applied Forestry 9: 137-140.
    • Bush, A.O. and W.M. Samuel. 1981. A review of helminth communities in beaver (Castor spp.) with a survey of Castor candensis in Alberta, Canada. In: Chapman, J.A. and D. Pursley, eds. Proceedings, worldwide furbearer conference; 1980 August 8-11; Frostburg, MD. Frostburg, MD: Worldwide Furbearer Conference, Inc.: 678-689.
    • Busher, P.E. 1975. Movements and activities of beavers, Castor canadensis, on Sagehen Creek, California. M.A. thesis, San Francisco State University, California. 86pp.
    • Busher, P.E. 1983. Interactions between beavers in a montane population in California. Acta Zoologica Fennica. 174: 109-110.
    • Busher, P.E. 1987. Population parameters and family composition of beaver in California. Journal of Mammalogy 68: 860.
    • Busher, P.E. 1991. Food caching behavior by the North American beaver, Castor canadensis, in western Massachusetts. Pages 111-114. In: Bobek, B., K. Perzanowski, and W. Regelin (eds), Global Trends in Wildlife Management. Transactions of the 18th IUGB Congress, Swiat Press, Krakow-Warszawa, Poland.
    • Busher, P.E. 1996. Food caching behavior of beavers (Castor canadensis)-selection and use of woody species. American Midland Naturalist 135: 343-348.
    • Busher, P.E. and S.H. Jenkins. 1985. Behavioral patterns of a beaver family in California. Biology of Behavior 10(1): 41-54.
    • Busher, P.E., R.J. Warner, J. Randall, and S.H. Jenkins. 1983. Population density, colony composition, and local movements in two Sierra Nevadan beaver populations. Journal of Mammalogy 64(2): 314-318.
    • Butler, D.R. 1989. The failure of beaver dams and resulting outburst flooding: a geomorphic hazard of the southeastern Piedmont. Geogr. Bull. 31: 29-38.
    • Butler, D.R. 1991. The changing south: the reintroduction of the beaver into the south. Southeastern Geographer 31: 39-43.
    • Butler, D.R. 1991a. Beavers as agents of biogeomorphic change: a review and suggestions for teaching excercises. J. Geogr. 90: 210-217.
    • Butler, D.R. and G.P. Malanson. 1993b. Canadian landform examples-beaver landforms. Can. Geogr. 38: 76-79.
    • Butler, D.R. and G.P. Malanson. 1995. Sedimentation rates and patterns in beaver ponds in a mountain environment. Geomorphology 13: 255-269.
    • Call, M.W. 1970. Beaver pond ecology and beaver-trout relationships in southeastern Wyoming. University of Wyoming, Laramie, Wyoming.
    • Carlsen, T. and R. Northrup. 1992. Canyon Ferry Wildlife Management Area Final Draft Management Plan. March 1992.
    • Casagranda, L.G. 1955. A study of beaver-waterfowl relations in the mountainous area of Beaverhead County, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 33 p.
    • Casagranda, Lloyd G., 1955, Beaver-waterfowl relationships in the Jefferson Basin. W-49-R-4, IId, August 5, 1955. A Study of beaver-waterfowl relations in the mountainous area of Beaverhead County, Montana.(thesis)
    • Chabreck, R.H. 1958. Beaver-forest relationships in St. Tammany Parish, Louisiana. Journal of Wildlife Management 22: 179-183.
    • Chaney, E., W. Elmore, et al. 1990. Livestock grazing on western riparian areas. U.S. Environmental Protection Agency, Washington , D.C.
    • Churchill, J.E. 1980. Beaver are killing our trout streams. Trout 21(4): 22-25.
    • Cirmo, C.P. 1993. Beaver pond biogeochemistry: acid neutralizing capacity generation in a headwater wetland. Wetlands 13(4): 277-292.
    • Clements, C. 1991. Beavers and riparian ecosystems. Rangelands 13(6): 277-279.
    • Cliff, E.P. 1936. The beaver, empire builder and conservationist. Utah Juniper 7: 23-26.
    • Clifford, H. F., G. M. Wiley, and R. J. Casey. 1993. Macroinvertebrates of a beaver-altered boreal stream of Alberta, Canada, with special reference to the fauna on the dams. Canadian Journal of Zoology 71: 1439-1447.
    • Coles, R.W. 1966. Thermoregulation of beaver. Ph.D. Dissertation, Harvard University, Cambridge, Massachusetts.
    • Collen, P. 1995. The reintroduction of beaver (Castor fiber L.) to Scotland: an opportunity to promote the development of suitable habitat. Scottish Forestry 49: 206-217.
    • Collen, P. and R. J. Gibson. 2001. The general ecology of beavers (Castor spp.), as related to their influence on stream ecosystems and riparian habitats, and the subsequent effects on fish - a review. Reviews in Biology and Fisheries 10: 439-461.
    • Collins, T. 1993. The role of the beaver in riparian habitat management. Habitat Extension Bulletin No. 38, Wyoming Game and Fish Department, Cheyenne.
    • Collins, T.C. 1976a. Population characteristics and habitat relationships of beaver, Castor canadensis, in northwest Wyoming. Ph.D. thesis, University of Wyoming, Laramie. 188pp.
    • Collins, T.C. 1976b. Stream flow effects on beaver populations in Grand Teton National Park. In: Proceedings, first conference on scientific research in the National Parks; 1976 November 9-13; New Orleans, LA. Arlington, VA: American Institute of Biological Sciences: 5(1): 349-352.
    • Compton, H.O., 1976, II Game Birds and Fur Survey and Inventory - Region 5. Statewide Wildlife Survey and Inventory. W-130-R-7, II, 5, July 1, 1975 - June 30, 1976.
    • Confluence Consulting Inc. 2010. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT.
    • Confluence Consulting Inc. 2011. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT.
    • Confluence Consulting Inc. 2012. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT.
    • Confluence Consulting Inc. 2013. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT.
    • Constan, Kerry, 1986, Game Birds and Fur (Small Game) Survey and Inventory (Fur Survey Segment). Statewide Wildlife Survey and Inventory. W-130-R-17, II-4, July 1, 1985 - June 30, 1986.
    • Correll, D.L. 2000. Beaver pond biogeochemical effects in the Maryland coastal plain. Biogeochemistry 49(3): 217-239.
    • Courcelles, R. and R. Nault. 1983. Beaver programs in the James Bay area, Quebec, Cananda. Acta Zoologic Fennica 174: 129-131.
    • Cowan, I.M., A.J. Wood, and W.D. Kitts. 1957. Feed requirements of deer, beaver, bear, and mink for growth and maintenance. Trans. N. Am. Wildl. Conf. 22: 179-188.
    • Cowardin, L.M., V. Carter, F.C. Golet, and E.T. LaRoe. 1979. Classification of wetlands and deepwater habitats of the United States. Washington, D.C.: U.S. Fish and Wildlife Service. 103pp.
    • Cowell, D.W. 1984. The Canadian beaver, Castor canadensis, as a geomorphic agent in karst terrain. The Canadian Field-Naturalist 98(2): 227-230.
    • Crawford, H.S., R.G. Hooper, and R.F. Harlow. 1976. Woody plants selected by beavers in the Appalachian Ridge and Valley Province. USDA Forest Service Research Paper NE-346.
    • Crawford, J.S. 1978. The subarctic beaver. Alaska 44: 40-41.
    • Cross, James, and Richard P. Weckwerth, 1971, Upland Game Bird (Fur Survey Inventory). Wildlife Investigations, Region One. W-130-R-2, Job No. II-1, July 1, 1970 - June 30, 1971.
    • Cross, James, and Richard P. Weckwerth, 1972, Upland Game Bird (Fur Survey Inventory). Wildlife Investigations, Region One. W-130-R-3, Job No. II-1, July 1, 1971 - June 30, 1972.
    • Cross, James, and Richard P. Weckwerth, 1974, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-5, Job No. II-1, July 1, 1973 - June 30, 1974.
    • Cross, James, and Richard P. Weckwerth, 1977, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-8, Job No. II-1, July 1, 1976 - June 30, 1977.
    • Cross, James, and Richard P. Weckwerth, 1978, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-9, Job No. II-1, July 1, 1977 - June 30, 1978.
    • Cross, James, and Richard P. Weckwerth, 1980, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-11, Job No. II-1, July 1, 1979 - June 30, 1980.
    • Cross, James, and Richard P. Weckwerth, 1982, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-13, Job No. II-1, July 1, 1981 - June 30, 1982.
    • Cross, James, and Richard P. Weckwerth, 1983, Upland Game Bird (and Fur Survey) Inventory. Wildlife Investigations, Region One. W-130-R-14, Job No. II-1, July 1, 1982 - June 30, 1983.
    • Culp, J.M. and R.W. Davies. 1985. Responses of benthic macroinvertebrate species to manipulation of interstitial detritus in Carnation Creek, British Columbia. Canadian Journal of Fisheries and Aquatic Sciences 42: 139-146.
    • Cunningham, J.M., A.J.K. Calhoun, and W.E. Glanz. 2002. The effect of beaver on the spatial and temporal distribution of pond-breeding amphibian species. Society for Conservation Biology, 2002 Annual Meeting: 3-4.
    • Currier, A., W.D. Kitts, and I.M. Cowan. 1960. Cellulose digestion in the beaver (Castor canadensis). Canadian Journal of Zoology 38: 1109-1116.
    • Curry-Lindahl, K. 1967. The beaver, Castor fiber Linnaeus, 1758 in Sweden - extermination and reappearance. Acta Theriologica 12: 1-15.
    • Cutting, K.A., J.M. Ferguson, M.L. Anderson, K. Cook, S.C. Davis, and R. Levine. 2018. Linking beaver dam affected flow dynamics to upstream passage of Arctic Grayling. Ecology and Evolution 2018; 8:12905-12917.
    • Dahm, C.N. and J.R. Sedell. 1986. The role of beaver on nutrient cycling in streams. Journal of the Colorado-Wyoming Academy of Science 18(1): 32.
    • D'Aulaire, E. and O. D'Aulaire. 1973. The beaver is back. National Wildlife 11(4): 10-13.
    • Davis, J.R. 1984. Movement and behavior patterns of beaver in the Piedmont of South Carolina. M.S. thesis, Clemson University, Clemson, South Carolina. 83pp.
    • Davis, J.R. and D.C. Guynn, Jr. 1993. Activity and habitat utilization of beaver colonies in South Carolina. Proc. Ann. Conf. Southeast Fish and Wildl. Agencies 47: 299-310.
    • Davis, J.R., A.F. von Recum, D.D. Smith, and D.C. Guynn, Jr. 1984. Implantable telemetry in beaver. Wildlife Society Bulletin 12: 322-324.
    • Deems, E.F., Jr. and D. Pursley (eds). 1978. North American furbearers: their management, research and harvest status in 1976. Int. Assoc. Fish and Wildlife Agencies and University of Maryland. 171 p.
    • Denney, R.N. 1952. A summary of North American beaver management, 1946-1948. Colorado Game and Fish Department, Curr. Rep. 28. 58pp.
    • Dennington, M. and B. Johnson. 1974. Studies of beaver habitat in the Mackenzie Valley and northern Yukon. Report 74-39. Ottawa, ON: Canadian Wildlife Service, Department of the Environment. 172p.
    • Devito, K.J. and P.J. Dillon. 1993. Importance of runoff and winter anoxia to the P and N dynamics of a beaver pond. Canadian Journal of Fish and Aquatic Sciences 50: 2222-2234.
    • Dice, L.R. 1923. Mammal associations and habitats of the Flathead Lake Region, Montana. Ecology 4(3): 247-260.
    • Dickinson, N.R. 1971. Aerial photographs as an aid in beaver management. New York Fish and Game Journal 18: 57-61.
    • Dieter, C.D. 1987. Habitat use by beaver along the Big Sioux River. M.S. Thesis, South Dakota State University, Brookings. 60p.
    • Dieter, C.D. and T.R. McCabe. 1989. Factors influencing beaver lodge-site selection on a prairie river. American Midland Naturalist 122: 408-411.
    • Donkor, N. T. and J. M. Fryxell. 1999. Impact of beaver foraging on structure of lowland boreal forests of Algonquin Provincial Park, Ontario. Forest Ecology and Management 118: 83-92.
    • Donkor, N.T. and J.M. Fryxell. 2000. Lowland boreal forests characterization in Algonquin Provincial Park relative to beaver (Castor canadensis) foraging and edaphic factors. Plant Ecology 148(1): 1-12.
    • Dood, A.R. 1980. Terry Badlands nongame survey and inventory final report. Montana Department of Fish, Wildlife, and Parks and Bureau of Land Management, Helena, MT. 70 pp.
    • Dorn, R.D. 1969. Big game research: relations of moose, cattle, and willows in southwestern Montana. Montana Fish and Game Department, Project # MONT. W-098-R-09/W.K.P.L. B/Job 13/PT1; MONT. W-073-R-14/W.K.P.L. B/JOB 13/PT1. 79p.
    • Doucet, D.M. and J.M. Fryxell. 1993. The effect of nutritional quality on forage preference by beavers. Oikos 67: 201-208.
    • Douglass, Richard J., DMI Ecological Research, Butte, MT., 1995, Small animals potentially living in the proposed Cottonwood Mining Area. August 1995. In Gem River Corporation Application for Operating Permit and Plan of Operations. Marc I Mine, Dry Cottonwood Creek, Deer Lodge County, Montana. Vol. 2, Apps. App. L. No date.
