Eastern Red Bat - Lasiurus borealis
State Rank Reason (see State Rank above)
Recent surveys using acoustic detectors have shown this species to be