    • Dudley, T. and N.H. Anderson. 1982. A survey of invertebrates associated with wood debris in aquatic habitats. Melanderia No. 39: 1-21.
    • Dugmore, A.R. 1914. The romance of the beaver: being the history of the beaver in the Western Hemisphere. London: William Heinemann.
    • Duncan, S.L. 1984. Leaving it to beaver. Environment 26: 41.
    • Dunning, J.B., B.J. Danielson, et al. 1992. Ecological processes that affect populations in complex landscapes. Oikos 65: 169-175.
    • Dyck, A.P. and R.A. MacArthur. 1993. Daily energy requirements of beaver (Castor candensis) in a similated winter microhabitat. Canadian Journal of Zoology 71: 2131-2135.
    • Easter-Pilcher, A. 1990. Cache size as an index to beaver colony size in northwestern Montana. Wildlife Society Bulletin 18(2): 110-113.
    • Easter-Pilcher, A.L. 1987. Forage utilization, habitat selection and population indices of beaver in northwestern Montana. University of Montana, Missoula, Montana.
    • ECON, Inc. (Ecological Consulting Service), Helena, MT., 1976, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1976. Proj. 135-85-A. December 31, 1976.
    • Edwards, J.K. and D.C. Guynn, Jr. 1984. Utilization of woody vegetation by beaver within the South Carolina Piedmaont. Department of Forestry, Clemson University, South Carolina. Forestry Bulletin No. 42: 1-6.
    • Enck, J.W., P.G. Bishop, T.L. Brown, and J.E. Lamendola. 1992. Beaver related attitudes, experiences, and knowledge of key stakeholders in wildlife management unit 21. HDRU Series No. 92-7. Human Dimensions Research Unit, Department of Natural Resources, Cornell University, Ithica, New York.
    • Eng, R.L. 1976. Wildlife Baseline Study [for West Fork of the Stillwater and Picket Pin drainages]
    • Engelhart, A. and D. Muller-Schwartze. 1995. Responses of beaver (Castor canadensis KUHL) to predator chemicals. Journal of Chemical Ecology 21(9): 1349-1362.
    • Erickson, Glenn, 1977, Fur Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-7, II-4, July 1, 1975 - June 30, 1976.
    • Ermala, A., M. Helminen, and S. Lahti. 1989. Some aspects of the occurrence, abundance, and future of the Finnish beaver population. Suomen Riista 35: 108-118.
    • Ermer, D.C. 1973. Upstream changes in fish populations following impoundment of Sagehen Creek, California. Transactions of the American Fisheries Society 102: 626-629.
    • Ermer, E.M. 1980. Beaver management studies in Region 9. New York Federal Aid in Fish and Wildlife Restoration Project W-1139-D (annual report for Job 1-7). New York State Department of Environmental Conservation.
    • Ermer, E.M. 1984. Analysis of benefits and management costs associated with beaver in western New York. New York Fish and Game Journal 31(2): 119-132.
    • Ermer, E.M. 1988. Managing beaver in New York. The Conservationist (State of New York, Department of Environmental Conservation), March-April, pp. 36-39.
    • Euch, R.R. 1985. Beaver in water impoundment: understanding a problem of water-level management. In: Knighton, M.D., compiler. Water Impoundments for Wildlife: A Habitat Management workshop, 1982 August 31-September 2, Bemidji, MN. USDA Forest Service, General Technician Report NC-100: 95-105.
    • Eustace, Charles D., 1980, II Game Birds and Fur Survey and Inventory - Region 5. Statewide Wildlife Survey and Inventory. W-130-R-11, II, 5, July 1, 1979 - June 30, 1980.
    • Eustace, Charles D., 1981, II Game Birds and Fur Survey and Inventory - Region 5. Statewide Wildlife Survey and Inventory. W-130-R-12, II, 5, July 1, 1980 - June 30, 1981.
    • Evans, T.R. 1948. Beaver-trout-waterfowl problems. Proceedings of the Midwest Wildlife Conference 10: 1-15.
    • Fallau, S.S. 1995. Seasonal streamflow effects on salmonid habitat, and observations of fish movement in Beaver Creek, Idaho-Utah. Utah State University, Logan, Utah.
    • Farmer, Patrick J., and Thomas W. Butts, Western Technology & Eng., Inc., Helena, MT., 1994, McDonald Project Terrestrial Wildlife Study, November 1989 - November 1993. April 1994. In McDonald Gold Project: Wildlife & Fisheries. [#18]. Seven-up Pete Joint Venture, Lincoln, MT. Unpub. No date.
    • Farmer, Patrick. J., et al., Western Technology and Eng., Inc., Helena, MT., 1984, Montana Tunnels Project Baseline Terrestrial Wildlife Study. December 14, 1984. In Application for a Hard Rock Operating Permit, Montana Tunnels Project, Jefferson County, Montana. Vol. 3. Environmental Baseline Reports. (Centennial Minerals, Inc., Hydrometrics, 1984?)
    • Farrar, G.B. 1971. The beaver: the conservationist! Defenders of Wildlife News 46(2): 205-206.
    • Feigley, H.P. 1981. Studies on native small mammals as intermediate hosts of Echinococcus multilocularis. M.Sc. Thesis. Bozeman, Montana: Montana State University. 50 p.
    • Feldmann, A. 1995. The effects of beaver (Castor canadensis) impoundment on plant diversity and community composition in the Coastal Plain of South Carolina. M.S. Thesis, University of Georgia, Athens.
    • Ffolliott, P.F., W.P. Clary, and F.R. Larson. 1976. Observations of beaver activity in an extreme environment. Southwestern Naturalist 21(1): 131-133.
    • Firebaugh, John E., 1980, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-11, Job No. 2, July 1, 1979 - June 30, 1980.
    • Fisher, P.H. 1986. Keeping beavers from plugging culvert inlets. Engineering Field Notes. Washington, DC: USDA Forest Service; 18: 9-13.
    • Ford, T.E. and R.J. Naiman. 1988. Alteration of carbon cycling by beaver: methane evasion rates from boreal forest streams and rivers. Canadian Journal of Zoology 66: 529-533.
    • Foresman, K.R. 2001. The wild mammals of Montana. American Society of Mammalogists, Special Publication Number 12. Lawrence, KS. 278 pp.
    • Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
    • Forman, R.T.T. and M. Godron. 1981. Patches and structural components for a landscape ecology. BioScience 31: 733-739.
    • France, R. L. 1997. The importance of beaver lodges in structuring littoral communities in boreal headwater lakes. Canadian Journal of Zoology 75: 1009-1013.
    • Francis, M.M., R.J. Naiman, and J.M. Melillo. 1985. Nitrogen fixation in subarctic streams influenced by beaver (Castor canadensis). Hydrobiologia 121: 193-202.
    • Fries, C. 1940. Baverland. Nordisk Rotogravyr, Stockholm, Sweden.
    • Frisina, Michael R., 1982, Mt. Haggin Big Game Study. Big Game Survey and Inventory, Region III. W-130-R-13, I-3.2, July 1,1981 - June 30, 1982.
    • Fryxell, J. 1992. Space use by beavers in relation to resource abundance. Oikos 64: 474-478.
    • Fryxell, J. M. 2001. Habitat suitability and source-sink dynamics of beavers. Journal of Animal Ecology 70: 310-316.
    • Fryxell, J. M. and C. M. Doucet. 1993. Diet Choice and the Functional Response of Beavers. Ecology 74: 1297-1306.
    • Fryxell, J.M. and C.M. Doucet. 1991. Provisioning time and central place foraging in beavers. Canadian Journal of Zoology 69: 1308-1313.
    • Fuller, T.K., and J.A. Markl. 1987. Beaver, Castor canadensis, colony density in the Bearville Study Area, northcentral Minnesota. Canadian Field-Naturalist 101:597-598.
    • Fuller, W.A. 1953. Aerial surveys for beaver in the Mackenzie District, Northwest Territories. Trans. North Am. Wildl. Conf. 18: 329-335.
    • Gallo-Reynoso, J.P., G. Suarez-Gracida, H. Cabrera-Santiago, E. Coria-Galindo, J. Egido-Villarreal, and L.C. Ortiz. 2002. Status of beavers (Castor canadensis frondator) in Rio Bavispe, Sonora, Mexico. The Southwestern Naturalist 47(3): 501-504.
    • Gard, R. 1961. Effects of beaver on trout in Sagehen Creek, California. Journal of Wildlife Management 25: 221-242.
    • Gerhart, B. 1979. The land along the water: riparian zones are critical for wildlife survival. Wyoming Wildlife 43(11): 20-23.
    • Gese, E.C. and A.R. Shadle. 1943. Reforestation of aspen after complete cutting by beavers. Journal of Wildlife Management 7: 223-228.
    • Gill, D. 1972. The evolution of a discrete beaver habitat in the Mackenzie River Delta, Northwest Territories. The Canadian Field-Naturalist 86(3): 233-239.
    • Gniadek, S. 1983. Southwest Glendive Wildlife Baseline Inventory. Miles City, Mont: Bureau of Land Management, Miles City District Office. 56 pp with appendices.
    • Godbee, J. and T. Price. 1975. Beaver damage survey. Georgia Forestry Commission, Macon, Georgia. 24pp.
    • Gotie, R.F. and D.L. Jenks. 1984. Assessment of the use of wetlands inventory maps for determining potential beaver habitat. New York Fish and Game Journal 31(1): 55-62.
    • Gowan, C. and K.D. Fausch. 1996a. Long term demographic responses of trout populations to habitat manipulation in six Colorado streams. Ecological Applications 63: 931-946.
    • Gowan, C. and K.D. Fausch. 1996b. Mobile brook trout in two high elevation Colorado streams: re-evaluating the concept of restricted movement. Canadian Journal of Fisheries and Aquatic Sciences 53: 1370-1381.
    • Gowan, C., M.K. Young, K.D. Fausch, and S.C. Riley. 1994. Restricted movement in resident stream salmonids: a paradigm lost? Canadian Journal of Fisheries and Aquatic Sciences 51:2626-2637.
    • Grasse, J.E. 1951. Beaver ecology and management in the Rockies. J. For. 49(1):3-6.
    • Grasse, J.E. and E.F. Putnam. 1950. Beaver management and ecology in Wyoming. Wyoming Game and Fish Commission, Bulletin 6. 52pp.
    • Gray, M.T. 1990. Denver's urban beavers--a gnawing problem. Colorado Outdoors 39(1): 27-29.
    • Green, H.V. 1936. The beaver of the Riding Mountain, Manitoba: an ecological study and commentary. Canadian Field Naturalist 50: 1-8, 21-23, 36-50, 1-67, 85-92.
    • Grover, A.M. 1995. Bird species richness within beaver ponds in south-central New York. Wetlands 15(2): 108-118.
    • Guenther, G.E. 1989. Ecological relationships of bitterbrush communities on the Mount Haggin Wildlife Management Area. M.Sc. Thesis. Bozeman, MT: Montana State University. 73 p.
    • Guenther, S. 1956. A beaver baffler. Washington State Game Bulletin 8(3): 3,6.
    • Gunson, J.R. 1970. Dynamics of the beaver of Saskatchewan's northern forest. M.S. thesis, University of Alberta, Edmondton. 122pp.
    • Gurnell, A.M. 1998. The hydrogeomorphological effects of beaver dam-building activity. Progress in Physical Geography 22(2): 167-189.
    • Gurney, W.S.C. and J.H. Lawton. 1996. The population dynamics of ecosystem engineers. Oikos 76: 273-283.
    • Hacker, A.L. and B.E. Coblentz. 1993. Habitat selection by mountain beavers recolonizing Oregon coast range clearcuts. Journal of Wildlife Management 57: 847-853.
    • Hagglund, A. and G. Sjoberg. 1999. Effects of beaver dams on the fauna of forest streams. Forest Ecology and Management 115: 259-266.
    • Hakala, J.B. 1952. The life history and general ecology of the beaver (Castor canadensis Kuhl) in interior Alaska. M.S. thesis, University of Alaska, Fairbanks. 181pp.
    • Hale, J.G. 1966. Influence of beaver on some trout streams along the Minnesota north shore of Lake Superior. Minnesota Department of Conservation, Division of Game and Fish, Section on Research and Planning Investigation Report 244.
    • Hall, A.M. 1971. Ecology of beaver and selection of prey by wolves in central Ontario. M.S. thesis, University of Toronto, Ontario. 116pp.
    • Hall, J. G. 1960. Willow and Aspen in the Ecology of Beaver on Sagehen Creek, California. Ecology 41: 484-494.
    • Hammerson, G.A. 1994. Beaver (Castor candensis): Ecosystem alterations, management, and monitoring. Natural Areas Journal 14(1): 44-57.
    • Hammond, M.C. 1943. Beaver on the Lower Souris Refuge. Journal of Wildlife Management 7: 316-321.
    • Hand, D. 1984. The beaver's tale: out of the woods and into hot water. Smithsonian 15(8): 162-170.
    • Hanson, W.D. and R.S. Campbell. 1963. The effects of pool size and beaver activity on distribution and abundance of warm-water fishes in a north Missouri stream. American Midland Naturalist 69: 136-149.
    • Harris, D. and S.E. Aldous. 1946. Beaver management in the northern Black Hills of South Dakota. Journal of Wildlife Management 10: 348-355.
    • Harris, H.T. 1991. Habitat use by dispersing and transplanted beavers in western Montana. M.S. Thesis, University of Montana, Missoula, MT. 40p.
    • Hartman, A.M. 1975. Analysis of conditions leading to the regulation of water flow by a beaver. The Psychological Record 25: 427-431.
    • Hartman, G. 1994a. Ecological studies of reintroduced beaver (Castor fiber) population. Dissertation, Swedish University of Agricultural Sciences, Department of Agricultural Sciences.
    • Hartman, G. 1994b. Long -term population development of a reintroduced beaver (Castor fiber) population in Sweden. Conservation Biology 8: 713-717.
    • Hartman, G. 1996. Habitat selection by European beaver (Castor fiber) colonizing a boreal landscape. Journal of Zoology 240(2): 317-325.
    • Hawkes, F.W. 1973. Elk, moose, and Beaver. Pacific Discovery 26(3): 12-15.
    • Hay, K.G. 1958. Beaver census methods in the Rocky Mountain region. Journal of Wildlife Management 22: 395-402.
    • Heggenes, J., T.G. Northcote, and A. Peter. 1990. Spatial stability of cutthroat trout (Oncorhynchus clarki) in a small, coastal stream. Can. J. Fish. Aquat. Sci. 48: 757-762.
    • Hendricks, P., S. Lenard, D.M. Stagliano, and B.A. Maxell. 2013. Baseline nongame wildlife surveys on the Fort Peck Indian Reservation. Report to the Assiniboine and Sioux Tribes of the Fort Peck Indian Reservation. Montana Natural Heritage Program, Helena, MT. 83 p.
    • Henry, D.B. 1967. Age structure, productivity, and habitat characteristics of beaver in northeastern Ohio. M.S. Thesis, Ohio State University, Columbus. 68p.
    • Henry, D.B. and T.A. Bookhout. 1969. Productivity of beavers in northeastern Ohio. Journal of Wildlife Management 33: 927.
    • Hensley, A.L. and B.C. Fox. 1948. Experiments on the management of Colorado River beaver. Calif. Fish Game 43: 115.
    • Heter, E.W. 1950. Transplanting beavers by airplane and parachute. Journal of Wildlife Management 14: 143-147.
    • Hibbard, E.A. 1958. Movements of beaver transplanted in North Dakota. Journal of Wildlife Management 22: 209-211.
    • Hilfiker, E.L. 1991. Beavers: water, wildlife, and history. Windswept Press, Interlaken, New York. 198pp.
    • Hill, E.P. 1976. Control methods for nuisance beavers in the southeastern United States. Proceedings of the 7th Vertebrate Pest Conference, Monterey, California: 85-98.
    • Hill, E.P. 1987. Beaver restoration. Pages 281-286 in Restoring America's wildlife 1937-1987: The first fifty years of the Federal Aid in Wildlife Restoration (Pittman-Robertson) Act. Edited by H. Kallman, C.P. Agee, W.R. Goforth, J.P. Linduski, and N. Rollison. USDI Fish and Wildlife Service, U.S. Government Printing Office, Washington, DC.
    • Hill, E.P. and N.S. Novakowski. 1984. Beaver management and economics in North America. Acta Zoologica Fernica 1972: 259-262.
    • Hiner, L.E. 1938. Observations on the foraging habits of beavers. Journal of Mammalogy 19: 317-319.
    • Hodgdon, H.E. and J.S. Larson. 1973. Some sexual differences in behavior within a colony of marked beavers (Castor canadensis). Animal Behavior 21: 147.
    • Hodgdon, H.E. and J.S. Larson. 1980. A bilbliography of the recent literature on beaver. Mass. Agric. Exp. Sta., Research Bulletin No. 665, p128.
    • Hodgdon, H.E. and R.A. Lancia. 1983. Behavior of the North American beaver, Castor canadensis. Acta Zoologica Fennica 174: 99-103.
    • Hodgdon, K.W. and J.H. Hunt. 1953(1966?). Beaver management in Maine. Maine Department of Inland Fish and Game, Game Division Bulletin No. 3.
    • Hodkinson, I.D. 1975. A community analysis of the benthic insect fauna of an abandoned beaver pond. Journal of Animal Ecology 44(2): 533-551.
    • Hodkinson, I.D. 1975. Dry weight loss and chemical changes in vascular plant litter of terrestrial origin occuring in a beaver pond ecosystem. Journal of Ecology 63(1975a): 131-142.
    • Hodkinson, I.D. 1975. Energy flow and organic matter decomposition in an abandoned beaver pond ecosystem. Oecologia (Berlin) 21: 131-139.
    • Hoffmann, R.S. and D.L. Pattie. 1968. A guide to Montana mammals: identification, habitat, distribution, and abundance. Missoula, MT: University of Montana. 133 p.
    • Hoover, R.L. 1955. Beaver ecology in the Longs Peak area of Colorado. M.S. thesis, Colorado A7M College, Fort Collins. 262pp.
    • Hoover, W.H. and S.D. Clark. 1972. Fiber digestion in beavers. J. Nutr. 102: 9-16.
    • Horstman, L.P. 1979. Evaluation of beaver depredation and control in the Edmonton Fish and Wildlife Region, 1979. Edmonton, AB: Alberta Energy and Natural Resources, Fish and Wildlife Division. 36p.
    • Hossack,B.R., W.R. Gould, D.A. Patla, E. Muths, R. Daley, K. Legg, and P.S. Corn. 2015. Trends in Rocky Mountain amphibians and the role of beaver as a keystone species. Biological Conservation 187:260-269.
    • Howard, R.J. 1982. Beaver habitat classification in Massachusetts. M.S. thesis, University of Massechusetts, Amherst.
    • Howard, R.J. and J.S. Larson. 1985. A stream habitat classification system for beaver. Journal of Wildlife Management 49(1): 19-25.
    • Huey, W.S. and W.H. Wolfrum. 1956. Beaver-trout relations in New Mexico. Progressive Fish-Culturalist 18: 70-74.
    • Hutchinson, M.D., S. Olson, K. Lewis, and T. Vogt. 1986. Beaver at LaRue Swamp, Union County, Illinois: an assessment of their influence upon the area. Unpublished report. Natural Land Institute, Illinois 62908. 48pp.
    • Ingle-Sidorowicz, H.M. 1982. Beaver increase in Ontario: result of changing environment. Mammalia 46: 167-175.
    • Ives, R.L. 1942. The beaver-meadow complex. Journal of Geomorphology 5: 191-203.
    • Jackson, M.D. 1990. Beaver dispersal in western Montana. Thesis. Missoula, Montana: University of Montana. 79 p.
    • Janson, Reuel G., 1977, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-8, Job No. 2, July 1, 1976 - June 30, 1977.
    • Janson, Reuel G., 1978, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-9, Job No. 2, July 1, 1977 - June 30, 1978.
    • Janson, Reul G., 1976, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-7, Job No. 2, July 1, 1975 - June 30, 1976.
    • Jeffress, J. 1975. The beaver (Castor canadensis). Nevada Outdoors and Wildlife Review 9(4): 1-6.
    • Jeglum, J.K. 1975. Vegetation-habitat changes caused by damming a peatland drainageway in northern Ontario. Canadian Field-Naturalist 89: 400-412.
    • Jenkins, S. H. and P. E. Busher. 1979. Castor Canadensis. Mamm. Species 120:1-8.
    • Jenkins, S.H. 1975. Food selection by beavers: a multidimensional contingency table analysis. Oecologia 21: 157-173.
    • Jenkins, S.H. 1978. Food selection by beavers: sampling behavior. Breviora 447: 1-6.
    • Jenkins, S.H. 1979. Seasonal and year-to-year differences in food selection by beavers. Oecologia (Berlin) 44: 112-116.
    • Jenkins, S.H. 1980. A size-distance relation in food selection by beavers. Ecology 61: 740-746.
    • Jenkins, S.H. 1981. Problems, progress, and prospects in studies of food selection by beavers. In: Chapman, J.A., D. Pursley, eds. Proceedings worldwide furbearer conference; 1980 August 8-11; Frostburg, MD. Frostburg, MD: Worldwide Furbearer Conference, Inc.: 559-579.
    • Johannsen, N. 1970. About beavers. Pacific Search 5(3): 8.
    • Johnson, D.R. and D.H. Chance. 1974. Presettlement overharvest of upper Columbia River beaver populations. Canadian Journal of Zoology 52: 1519-1512.
    • Johnson, J. 1981. Leave it to beaver. Natural History 90: 44-47.
    • Johnson, L.J. 1960. Mammal studies on the Lubrecht Forest, Montana: a preliminary report. Proc. Mont. Acad. Sci. 20: 40-47.
    • Johnson, P. 1984. The dam buildeer is at it again! National Wildlife, June/July, pp 9-15.
    • Johnson, R.B. 1983. Aspen utilization by beavers (Castor canadensis) in northern Wisconsin. Wisconsin Academy of Sciences. Arts and Letters 71 part 2: 82-86.
    • Johnson, R.C., F.J. Rahel, and W.A. Hubert. 1992. Factors influencing the size structure of brook trout populations in beaver ponds in Wyoming. N. Amer. J. Fish. Manage. 12: 118.
    • Johnson, R.C., J.W. Preacher, J.R. Gwaltney, and J.E. Kennamer. 1976. Evaluation of habitat manipulation for ducks in an Alabama beaver pond complex. Proc. Ann. Conf. Southeast Assoc. Game Fish Comm. 29: 512.
    • Johnston, C.A. and R.J. Naiman. 1987. Boundary dynamics at the aquatic-terrestrial interface: The influence of beaver and geomorphology. Landscape Ecology 1(1): 47-57.
    • Johnston, C.A. and R.J. Naiman. 1990. Aquatic patch creation in relation to beaver population trends. Ecology 71: 1617-1621.
    • Johnston, C.A. and R.J. Naiman. 1990. The use of a geographic information system to analyze long-term landscape alteration by beaver. Ecology 71: 5-19.
    • Johnston, C.A., G. Pinay, C. Arens, and R.J. Naiman. 1995. Influence of soil properties on the biogeochemistry of a beaver meadow hydrosequence. Soil Science Society of America Journal 59(6): 1789-1799.
    • Johnston, C.A., J. Pastor, and R.J. Naiman. 1993. Effects of beaver and moose on boreal forest landscapes. In: Cousins, S.H., R. Haines-Young, and D. Green; editors. Landscape and Geographical Information Systems. Taylor and Francis, London, pp. 237-254.
    • Jonas, R. J. 1955. A population and ecological study of the beaver (Castor canadensis) of Yellowstone National Park. M.S. Thesis, University of Idaho, Moscow.
    • Jonas, R. J. 1959. Beaver. Naturalist 10(2): 60-61.
    • Jones, C.G., J.H. Lawton, and M. Shachak. 1994. Organisms as ecosystem engineers. Oikos 69: 373-386.
    • Jones, C.G., J.H. Lawton, and M. Shachak. 1997. Positive and negative effects of organisms as physical ecosystem engineers. Ecology 78: 1946-1957.
    • Joslin, Gayle, and Heidi B. Youmans. 1999. Effects of recreation on Rocky Mountain wildlife: a review for Montana. [Montana]: Montana Chapter of the Wildlife Society.
    • Kafcas, E.N. 1987. Census and exploitation of a discrete beaver population in Michigan. M.S. thesis, Central Michigan University, Mount Pleasant.
    • Karston, P. 1983. Displaying natural rearing of beaver (Castor canadensis). Zoologische Garten 53(6): 404-408.
    • Kay, C.E. 1994. The impact of native ungulates and beaver on riparian communities in the intermountain west. Natural Resources and Environmental Issues 1: 23-44.
    • Kaye, C.A. 1962. Early postglacial beavers in southeastern New England. Science 138: 906-907.
    • Keast, A. and M.G. Fox. 1990. Fish community structure, spatial distribution, and feeding ecology in a beaver pond. Environmental Biology of Fishes 27: 201-214.
    • Keigley, R.B. 2000. Elk, Beaver, and the persistence of willows in national parks: comment on Singer et al. (1998). Wildlife Society Bulletin 28(2): 448-450.
    • Keller, E.A. and F.J. Swanson. 1979. Effects of large organic material on channel form and fluvial processes. Earth Surface Processes 4: 361-380.
    • Kienzler, J.M. 1971. Woody plant utilization by beaver in naturally acid and acid mine water. M.S. thesis, West Virginia University, Morgantown.
    • Kindschy, R.R. 1985. Response of red willow to beaver use in southeastern Oregon. Journal of Wildlife Management 49(1): 26-28.
    • King, S.L., B.D. Keeland, and J.L. Moore. 1998. Beaver lodge distribution and damage assessments in a forested wetland ecosystem in the southern United States. Forest Ecology and Management 108: 1-7.
    • Kirby, R.E. 1976. Mapping wetlands on beaver flowages with 35-mm photography. Canadian Field-Naturalist 90: 423-431.
    • Klotz, R.L. 1998. Influence of beaver ponds on the phosphorus concentration of stream water. Canadian Journal of Fisheries and Aquatic Sciences 55: 1228-1235.
    • Knapp, Stephen, and Bernie Hildebrand, 1984, Upland Game Bird and Fur Surveys and Inventory - Region 7. Statewide Wildlife Survey & Inventory. W-130-R-15, II-7, July 1, 1983 - June 30, 1984.
    • Knapp, Steven, and Bernie Hildebrand, 1982, Upland Game Bird and Fur Surveys and Inventory - Region 7. Statewide Wildlife Survey and Inventory. W-130-R-13, II-7, July 1, 1981 - June 30, 1982.
    • Knudsen, G.J. 1954. Beaver damage to forests and forest reproduction. Wisconsin Wildlife Research 13: 121-129.
    • Knudsen, G.J. 1962. Relationship of beaver to forests, trout, and wildlife in Wisconsin. Wisconsin Conservation Department Technical Wildlife Bulletin No. 25. 52pp.
    • Knudsen, G.J. and J.B. Hale. 1965. Movements of transplanted beaver in Wisconsin. Journal of Wildlife Management 29: 685-688.
    • Koch, R.R., R. Curry, and M. Weber. 1977. The effect of altered streamflow on the hydrology and geomorphology of the Yellowstone River Basin, Montana. Montana Department of Natural Resources and Conservation, Yellowstone Impact Study Tech. Report 2. 163pp.
    • Komadina-Douthwright, S.M. 1994. Effects of beaver (Castor canadensis) activity on stream water quality under conditions of prolonged snow and ice cover (winter 1991-1992). Canadian Technical Report of Fisheries and Aquatic Sciences 1(6-7): 1-23.
    • LaBastille, A. 1979. The best dam builder around. National Wildlife 17(3): 26-33.
    • LaBastille, A. 1986. Beaver dammed again. Adirondack Life, March/April.
    • Lahti, S. and M. Helminen. 1974. The beaver, Castor fiber L., and Castor canadensis Kuhl in Finland. Acta Theriologica 19: 177-189.
    • Lambeth, R. H. 1953. New beaver law provides for open season. Montana Wildl. Winter.
    • Lampe, R.P., J.K. Jones Jr., R.S. Hoffmann, and E.C. Birney. 1974. The mammals of Carter County, southeastern Montana. Occa. Pap. Mus. Nat. Hist. Univ. Kan. 25:1-39.
    • Lancia, R.A. and J.W. Bishir. 1985. Mortality rates of beaver in Newfoundland: a comment. Journal of Wildlife Management 49: 879-881.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Big Spring Creek, Lewistown, Montana. Proj. No. 130091.029. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Lawrence Park, Kalispell, Montana. Proj. No. 130091.008. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Lang, G.E. and R.K. Wieder. 1984. The role of beaver in vegetation patterning and development in sphagnum-dominated wetlands in West Virginia. Bulletin of the Ecological Society of America 65: 243.
    • Laramie, H.A., Jr. 1963. A device for the control of problem beavers. Journal of Wildlife Management 27: 471-476.
    • Larson, J.S. 1967. Age structure and sexual maturity within a western Maryland beaver (Castor canadensis) population. Journal of Wildlife Management 32: 93-103.
    • Larson, J.S. and J.R. Gunson. 1983. Status of the beaver in North America. Acta Zoologica Fennica 174: 91-93.
    • Lawrence, W., L. Fay, and S. Graham. 1956. A report on the beaver die-off in Michigan. Journal of Wildlife Management 20: 184-187.
    • Lawrence, W.H. 1952. Evidence of the age of beaver ponds. Journal of Wildlife Management 16: 69-79.
    • Lawrence, W.H. 1954. Michigan beaver populations as influenced by fire and logging. Dissertation, University of Michigan, Ann Arbor, Michigan.
    • Leege, T.A. 1968. Beavers on the move. Idaho Wildlife Review 20(5): 14-16.
    • Leege, T.A. 1968. Natural movements of beavers in southeastern Idaho. Journal of Wildlife Management 32: 973-976.
    • Leege, T.A. and R.M. Williams. 1967. Beaver productivity in Idaho. Journal of Wildlife Management 31(2): 326-332.
    • Leidholt, K.L., W. McComb, and D.E. Hibbs. 1989. The effect of beaver on stream and stream-side characteristics and coho populations in western Oregon. Northwest Science (Abstract) 63: 71.
    • Leidholt-Bruner, K., D.E. Hibbs, and W.C. McComb. 1992. Beaver dam locations and their effects on distribution and abundance of Coho salmon fry in two coastal Oregon streams. Northwest Science 66(4): 218-223.
    • Leighton, A.H. 1933. Notes on the relation of beavers to one another and to the muskrat. Journal of Mammalogy 14: 27-35.
    • Levin, S.A. and R.T. Paine. 1974. Disturbance, patch formation, and community structure. Proc. Nat. Acad. Sci. 71: 2744-2747.
    • Libby, W.L. 1957. Observations on beaver movements in Alaska. Journal of Mammalogy 38: 269.
    • Lizarralde, M.S. 1993. Current status of the introduced beaver (Castor canadensis) population in Tierra del Fuego, Argentina. Ambio 22(6): 351-358.
    • Lizarralde, M.S. 1996. Effects of beaver (Castor candensis) on the nutrient dynamics of the Southern Beech forest of Tierra del Fuego (Argentina). Ecologia Australia 6(2): 101-105.
    • Llred, M. 1980. A re-emphasis on the value of the beaver in natural resource conservation. Journal of Idaho Academy of Science 16: 3-10.
    • Llred, M. 1981. The potential use of beaver population behaviour in beaver resource management. Journal of Idaho Academy of Science 17: 14-24.
    • Lochmiller, R.L. 1979. Use of beaver ponds by southeastern woodpeckers in winter. Journal of Wildlife Management 43: 262-266.
    • Longley, W.H. and J.B. Moyle. 1963. Beaver in Minnesota. Minnesota Department of Conservation Technical Bulletin 6. 63pp.
    • Lovallo, M.J., J.H. Gilbert, and M. Gehring. 1993. Bobcat (Felis rufus) dens in an abandoned beaver (Castor canadensis) lodge. The Canadian Field-Naturalist 107(1): 108-109.
    • Lowry, M.M. 1993. Groundwater elevations and temperature adjacent to a beaver pond in central Oregon. Unpublished M.S. thesis, University of Oregon.
    • Lulman, P.D. 1974. Moose and muskeg, birch and beaver, natural environment and wildlife. Occas. Publ. 12. Edmonton, AB: University of Alberta, Boreal Institute for Northern Studies: 232-32.
    • Lundberg, J. and F. Moberg. 2003. Mobile Link Organisms and Ecosystem Functioning: Implications for Ecosystem Resilience and Management. Ecosystems 6: 87-98.
    • MacArthur, R.A. 1989. Energy metabolism and thermoregulation of beaver (Castor canadensis). Canadian Journal of Zoology 67: 651-657.
    • MacArthur, R.A. and A.P. Dyck. 1990. Aquatic thermoregulation of captive and free-ranging beavers (Castor canadensis). Canadian Journal of Zoology 68: 2409-2416.
    • MacDonald, D.W., F.H. Tattersall, E.D. Brown, and D. Balharry. 1995. Reintroducing the European beaver to Britain - Nostalgic meddling or restoring biodiversity? Mammal Review 25: 161-200.
    • Maret, T.J. 1985. The effect of beaver ponds on the water quality of Currant Creek, Wyoming. University of Wyoming, Laramie, Wyoming.
    • Maret, T.J., M. Parker, and T.E. Fannin. 1987. The effect of beaver ponds on the nonpoint source water quality of a stream in southwestern Wyoming. Wat. Res. 21(3): 263-268.
    • Margolis, B.E., M.S. Castro, and R.L. Raesly. 2001. The impact of beaver impoundments on the water chemistry of two Appalacian streams. Canadian Journal of Fisheries and Aquatic Sciences 58: 2271-2283.
    • Marston, R.A. 1994. River enterenchment in of small mountain valleys of the western USA; influence of beaver, grazing, and clearcutt logging. Rev. geogr. Lyon 69: 11-15.
    • Martin, P.R., K. Dubois and H.B. Youmans. 1981. Terrestrial wildlife inventory in selected coal areas, Powder River resources area final report. Montana Department of Fish, Wildlife and Parks and Bureau of Land Management, Helena, MT. No. YA-553-CTO- 24. 288 p.
    • Martinsen, G. D., E. M. Driebe, and T. G. Whitham. 1998. Indirect interactions mediated by changing plant chemistry: beaver browsing benefits beetles. Ecology 79(1): 192-200.
    • Masslich, W.J., J.D. Brotherson, and R.G. Cates. 1988. Realtionship of aspen to foraging patterns of beaver in the Strawberry Valley of central Utah. Great Basin Naturalist 48: 250.
    • Matthews, W.L. 1980a. Wibaux-Beach comparison study: Sydney, Glendive and Plevna Study Areas. Bureau of Land Management, Miles City, MT. 50 p.
    • Maxell, B.A. 2016. Northern Goshawk surveys on the Beartooth, Ashland, and Sioux Districts of the Custer-Gallatin National Forest: 2012-2014. Montana Natural Heritage Program. Helena, MT. 114pp.
    • Maxim Technologies, Inc., 2002, Western Energy Company Rosebud Mine, Colstrip, Montana: 2002 Annual Wildlife Monitoring Report; December 1, 2001 - November 30, 2002. Febr. 24, 2002.
    • Maxwell, J. 1994. Leave it to beavers. Audubon March/April: 104-109.
    • McCall, T.C. 1996. Beaver populations and their relation to wetland habitat and breeding waterfowl in Maine. Wetlands 16(2): 163-172.
    • McColley, S.D. 2007. Restoring aspen riparian stands with beaver on the northern Yellowstone winter range. M.Sc. Thesis. Bozeman, MT: Montana State University. 67 p.
    • McComb, W.C., J.R. Sedell, and T.D. Buchholz. 1990. Dam-site selection by beavers in an eastern Oregon basin. Great Basin Naturalist 50: 273-281.
    • McDonald, D. 1956. Beaver carying capacity of certain mountain streams in North Park, Colorado. M.S. thesis, Colorado State University, Fort Collins.
    • McDowell, D.M. and R.J. Naiman. 1986. Structure and function of a benthic invertebrate stream community as influenced by beaver (Castor candensis). Oecologia (Berlin) 68: 481-489.
    • McDowell, R.A. 1975. A study of the Pole Mountain fishery: beaver pond, artificial impoundment and stream investigations. Laramie, Fish Division of the Wyoming Game and Fish Department: 174.
    • McGinley, M.A. and T.G. Whitham. 1985. Central place foraging by beavers (Castor canadensis): a test of foraging predictions and the impact of selective feeding on the growth form of cottonwoods (Populus fremontii). Oecologia 66: 558-562.
    • McIntre, R. 1981. Death of a beaver pond. Alaska 47: 50.
    • McKinstry, M. C. and S. H. Anderson. 1998. Using Snares to Live-Capture Beaver, Castor canadensis. The Canadian Field-Naturalist 112: 469-473.
    • McKinstry, M. C. and S. H. Anderson. 2002. Survival, Fates, and Success of Transplanted Beavers, Castor canadensis, in Wyoming. The Canadian Field-Naturalist 116: 60-68.
    • McKinstry, M.C. 2001. Using beavers to create wetlands in Wyoming. Birdscapes Spring/Summer: 8-9.
    • McKinstry, M.C. and S.H. Anderson. 1997. Use of beaver to improve riparian areas in Wyoming. Pages 128-134 in Wyoming Water 1997: Applied Research for Management of Wyoming's Water Resources. Edited by C. Goertler, C. Rumsey, T. Bray, and D. Boyson. Wyoming Water Research Center, Laramie, Wyoming.
    • McKinstry, M.C. and S.H. Anderson. 1999. Attitudes of private and public land managers in Wyoming, USA, toward beaver. Environmental Management 23(1): 95-101.
    • McKinstry, M.C. and S.H. Anderson. In press. Predation on introduced beaver in Wyoming. Greater Yellowstone Predators: Ecology and Conservation in a Changing Landscape. Symposium Proceedings.
    • McKinstry, M.C., P. Caffrey, and S.H. Anderson. 2000. The importance of beaver to duck populations in Wyoming. Pages 95-100 in Riparian Ecology and Management in Multi-Land Use Watersheds. Edited by P.J. Wigington and R.L. Beschta. American Water Resources Association, Middleburg, Virginia.
    • McKinstry, M.C., P. Caffrey, and S.H. Anderson. 2001. The importance of beaver to wetland habitats and waterfowl in Wyoming. Journal American Water Resources Association 37: 1571-1577.
    • McMaster, R.T. 1997. Floristics, zonation, and succession of vascular vegetation in fifteen beaver-impacted wetlands in western Massachusetts. Ph.D. dissertation, University of Massachusetts, Amherst.
    • McMaster, R.T. and N.D. McMaster. 2000. Vascular flora of beaver wetlands in western Massachusetts. Rhodora 102(910): 175-197.
    • McRae, G. and C.J. Edwards. 1994. Thermal characteristics of Wisconsin headwater streams occupied by beaver: Implications for brook trout habitat. Transactions of the American Fisheries Society 123: 641-656.
    • Medin, D. E. and W. P. Clary. 1991. Small mammals of a beaver pond ecosystem and adjacent riparian habitat in Idaho. USDA, Forest Service, Res. Paper INT-445.
    • Medin, D.E. and K.E. Torquemada. 1988. Beaver in western North America: An annotated bibliography, 1966 to 1986. USDA Forest Service, Intermountain Research Station, General Technician Report INT-242. 17pp.
    • Medin, D.E. and W.P. Clary. 1990. Bird populations in and adjacent to a beaver pond ecosystem and adjacent riparian habitat in Idaho. Intermountain For. And Range Exp. Sta., USDA Forest Service, Ogen, Utah.
    • Medwecka-Kornas, A. and R. Hawro. 1993. Vegetation on beaver dams in the Ojcow National Park (southern Poland). Phytocoenologia 23: 611-618.
    • Meentemyer, R.K. and D.R. Butler. 1999. Hyrogeomorphic effects of beaver dams in Glacier National Park, Montana. Physical Geography 20(5): 436-446.
    • Metts, B.S. 2001. Evaluation of herpetofaunal communities on upland streams and beaver-impounded streams in the upper Piedmont of South Carolina. American Midland Naturalist 145(1): 54-65.
    • Michelmore, P. 1984. The amazing beavers of Currant Creek. Reader's Digest 124(744): 109-113.
    • Miller, K.L. 1967. Microclimate of northern beaver: a constructed habitat. In: Fourth international biometerological congress; 1966 August 26-September 2; New Brunswick, NJ. New Brunswick, NJ: Rutgers University: 288.
    • Mitchell, J. L. 1958. Beaver age and sex ratios. Montana Fish and Game Dept., Job Compl. Rep., Fed. Aid Proj. W-49-R-7:16-36.
    • Molini, J.J., R.A. Lancia, J. Bishir, and H.E. Hodgdon. 1980. A stochastic model of beaver population growth. In: Chapman, J.A. and D. Pursley, eds. Proceedings, worldwide furbearer conference; 1980 August 8-11; Frostburg, MD. Frostburg, MD: Worldwide Furbearer Conference, Inc.: 1215-1245.
    • Montana Dept. of Fish, Wildlife & Parks, 1964, Beaver population and distribution trends - Aerial Survey. Wildlife Investigations - District Six. W-76-R-9, D-2 through W-76-R-13, D-2, 1963-1969.
    • Montana Dept. of Fish, Wildlife & Parks, 1985, Upland Game Bird and Furbearer Survey and Inventory. Wildlife Investigations, Region One. W-130-R-16, Job No. II-1, July 1, 1984 - June 30, 1985.
    • Montana Dept. of Fish, Wildlife & Parks, 1986, Upland Game Bird and Furbearer Survey and Inventory. Wildlife Investigations, Region One. W-130-R-17, Job No. II-1, July 1, 1985 - June 30, 1986.
    • Montana Dept. of Fish, Wildlife & Parks, 1987, Upland Game Bird and Furbearer Survey and Inventory. Wildlife Investigations, Region One. W-130-R-18, Job No. II-1, July 1, 1986 - June 30, 1987.
    • Montana Dept. of Transportation., 200?, Montana Dept. of Transportation Biological Resources Report: Wetland mitigation east of Browning. Montana Wetland ?? Proj. No. NH 0002(232) CN 0703. In Perry Ranch - East of Browning Wetland Mitigation Site, Glacier County. US#3. Man.? Fin Dist3 Admin.Dist3
    • Moore, G.C. and E.C. Martin. 1949. Status of beaver in Alabama. Alabama Department of Conservation: Game Fish and Sea Foods Division. Pittman-Robertson Project. 30p.
    • Moore, P. D. 1999. Ecology: Sprucing up beaver meadows. Nature 400: 622-623.
    • Moran, S.S. 2001. Survey of habitat & fish communities in segments of Cherry Creek, a proposed site for re-introduction of Westslope Cutthroat Trout. M.Sc. Thesis. Bozeman, MT: Montana State University. 121 p.
    • Morgan, L.H. 1868. The American Beaver. Dover Publications, Inc. 384pp.
    • Mosher, N.D. 1981. Floristic and successional development of three abandoned beaver meadows in western Massachusetts. M.A. thesis, Smith College, Northampton, MA.
    • Muchmore, D. 1975. Beaver, if you will. Wyoming Wildlife 39(1): 16-21, 34.
    • Muller-Schwartze, D. et al. No date. On a study of the behavioral and population ecology of the beaver at Acadia National Park. National Park Service Cooperative Research Unit, State University of New York, College of Environmental Science and Forestry, Institute of Environmental Program Affairs, Syracuse, New York. *6pp.
    • Muller-Schwarze, D. and S. Hickman. 1980. The social role of scent marking in beaver (Castor canadensis). Journal of Chemical Ecology 6: 81.
    • Munther, G.L. 1982. Beaver management in grazed riparian ecosystems. Wildlife-livestock relationships symposium: proceedings 10, Moscow, ID, University of Idaho.
    • Munther, G.L. 1983. Integration of beaver into forest management. Proc. Annu. Meeting Colo.-Wyo. Chapter Am. Fish. Soc. 18: 73-80.
    • Naiman, R. J., J. M. Melillo, and J. E. Hobbie. 1986. Ecosystem alteration of boreal forest streams by beaver (Castor canadensis). Ecology 67: 1254-1269.
    • Naiman, R.J. 1988. Animal influences on ecosystem dynamics. BioScience 38(11): 750-752.
    • Naiman, R.J. and J.M. Melillo. 1984. Nitrogen budget of a subarctic stream altered by beaver (Castor canadensis). Oecologia (Berlin) 62: 150-155.
    • Naiman, R.J., B.S. Farr, M.M. Francis, J.M. Melillo, and J.E. Hobbie. 1982. The role of beaver in shaping aquatic ecosystem dynamics. In: Naiman, R.J. (editor). The matamek Research Program: Annual Report for 1981. Woods Hole Oceanographic Institution Tech Rept 81-29, pp 90-117.
    • Naiman, R.J., D.M. McDowell, and B.S. Farr. 1984. The influence of beaver (Castor canadensis) on the production dynamics of aquatic insects. Vehr. Internat. Verein. Limnol. 22: 1801-1810.
    • Naiman, R.J., G. Pinay, C.A. Johnston, and J. Pasto. 1994. Beaver influences on the long-term biogeochemical characteristics of boreal forest drainage networks. Ecology 75: 905-921.
    • Naiman, R.J., T. Manning, and C.A. Johnston. 1991. Beaver population fluctuations and tropospheric methane emissions in boreal wetlands. Biogeochemistry 12: 1-15.
    • Naiman, Robert J., Carol A. Johnston, and James C. Kelley. 1988. Alteration of North American streams by beaver. BioScience 38(11): 753-762.
    • Nash, J.B. 1951. An investigation of some problems of ecology of the beaver, Castor canadensis Kuhl, in northern Manitoba. M.S. thesis, University of Manitoba. 64pp.
    • Nawrocki, T., C.A. Johnson, and J. Sales. 1994. GIS and modeling in ecological studies: Analysis of beaver pond impacts on runoff and its quality. Technician Report NRRI/TR-94/01, Natural Resource Research Institute, Duluth, MN.
    • Neff, D.J. 1957. Ecological effects of beaver habitat abandonment in the Colorado Rockies. Journal of Wildlife Management 21(1): 80-84.
    • Neff, D.J. 1959. A seventy-year history of a Colorado beaver colony. Journal of Mammalogy 40: 381-387.
    • Nevers, H.P. 1968. Waterfowl utilization of beaver impoundments in southeastern New Hampshire. Trans. Northeast Fish and Wildl. Conf. 25: 105-120.
    • New York Department of Environmental Conservation. 1991. Status of beaver management in New York. Draft report.
    • Newby, F. E. 1955. Beaver ecology in the Jefferson Basin - fur resources survey and investigation. Montana Fish and Game Department, P-R Job Compl. Rept. Proj. W-49R- 11, Job No. II-B, Segment 1.
    • Nielsen, Lyn S., 1983, II Game Bird and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. Region Two Survey and Inventory. W-130-R-14, Job No. 2, July 1, 1982 - June 30, 1983.
    • Nixon, C.M. and J. Ely. 1969. Foods eaten by a beaver colony in southeast Ohio. Ohio Journal of Science 69: 313-319.
    • Nolet, B.A. and F. Rosell. 1994. Territoriality and time budgets in beavers during sequential settlement. Canadian Journal of Zoology 72: 1227-1237.
    • Nolet, B.A. and F. Rosell. 1998. Comeback of the beaver Castor fiber: an overview of old and new conservation problems. Biological Conservation 83(2): 165-173.
    • Nolet, B.A. and J.M. Baveco. 1996. Development and viability of a translocated beaver, Castor fiber, Population in the Netherlands. Biological Conservation 75: 125-137.
    • Nolet, B.A., A. Hoekstra, and M.M. Ottenheim. 1994. Selective foraging on woody species by the beaver (Castor fiber) and its impact on a riparian willow forest. Biological Conservation 70: 117-128.
    • Nordstrom, W.R. 1972. Comparison of trapped and untrapped beaver populations in New Brunswick. M.S. thesis, University of New Brunswick, Frederickton. 104pp.
    • Northcott, T.H. 1964. An investigation of the factors affecting carrying capacity of selected areas in Newfoundland for the beaver, Castor canadensis caecator Bangs, 1913. M.S. thesis, Memorial University, ST. John's, Newfoundland. 133pp.
    • Northcott, T.H. 1971. Feeding habits of beaver in Newfoundland. Oikos 22: 407-410.
    • Northrop, Devine & Tarbell, Inc. 1994. Cabinet Gorge and Noxon Rapids hydroelectric developments: 1993 wildlife study. Unpublished report to the Washington Water Power Company, Spokane. Vancouver, Washington and Portland, Maine. 144 pp. plus appendices.
    • Northrop, Devine and Tarbell, Inc. 1995. Cabinet Gorge and Noxon Rapids Hydroelectric Developments, 1994 Wetland Mapping and Assessment Study, Volume I of II. 27 pp. plus appendices.
    • Novak, M. 1976. The beaver in Ontario. Spec. Publ. , Ontario Ministry of Natural Resources.
    • Novak, M. 1977. Determining the average size and composition of beaver families. Journal of Wildlife Management 41: 751-754.
    • Novakowski, N.S. 1965. Population dynamics of a beaver population in northern latitudes. Ph.D. dissertation. University of Saskatchewan, Saskatoon. 154pp.
    • Novakowski, N.S. 1967. The winter bioenergetics of a beaver population in northern latitudes. Canadian Journal of Zoology 45(6, part 1): 1107-1118.
    • Novakowski, N.S. 1969. The influence of vocalization on the behavior of beaver, Castor canadensis. American Midland Naturalist 81(1): 198-204.
    • Nummi, P. 1989. Simulated effects of the beaver on vegetation, invertebrates and ducks. Ann. Zool. Fennici 26: 43-52.
    • Nummi, P. 1992. The importance of beaver ponds to waterfowl broods: an experiment and natural tests. Ann. Zool. Fennici 29: 47-55.
    • Nummi, P. and H. Poysa. 1997. Population and community level responses in Anas-species to patch disturbance caused by ecosystem engineer, the beaver. Ecography 20: 580-584.
    • Oechsli, L.M. 2000. Ex-urban development in the Rocky Mountain West: consequences for native vegetation, wildlife diversity, and land-use planning in Big Sky, Montana. M.Sc. Thesis. Montana State University, Bozeman. 73 p.
    • Oertli, E.F. 1976. The beavers of Kananaskis Forest. Nature Canada 5(1): 3-8.
    • Olson, Gary R., 1978, Game Birds and Fur (Small Game) Survey and Inventory (Fur Survey Segment). Statewide Wildlife Survey and Inventory. W-130-R-9, II-4, July 1, 1977 - June 30, 1978.
    • Olson, Gary R., 1979, Game Birds and Fur (Small Game) Survey and Inventory (Fur Survey Segment). Statewide Wildlife Survey and Inventory. W-130-R-10, II-4, July 1, 1978 - June 30, 1979.
    • Olson, Gary R., 1980, Game Birds and Fur (Small Game) Survey and Inventory (Fur Survey Segment). Statewide Wildlife Survey and Inventory. W-130-R-11, II-4, July 1, 1979 - June 30, 1980.
    • Olson, Gary R., 1981, Game Birds and Fur (Small Game) Survey and Inventory (Fur Survey Segment). Statewide Wildlife Survey and Inventory. W-130-R-12, II-4, July 1, 1980 - June 30, 1981.
    • Olson, Gary R., 1984, Game Birds and Fur (Small Game) Survey and Inventory (Fur Survey Segment). Statewide Wildlife Survey and Inventory. W-130-R-15, II-4, July 1, 1983 - June 30, 1984.
    • Olson, R. and W.A. Hubert. 1994. Beaver: Water resources and riparian habitat manager. University of Wyoming, Laramie, Wyoming. 48pp.
    • Osmundson, C.L. 1990. Dynamics of beaver food caches and cache size as a predictor of colony size in Wyoming. M.S. thesis, University of Wyoming, Laramie.
    • Owen, C.N., D.L. Adams, and T.B. Wigley. 1984. Innefficacy of a deer repellent on beavers. The Wildlife Society Bulletin 12(4): 405-408.
    • Packard, F.M. 1940. Beaver killed by coyotes. Journal of Mammalogy 21: 359-360.
    • Packard, F.M. 1947. A survey of the beaver population of Rocky Mountain National Park, Colorado. Journal of Mammalogy 28: 219-227.
    • Paine, R.T. and S.A. Levin. 1981. Intertidal landscapes: disturbance and the dynamics of pattern. Ecol. Monogr. 51: 145-178.
    • Parker, M. 1986. Beaver, water quality, and riparian systems. Proceedings, Wyoming Water and Stream Zone Conference, Laramie, WY, University of Wyoming, Agricultural Extension Service.
    • Parker, M., F.J. Wood, Jr., B.H. Smith, and G. Robert. 1985. Erosional downcutting in lower order riparian ecosystems: have historical changes been caused by the removal of beaver? Riparian Ecosystems and their Management: Reconciling Conflicting Uses, University of Arizona, Tucson, USDA Forest Service.
    • Parker, R. R., E. A. Steinhaus, G. M. Kohls, and W. L. Jellison. 1951. Contamination of natural waters and mud with Pasteurella tularemsis and tularemia in beavers and muskrats in the northwestern United States. National Inst. of Health Bull. No. 193. 61 pp.
    • Parker, T.G., P.L. Hansen, R.C. Ehrhart, B. Thompson, and B. Haglan. Riparian and wetland ecological health evaluation of selected streams on the Charles M. Russell National Wildlife Refuge. Intermountain Journal of Science 3(4): 145.
    • Parsons, G.R. and M.K. Brown. 1978. An assessment of aerial photograph interpretation for recognizing potential beaver colony sites. New York Fish and Game Journal 25: 175-177.
    • Parsons, G.R. and M.K. Brown. 1979. Yearling reproduction in beaver as related to population density in portions of New York. Transactions of Northeast Fish and Wildlife Conference 16: 180-191.
    • Patterson, D. 1950. Beaver-trout relationships. Wisconsin Conservation Bulletin 15: 9-11.
    • Pattie, D.L. and N.A. M. Verbeek. 1967. Alpine mammals of the Beartooth Plateau. Northwest Science 41(3): 110-117.
    • Payne, N.F. 1975. Trapline management and population biology of Newfoundland beaver. Ph.D. dissertation, Utah State University, Logan. 178pp.
    • Payne, N.F. 1979. Relationship of pelt size, weight, and age for beaver. The Journal of Wildlife Management 43: 804-806.
    • Payne, N.F. 1981. Accuracy of aerial censusing for beaver colonies in Newfoundland. Journal of Wildlife Management 45: 1014-1016.
    • Payne, N.F. 1982. Colony size, age, and sex structure of Newfoundland beaver. Journal of Wildlife Management 46: 655-661.
    • Payne, N.F. 1984. Mortality rates of beaver in Newfoundland. The Journal of Wildlife Manangement 48: 117-126.
    • Payne, N.F. 1984. Population dynamics of beaver in North America. Acta Zoologica Fennica 172: 263-266.
    • Payne, N.F. and R.P. Peterson. 1986. Trends in complaints of beaver damage in Wisconsin. Wildlife Society Bulletin 14: 303-307.
    • Pearson, A.M. 1960. A study of the growth and reproduction of the beaver (Castor canadensis Kuhl) correlated with the quality and quantity of some habitat factors. M.S. thesis, University of British Columbia, Vancouver.
    • Perkins, T.E. 2000. The spatial distribution of beaver (Castor canadensis): Impoundments and effects on plant community structure in the lower Alsea drainage of the Oregon Coast Range. Master of Science Thesis.
    • Peterson, Joel, 1981, II Upland Game Bird Survey and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-12, Job No. II-3, July 1, 1980 - June 30, 1981.
    • Peterson, Joel, and Michael R. Frisina, 1985, II Upland Game Bird Survey and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-16, Job No. II-3, July 1, 1984 - June 30, 1985.
    • Peterson, Joel, and Michael R. Frisina, 1986, II Upland Game Bird Survey and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-17, Job No. II-3, July 1, 1985 - June 30, 1986.
    • Peterson, Joel, and Michael R. Frisina, 1987, II Upland Game Bird and Inventory, Region Three, Game Birds and Fur (Small Game) Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-18, Job No. II-3, July 1, 1986 - June 30, 1987.
    • Peterson, R.P. 1979. Nuisance beaver biology and control in north-central Wisconsin. M.S. thesis, University of Wisconsin, Steven's Point.
    • Peterson, R.P. and N.F. Payne. 1986. Productivity, size, age, and sex structure of nuisance beaver colonies in Wisconsin. The Journal of Wildlife Management 50: 265-268.
    • Pickett, S.T.A. and P.S. White. 1985. The ecology of natural disturbance and patch dynamics. Academic Press, New York.
    • Pinay, G. and R.J. Naiman. 1991. Short term hydrologic variations and nitrogen dynamics in beaver created meadows. Arch. Hydrobiol. 123: 187-205.
    • Pinkowski, B. 1983. Foraging behavior of beavers (Castor canadensis) in North Dakota. Journal of Mammalogy 64: 312-314.
    • Pollock, M.M., C.R. Pess, and T.J. Beeche. 2003. The importance of beaver ponds to coho salmon production in the Stillaguamish River Basin, Washington, USA. National Oceanic and Atmospheric Administration, Northwest Fisheries Science Center, Seattle, WA.
    • Pollock, M.M., R.J. Naiman, H.E. Erickson, C.A. Johnston, J. Pastor, and G. Pinay. 1995. Beaver as engineers: Influences on biotic and abiotic characteristics of drainage basins. Linking Species & Ecosystems. In: Jones, C.G. and J.H. Lawton (Ed.). Chapter 12: 117-126
    • Potvin, C.L. and M. Bovet. 1975. Annual cycle of patterns of activity rythms in beaver colonies (Castor canadensis). J. Comp. Physiol. 98: 248-256.
    • Potvin, F., L. Breton, and M. MacQuart. 1992. Impact of an experimental wolf reduction on beaver in Papineau-LaBelle Reserve, Quebec. Canadian Journal of Zoology 70: 180-183.
    • Powder River Eagle Studies, Inc., Gillette, WY., 1996, Spring Creek Mine 1995 Wildlife Monitoring Studies. Spring Creek Coal Company 1995-1996 Mining Annual Report. Vol. I, App. I. May 1996.
    • Pringle, C.M. 1997. Exploring how disturbance is transmitted upstream: going against the flow. J.N. American Benthological Society 16(2): 425-438.
    • Pritchard, G. and H.A. Hall. 1971. An introduction to the biology of craneflies in a series of abandoned beaver ponds, with an account of the life cycle of Tipula sacra Alexander (Diptera: Tipulidae). Canadian Journal of Zoology 49: 467-482.
    • Provost, E.E. 1958. Studies on reproduction and population dynamics in beaver. Unpubl. Ph.D. Diss., Washington State University, Pullman. 78p.
    • Pullen, T.M., Jr. 1971. Some effects of beaver (Castor canadensis) and beaver pond management on the ecology and utilization of fish populations along warm-water streams in Georgia and South Carolina. Unpubl. Ph.D. dissertation, University of Georgia, Athens. 84pp.
    • Purdy, K.G., D.J. Decker, R.A. Malechi, and J.C. Proud. 1985. Landowner tolerance of beavers: implications for damage management and control. Pages 83-88 in Proceedings of the Second Eastern Wildlife Damage Control Conference, North Carolina State University, Raleigh, North Carolina, USA.
    • Rabe, F.W. 1970. Brook trout populations in Colorado beaver ponds. Hydrobiologia 35(3/4): 431-448.
    • Randall, D. 1985. A new role for beavers. Defenders 58(3): 29.
    • Rasmussen, D.I. 1940. Beaver-trout relationship in the Rocky Mountain region. Transactions of the North American Wildlife Conference 5: 256-263.
    • Rawinski, T.J. and B.P. Lapin. 1990. Some affects of flooding on rare swamp birch (Betula pumila L.) vegetation in Connecticut: 1984-1987. Pages 244-247 in; Ecosystem Management: are species and significant habitats. New York State Museum Bulletin 471.
    • Rawley, E.V. 1954. Utah Beaver Transplanting Manual. Department Information Bulletin No. 12. Utah State Department of Fish and Game, Salt Lake City. 19pp.
    • Ray, A.M. 2001. Macrophyte succession in Minnesota beaver ponds. Canadian Journal of Botany 79(4): 487-499.
    • Rea, A.M. 1983. Once a river: bird life and habitat changes on the Middle Gila. Tucson, AR: University of Arizona Press. 285p.
    • Rebertus, A.J. 1982. The ecology of beavers in bog habitats of north central Minnesota. M.S. Thesis, Louisiana State University, Baton Rouge. 90p.
    • Reese, K.P. and J.D. Hair. 1976. Avian species diversity in relation to beaver pond habitats in the Piedmont region of South Carolina. Proc. Annual Conf. Southeast. Fish and Wildl. Agencies 30: 437-447.
    • Reid, F. 2006. Peterson Field Guide to Mammals of North America, 4th Edition. Houghton Mifflin Company: Boston and New York, 608 pp.
    • Reid, K.A. 1952. Effects of beavers on trout waters. Maryland Conserv. 29(4): 21-23.
    • Reinert, J.H. 2020. Biotic and physical responses to biomimicry structures in a Rocky Mountain incised stream. M.Sc. Thesis. Bozeman, MT: Montana State University. 45 p.
    • Reinhart, D.P. 1990. Grizzly bear habitat use on cutthroat trout spawning streams in tributaries of Yellowstone Lake. M.Sc. Thesis. Bozeman, MT: Montana State University. 128 p.
    • Remillard, M.M., G.K. Gruendling, and D.J. Bogucki. 1987. Disturbance by beaver (Castor candensis Kuhl) and increased landscape heterogeneity. Pages 103-122 in M.G. Turner, editor. Ecological Studies 64. Landscape Heterogeneity and disturbance. Springer-Verlag, New York.
    • Renouf, R.N. 1972. Waterfowl utilization of beaver ponds in New Brunswick. Journal of Wildlife Management 36: 740-744.
    • Retzer, J.L. 1955. Physical environmental effects on beavers in the Colorado Rockies. Western Assoc. State Game and Fish Comm. Proc. 35: 279-287.
    • Retzer, J.L., H.M. Swope, J.D. Remington, and W.H. Rutherford. 1956. Suitability of physical factors for beaver management in the Rocky Mountains of Colorado. Colorado Department of Game and Fish Tech. Bulletin 2. 33pp.
    • Reynolds, R.E. and J.C. Lewis. 1976. Evaluating beaver guards on restricted flow risers of flood control impoundments. Ann. Conf. Southeast. Assoc. Fish and Game Comm. 13: 455-462.
    • Richard, P.B. 1983. Mechanisms and adaption in the constructive behavior of the beaver C. fiber L. Acta Zool. Fennica 174: 105.
    • Riley, C. W. 1996. Evaluation of trout spawning gravel restoration in a placer-mined Montana stream. M.Sc. Thesis. Bozeman: Montana State University. 97 p.
    • Ripple, W.J. and E.J. Larson. 2000. Historic aspen recruitment, elk, and wolves in northern Yellowstone National Park, USA. Biological Conservation 95: 361-370.
    • Ritter, Torrey. 2018. Ecosystem pioneers: Beaver dispersal and settlement site selection in the context of habitat restoration. M.Sc. Thesis. Bozeman, MT: Montana State University. 212 p.
    • Robel, R.J. 1993. Comparison of aerial and ground survey techniques to determine beaver colony densities in Kansas. Southwestern Naturalist 38(4): 357-361.
    • Robel, R.J., L.B. Fox, and K.E. Kemp. 1993. Relationship between habitat suitability index values and ground counts of beaver colonies in Kansas. Wildlife Society Bulletin 21: 415-421.
    • Roberts, T.H. and D.H. Arner. 1984. Food habits of beaver in east-central Mississippi. Journal of Wildlife Management 48: 1414-1419.
    • Robertus, A.J. 1985. Beaver engineering in northern bogs. The Minnesota volunteer 48:17-20.
    • Robertus, A.J. 1986. Bogs as beaver habitat in north-central Minnesota. American Midland Naturalist 116(2): 240-245.
    • Robinson, D. 1978. Beaver builders: sturdy dams shaped our land and history. Defenders 54(3): 153-160.
    • Roblee, K.J. 1983. A wire mesh culvert for use in controlling water levels at nuisance beaver sites. Proceedings of the First Eastern Wildlife Damage Control Conference 1: 167-168.
    • Rolauffs, P., D. Hering, and S. Lohse. 2001. Composition, invertebrate community, and productivity of a beaver dam in comparison yo other stream habitat types. Hydrobiologia 459: 201-212.
    • Rosell, F. and B. Hovde. 2001. Methods of aquatic and terrestrial netting to capture Eurasian beavers. Wildlife Society Bulletin 29: 269-274.
    • Rosell, F. and J.K. Kvinlaug. 1998. Methods for live trapping beaver (Castor spp.). Fauna Norvegica Serie A 19: 1-28.
    • Rosell, F. and L. Sun. 1999. Use of anal gland secretion to distinguish the two beaver species, Castor canadensis and C. fiber. Wildlife Biology 5: 119-123.
    • Rothmeyer, S. W., M. C. McKinstry, and S. H. Anderson. 2002. Tail attachment of modified ear-tag radio transmitters on beavers. Wildlife Society Bulletin 30: 425-429.
    • Rowe, J.S. and G.W. Scotter. 1973. Fire in the boreal forest. Quaternary Research 3(3): 444-464.
    • Rudersdorf, W.J. 1952. The coactions of beaver and moose on a joint food supply in the Buffaloa River Meadows. Utah State University, Logan, Utah.
    • Rue, L.L. 1964. The world of the beaver. Living World Books, J.B. Lippincott Co., Philadelphia.
    • Ruedemann, R. and W.J. Schoonmaker. 1938. Beaver dams as geologic agents. Science 88: 523-525.
    • Runyon, Molly J. 2013. Effects of beaver reintroduction and ungulate browsing on aspen recovery in the Eagle Creek drainage of the northern Yellowstone winter range. Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 81 p.
    • Rupp, R.S. 1954. Beaver-trout relationship in the headwaters of Sunkhaze Stream, Maine. American Fisheries Society: 75-85.
    • Russell, K.R., C.E. Mooreman, J.K. Edwards, B.S. Metts, and D.C. Guynn, Jr. 1998. Amphibian and reptile communities associated with beaver (Castor candensis) ponds and unimpounded streams in the Piedmont of South Carolina. Clemson University, Clemson, South Carolina. p149-158.
    • Russell, K.R., C.E. Moorman, J.K. Edwards, B.S. Metts, D.C. Guynn, Jr. 1999a. Amphibian and reptile communities associated with beaver (Castor canadensis) ponds and unimpounded streams in the Piedmont of South Carolina. Journal of Freshwater Ecology 14(2): 149-158.
    • Rust, H. J. 1946. Mammals of northern Idaho. J. Mammal. 27(4): 308-327.
    • Rutherford, W.H. 1955. Wildlife and environmental relationships of beavers in Colorado forests. Journal of Forestry 53(11): 803-806.
    • Rutherford, W.H. 1964. The beaver in Colorado: its biology, ecology, management, and economics. Corado Game, Fish, and Parks Department Tech. Bulletin 17. 48pp.
    • Rutten, M.G. 1967. Flat-bottomed glacial valleys, braided rivers and the beaver. Geol. Mijnbouw 46: 356-360.
    • Ryden, H. 1988. The beaver is one smart rat. Audubon, September, pages 96-105.
    • Salyer, J.C. 1935. Preliminary report on the beaver-trout investigation. American Game 24(6): 13-15.
    • Sanner, C.J. 1987. Effects of beaver on Coho Salmon habitat, Kenai Peninsula, Alaska. M.S. thesis, Oregon State University, Corvallis. 81pp.
    • Sawyer, H. E. 1935. Studies on the rodents of Montana. M.A. thesis. University of Montana, Missoula. 111 pp.
    • Schlosser, I. J. and L. W. Kallemeyn. 2000. Spatial variation in fish assemblages across a beaver-influenced successional landscape. Ecology 81: 1371-1382.
    • Schlosser, I.J. 1995. Dispersal, boundary processes, and trophic-level interactions in streams adjacent to beaver ponds. Ecology 76: 908-925.
    • Schubloom, L.A. 1995. Lichens as air quality indicators in three areas of southwestern Montana: lichen floristics and elemental analysis. M.Sc. Thesis. Bozeman, MT: Montana State University. 130 p.
    • Schulte, B.A. 1993. Chemical communication and ecology of the North American beaver (Castor canadensis). Ph.D. dissertation, State University of New York, College of Environmental Science and Forestry, Syacruse, New York. 194pp.
    • Science '83. 1983. Beavers: a dam site better on erosion. Science 4(9): 7.
    • Science Digest. 1984. New faith in beaver ecology. Science Digest 92(7): 36.
    • Scrafford, M.A., D.B. Tyers, D.T. Patten, and B.F. Sowell. 2018. Rangeland Ecology & Management 72(2):266-273.
    • Semyonoff, B.T. 1951. The river beaver in Archangel Province. Pages 5-45 in Russian game report. Vol. 1. Can. Dep. North. Aff. And Nat. Resour., Ottawa, Ont.
    • Shadle, A.R. 1954. Sizes of wood cuttings handled by beaver. American Midland Naturalist 52: 510-512.
    • Shadle, A.R. And T.S. Austin. 1939. Fifteen months of beaver work at Allegany State Park, New York. Journal of mammalogy 20: 299-303.
    • Shadle, A.R., A.M. Nauth, E.C. Gese, and T.S. Austin. 1943. Comparison of tree cuttings of six beaver colonies in Allegany State Park, New York. Journal of Mammalogy 24: 32-39.
    • Shaw, S.P. 1948. The beaver in Massachusetts. Federal Aid in Wildlife Restoration. Massachusetts Department of Conservation Research Bulletin No. 11.
    • Shay, R. 1978. Oregon's beaver. Oregon Wildlife 33(2): 3-5.
    • Shelton, P.C. 1966. Ecological studies of beaver, wolves, and moose in Isle Royal National Park, Michigan. Ph.D. dissertation, Purdue University, Lafayette, Indiana.
    • Shelton, P.C. and R.O. Peterson. 1983. Beaver, wolf, and moose interactions in Isle Royale National park, U.S.A. Acta Zool. Fenn. 174: 265-266.
    • Sidorovich, V.E., B. Jedrzejewska, and W. Jedrzejewski. 1996. Winter distribution and abundance of mustelids and beavers in the river valleys of Bialowieza-Primeval-Forest. Acta Theriologica 41: 155-170.
    • Simonsen, T.A. 1973. Feeding ecology of the beaver (Castor fiber L.) (in Norwegian, summary in English). Papers of the Norwegian State Game Research Institute, Second Series, No. 39: 1-61.
    • Skelly, D.K. and L.K. Freidenburg. 2000. Effects of beaver on the thermal biology of an amphibian. Ecology Letters 3: 483-486.
    • Skinner, Q.D., J.E. Speck Jr., M. Smith, and J.C. Adams. 1984. Stream water qualityas influenced by beaver within grazing systems in Wyoming. Journal of Range Management 37(2): 142-146.
    • Slough, B.G. 1978. Beaver food cache structure and utilization. Journal of Wildlife Management 42(3): 644-646.
    • Slough, B.G. and R.M.F.S. Sadlier. 1977. A land capability classification system for beaver (Castor canadensis). Canadian Journal of Zoology 55: 1324-1335.
    • Smith, A.E. 1950. Effects of water run-off and gradient on beaver in mountain streams. M.S. thesis, University of Michigan, Ann Arbor.
    • Smith, B.H. 1980. Not all beaver are bad; or, an ecosystem approach to stream habitat management, with possible software applications. In: Proceedings 15th Annual Meeting of Colorado-Wyoming Chapter of American Fisheries Society (edited by Whaley, R.). pp 32-37.
    • Smith, D. 1997. Dispersal strategies and cooporative breeding in beavers. Dissertation Abstracts International 58(4), Part B: Science and Engineering: p1649.
    • Smith, D. W. 1998. Beaver Survey: Yellowstone National Park 1998. National Park Service Report.
    • Smith, D.W. and R.O. Peterson. 1987. A survey of beaver ecology in Grand Portage National Monument, Minnesota. Contract No. 6000-7-8022. 44pp.
    • Smith, D.W. and R.O. Peterson. 1988. The effects of regulated lake levels on beaver in Voyageurs National Park, Minnesota National Park Service Research/Researches Management Report MWR-11. 84pp.
    • Smith, D.W. and R.O. Peterson. 1991. Beaver ecology in Apostle Islands National Lakeshore. Final Report to U.S. National Park Service. 64pp.
    • Smith, D.W. and R.O. Peterson. 1991. Behavior of beaver in lakes with varying water levels in northern Minnesota. Environmental Management 15: 395-401.
    • Smith, D.W., D.R. Trauba, R.K. Anderson, and R.O. Peterson. 1994. Black bear predation on beavers on an island in Lake Superior. American Midland Naturalist 132: 248-255.
    • Smith, M.E., C.T. Driscoll, B.J. Wykowski, C.M. Brooks, and C.C. Costentini. 1991. Modification of stream ecosystem structure and function by beaver (Castor canadensis) in the Adirondack Mountains, New York. Canadian Journal of Zoology 69: 55-61.
    • Smith, M.M. and C.S. Goldberg. 2022. Facilitative interaction promotes occupancy of a desert amphibian across a climate gradient. Conservation Ecology original research. Oecologia. https://doi.org/10.1007/s00442-022-05127-6. 10 p.
    • Smock, L.A., G.M. Metzler, and J.E. Gladden. 1989. Role of debris dams in the structure and functioning of low-gradient headwater streams. Ecology 70: 764-775.
    • Snigg, M. 1986. Pole Mountain beaver pond re-inventory. Wyoming Game and Fish Department, Project Report 5085-7301, Cheyenne.
    • Snodgrass, J. W. 1997. Temporal and spatial dynamics of beaver-created patches as influenced by management practices in a south-eastern North American landscape. Journal of Applied Ecology 34: 1043-1056.
    • Snodgrass, J.W. 1996. The influence of beaver ponds on the temporal and spatial dynamics of southeastern fish assemblages. Ph.D. Dissertation, University of Georgia, Athens.
    • Snodgrass, J.W. and G.K. Meffe. 1998. Influence of beavers on stream fish assemblages: effects of pond age and watershed position. Ecology 79: 928-942.
    • Sonster-Alpin, M.S. and R.L. Klotz. 1995. A comparison of electron transport system activity in stream and beaver pond sediments. Can. J. Fish. Aquat. Sci. 52: 1318-1326.
    • South, A., S. Rushton, and D. Macdonald. 2000. Simulating the proposed reintroduction of the European beaver (Castor fiber) to Scotland. Biological Conservation 93: 103-116.
    • Spence, J.A. and H.B.N. Hynes. 1971. Differences in benthos upstream and downstream of an impoundment. J. Fish. Board. Can. 28: 35-43.
    • Spieth, H. T. 1979. The Virilis Group of Drosophila and the Beaver Castor. American Naturalist 114: 312-316.
    • Sprules, W.M. 1940. The effect of a beaver dam on the insect fauna of a trout stream. American Fisheries Society: 236-248.
    • Stenlund, M. 1953. Report of Minnesota beaver die-off, 1951-52. Journal of Wildlife Management 17: 376-377.
    • Stephenson, A.B. 1956. Preliminary studies on growth, nutrician, and blood chemistry of beavers. M.S. Thesis, University of British Columbia, Vancouver.
    • Stephenson, A.B. 1969. Temperatures within a beaver lodge in winter. Journal of Mammaology 50: 134-136.
    • Steubner, S. 1992. Leave it to beaver. Paonia, Colorado, High Country News 24(15).
    • Steubner, S. 1994. Bullish on beaver. National Wildlife 32(3): 24-27.
    • Stewart, Shawn T., 1978, II Game Birds and Fur Survey and Inventory - Region 5. Statewide Wildlife Survey and Inventory. W-130-R-8, II, 5, July 1, 1976 - June 30, 1977.
    • Stirton, R.A. 1935. A review of the Tertiary beavers. Calif. Publ. Bull. Dept. Geol. Sci. 23: 391-458.
    • Stock, A.D. 1970. Notes on mammals of southwestern Utah. Journal of Mammalogy 51(2): 429-433.
    • Stock, J.D. and I.J. Schlosser. 1991. Short-term effects of a catastrophic beaver dam collapse on a stream fish community. Environmental Biology of Fishes 31: 123-129.
    • Stocker, G. 1983. Problems in utilization of plant food resources by the beaver. Revue Suisse de Zoologie 90: 487-496.
    • Svendsen, G. E. 1980. Population Parameters and Colony Composition of Beaver (Castor Canadensis) in Southeast Ohio. American Midland Naturalist 104: 47-56.
    • Svendsen, G. E. 1989. Pair formation, duration of pair-bonds, and mate replacement in a population of beavers (Castor canadensis). Canadian Journal of Zoology 67: 336-340.
    • Svendson, G.E. 1978. Castor and anal glands of beaver (Castor canadensis). Journal of Mammalogy 59: 619-620.
    • Svendson, G.E. 1980. Seasonal change in feeding patterns of beaver in southeastern Ohio. Journal of Wildlife Management 44: 285-290.
    • Swank, W.E. 1949. Beaver ecology and management in West Virginia. Conserv. Comp. WV. 65pp.
    • Swank, W.G. and F.A. Glover. 1948. Beaver censusing by airplane. Journal of Wildlife Management 12: 214.
    • Swenson, J.E. and S.J. Knapp. 1980. Composition of beaver caches on the Tongue River in Montana. The Prarie Naturalist 12(2): 33-36.
    • Swenson, J.E., S.J. Knapp, P.R. Martin, and T.C. Hinz. 1983. Reliability of aerial cache surveys to monitor beaver population trends on prarie rivers in Montana. Journal of Wildlife Management 47(3): 697-703.
    • Syphard, A.D., and M.W. Garcia. 2001. Human and beaver-induced wetland changes in the Chickahominy River watershed from 1953-1994. Wetlands 21(3): 342-353.
    • Tappe, D.T. 1942. The status of beavers in California. California Division of Fish and Game. Game Bulletin No. 3. 59p.
    • Taylor, D. 1970. Growth, decline, and equilibrium in a beaver population at Sagehen Creek, California. Ph.D. dissertation, University of California, Berkeley.
    • Taylor, D. 1971. Beaver population studies at Sagehen Creek. In: Transactions California-Nevada Section, The Wildlife Society; 1971 January 29-30; Sacramento, CA. Smartsville, CA: The Wildlife Society: 18-19.
    • Taylor, K.P. 1983. Factors influencing beaver management in rural Alaska-northern Bristol Bay. Acta Zoologica Fennica 174: 127-128.
    • Teaford, J.W. 1986. Beaver pond management. Section 5.5.2, U.S. Army Corps of Engineers Wildlife Resources Manual, Technical Report EL-86-10, Waterways Experiment Station, P.O. Box 631, Vickburg, Mississippi 39180-0631.
    • Terwilliger, J. and J. Pastor. 1999. Small mammals, ectomycorrhizae, and conifer succession in beaver meadows. Oikos 85: 83-94.
    • Tevis, L., Jr. 1950. Summer behavior of a family of beavers in New York state. Journal of Mammalogy 31: 40-65.
    • Thomas, A.E. 1986. Riparian protection/enhancement in Idaho. Rangelands 8(5): 224-227.
    • Thompson, L.S. 1981. Circle West wildlife monitoring study: Third annual report. Technical report No. 8. Montana Department of Natural Resources and Conservation. Helena, Montana.
    • Thompson, Richard W., Western Resource Dev. Corp., Boulder, CO., 1996, Wildlife baseline report for the Montana [Montanore] Project, Lincoln and Sanders counties, Montana. In Application for a Hard Rock Operating Permit and Proposed Plan of Operation, Montanore Project, Lincoln and Sanders Counties, Montana. Vol. 5. Stroiazzo, John. Noranda Minerals Corp., Libby, MT. Revised September 1996.
    • Thorniley, M. 1972. The dam builders. Pacific Search 7(3): 4-5.
    • Todd, A.W. 1981. Ecological arguments for fur-trapping in boreal wilderness regions. Wildlife Society Bulletin 9(2): 116-124.
    • Todd, P.A. 1992. Mountain beaver habitat use and management implications in Yosemite National Park. Natural Areas Journal 12(1): 26-31.
    • Townsend, J. E. 1952. A study in Beaver ecology in Western Montana with special reference to movements. M.Sc. Thesis. Bozeman, Montana: Montana State University. 44 p.
    • Townsend, J. E. 1953. Survey of beaver distribution and status in the Jefferson Basin. Montana Fish and Game Dept., P-R Quarterly Report. October-December:62-68.
    • Townsend, J.E. 1953. Beaver ecology in western Montana with special referrence to movements. Journal of Mammalogy 34(4): 459-479.
    • Townsend, J.E. 1955. Fur resources surveys and investigations: beaver ecology in the Jefferson basin. Montana Fish and Game Department, Project # MONT W-049-R-04/W.K.P.L.02/JOB C. 8p.
    • Townsend, J.E. and F.E. Newby. 1955. Montana beaver management. Proc. 35th Ann. Conf. W. Assn. State Game & Fish Comms., Project# MT W-049-R. p 269-278.
    • Townsend, P.A. and D.R. Butler. 1996. Patterns of landscape use by beaver on the lower Roanoke River floodplain, North Carolina. Physical Geography 17: 253-269.
    • Trueblood, R. W. 1964-65. Beaver population and distribution trends - aerial survey. Montana Fish and Game Dept. P-R Job Compl. Repts. Proj. W-76-R-I0, 11, Job D-2 (Multilith).
    • Trueblood, Richard W., 1977, II Game Birds and Fur (Small Game Survey and Inventory), Region Six Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-8, II, 6, July 1, 1976 - June 30, 1977.
    • Trueblood, Richard W., 1979, II Game Birds and Fur (Small Game Survey and Inventory), Region Six Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-9, II, 6, July 1, 1977 - June 30, 1978.
    • Trueblood, Richard W., 1980, II Game Birds and Fur (Small Game Survey and Inventory), Region Six Survey and Inventory. Statewide Wildlife Survey and Inventory. W-130-R-11, II, 6, July 1, 1979 - June 30, 1980.
    • Tufts, P.D. 1967. A comparative study of some environmental factors affecting beaver colony site locations. M.S. Thesis, Acadia University, Wolfville, Nova Scotia. 174pp.
    • TVX Mineral Hill Mine, Amerikanuak, Inc., Gardiner, MT., 2002, Yearly summary of wildlife observation reports. 1990-2002 Letter reports.
    • Tweten, R.G. 1984. Baseline survey of furbearing mammals within the South Fork drainage Sun River, Montana. M.Sc. Thesis. Bozeman, MT: Montana State University. 63 p.
    • USDI Fish and Wildlife Service., 1961, A Detailed report on fish and wildlife resources affected by McNamara Dam and Reservoir, Blackfoot River Project, Montana. June 1961.
    • Van Deelen, T. R. 1991. Dispersal patterns of juvenile beavers in western Montana. Thesis. University of Montana, Missoula, Montana. 91 p.
    • Van Deelen, T. R. and D. H. Pletscher. 1996. Dispersal Characteristics of Two-year-old Beavers, Castor canadensis, in Western Montana. The Canadian Field-Naturalist 110: 318-321.
    • Vanden Berge, R.J. and P.A. Vohs, Jr. 1977. Population status of beaver on the free-running Missouri River in southeastern South Dakaota. Proc. S.D. Acad. Sci. 56: 230-236.
    • Vogt, B. 1981. What ails the river otter? National Wildlife 19(2): 25-28.
    • Vore, J. 1993. Guidelines for the reintroduction of beavers into southwest Montana streams. Montana Department of Fish, Wildlife and Parks, Townsend, Montana. 34pp.
    • Warren, E. R. 1926. A study of the beavers in the Yancey region of Yellowstone National Park. Roosevelt Wildlife Annals 1: 1-191.
    • Warren, E.R. 1905. Some interesting beaver dams in Colorado. Proc. Wash. Acad. Sci. 6: 429-437.
    • Warren, E.R. 1927. The beaver: its works and ways. London: Bailliere, Tindall & Cox.
    • Waterman, C.F. 2001. Beaver ponds and trout. Montana Outdoors 32(4): 19-22.
    • Weckwerth, R. P. 1957. The relationship between the marten population and the abundance of small mammals in Glacier National Park. M.S. thesis. University of Montana, Missoula. 76 pp.
    • WESTECH Env. Services, Inc., Helena, MT., 2000, Reconnaissance of Wildlife and Fisheries Resources at the Ruby Garnet Alder Gulch Property. May 24, 2000. In Application for an Operating Permit and Proposed Plan of Operations, Alder Gulch Mine Project, Mad
    • Westech, Inc. [Western Technology and Engineering]. 1989. Reconnaissance of terrestrial wildlife resources in the Pauper's Dream project vicinity, Aug. 1988. Prepared for Hydrometrics, Inc., Helena, MT. 22 pp.
    • Westech, Inc. [Western Technology and Engineering]. No date. Preliminary wildlife reconnaissance, Ruby and Little Ben mine areas, Little Rocky Mountains, Montana. Technical Report for Zortman and Landusky Mining Companies.
    • Western Technology and Engineering, Inc. (WESTECH). 1994. Wildlife Monitoring Absaloka Mine Area Annual Report, 1993. Montana SMP 85005. OSMP Montana 0007c. Mar. 12, 1994.
    • Western Technology and Engineering, Inc., Helena, MT., 1989, Reconnaissance of the wildlife resources in the vicinity of the Kendall Venture Mine. January 1989. In Kendall Venture North Moccasin Project: Amendment to Operating Permit 00122, Fergus County, Montana. Vol. 2, App. A, Feb., 1989.
    • Wetlands West, Inc., Bozeman, MT., and Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Lavina Wetland, Lavina, Montana. Proj. No. 130091.030. February 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Wetlands West, Inc., Bozeman, MT., and Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Stillwater River, Absarokee, Montana. Proj. No. 130091.032. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. II.
    • Wetlands West, Inc., Bozeman, MT., and Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Wyola-Sunlight Ranch Wetland, Wyola, Montana. Proj. No. 130091.035. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. II.
    • Wheatley, M. 1997. Beaver, Castor canadensis, Home Range Size and Patterns of Use in the Taiga of Southeastern Manitoba: II. Sex, Age, and Family Status. The Canadian Field-Naturalist 111: 211-216.
    • White, D.S. 1990. Biological relationships to convective flow patterns within stream beds. Hydrobiologia 196: 149-158.
    • White, P.S. and S.T.A. Pickett. 1985. Natural disturbance and patch dynamics: an introduction. In: Pickett, S.T.A. and P.S. White; editors, The ecology of natural disturbance and patch dynamics. Academic Press, pp. 3-13.
    • Whitehead, B.K. 2020. The response of parafluvial soils to beaver mimicry restoration in a montane stream. M.Sc. Thesis. Bozeman, MT: Montana State University. 114 p.
    • Wigley, T.B. and M.E. Garner. 1987. Impact of beavers in the Arkansas Ozarks. Report Series 298. Arkansas Agricultural Experiment Station, University of Arkansas, Fayetteville. 12pp.
    • Wigley, T.B., T.H. Roberts, and D.H. Arner. 1983. Reproductive characteristics of beaver in Mississippi. Journal of Wildlife Management 47: 1172-1177.
    • Wilde, S.A., C.T. Youngberg, and H.H. Hovind. 1950. Changes in compositions of ground water, soil fertility, and forest growth produced by the construction and removal of beaver dams. Journal of Wildlife Management 14: 123-128.
    • Wiley, G.M. 1991. Aquatic invertebrate communities of beaver dams in a brown-water stream of Alberta. M.S. thesis, University of Alberta, Edmonton, Alberta.
    • Willis, R.M. A beaver habitat classification system for the Truckee River. M.S. thesis, University of Nevada, Reno. 69pp.
    • Wilsson, L. 1971. Observations and experiments on the ethology of the European beaver (Castor fiber L.). Viltrevy 8(3): 115-266.
    • Winkle, P.L. and W.A. Hubert. 1990. Relations between brook trout standing stocks and habitat features in beaver ponds in southeastern Wyoming. North American Journal of Fisheries Management 10: 72-79.
    • Winston, M.R., C.M. Taylor, and J. Pigg. 1991. Upstream extirpation of four minnow species due to damming of a prairie stream. Transactions of the American Fisheries Society 120: 98-105.
    • Wolff, S.W., T.A. Wesch, and W.A. Hubert. 1989. Stream channel and habitat changes due to flow augmentation. Regulated Rivers, Research and Management 4: 225-233.
    • Woo, M. and J.M. Waddington. 1990. Effects of beaver dams on subarctic wetland hydrology. Arctic 43(3): 223-230.
    • Woodward, D.K. 1977. Status and ecology of the beaver (Castor canadensis) in South Carolina with emphasis on the Piedmont region. M.S. thesis, Clemson University, Clemson, South Carolina. 208pp.
    • Woodward, D.K. and R.B. Hazel. 1991. Beavers in North Carolina: ecology, utilization, and management. Publication No. AG-434. North Carolina State University, Coooperative Extension Service, Raleigh. 21pp.
    • Woodward, D.K., J.D. Hair, and B.P. Gaffney. 1976. Status of beaver in South Carolina as determined by a postal survey of landowners. Proceedings, Annual Conference, Southeastern Association of Fish and Wildlife Agencies 30: 448-454.
    • Woodward, D.K., R.B. Hazel, and B.P. Gaffney. 1985. Economic and environmental impacts of beaver in North Carolina. Pages 89-96 in P.T. Bromley (ed.), Proceedings of the Second Eastern Wildlife Damage Control Conference, Raleigh, North Carolina.
    • World Wildlife Illustrated. 1970. The American beaver (Castor candensis). World Wildlife Illustrated 2(1): 4-6.
    • Wotton, R.S. 1992. Animals that exploit lake outlets. Freshwater Forum 2: 62-76.
    • Wright, J.P., C.G. Jones, and A.S. Flecker. 2002. An Ecosystem engineer, the beaver, increases species richness at the landscape scale. Oecologia 132: 96-101.
    • Yavitt, J.B., G.E. Lang, and A.J. Sexstone. 1990. Methane fluxes in wetland and forest soils, beaver ponds, and low-order streams of a temperate forest ecosystem. J. Geophys. Res. 95: 22463-22474.
    • Yavitt, J.B., L.L. Angell, T.J. Fahey, C.P. Cirmo, and C.T. Driscoll. 1992. Methane fluxes, concentrations, and production in two Adirondack beaver impoundments. Limnology and Oceanography 37: 1057-1066.
    • Yeager, L.E. and K.G. Hay. 1955. A contribution toward a bibliography on the beaver. Tech Bull State of Colorado Department of Fish and Game 1 : 1-103.
    • Yeager, L.E. and R.R. Hill. 1954. Beaver management problems on western public lands. Transactions of the North American Wildlife and Natural Resources Conference 19: 462-479.
    • Yeager, L.E. and W.H. Rutherford. 1957. An ecological basis for beaver management in the Rocky Mountain Region. Transactions of the North Amrican Wildlife and Natural Resources Conference 22: 269-299.
    • Zurowski, W. 1992. Building activity of beavers. Acta Theriologica 37: 403-411.
    • Zurowski, W. and B. Kasperczyk. 1986. Characteristics of a European beaver population in the Suwalki Lakeland. Acta Theriologica 34: 311-321.
    • Zurowski, W. and B. Kasperczyk. 1988. Effects of reintroduction of European beaver in the lowlands of the Vistula Basin. Acta Theriologica 33: 325-338.
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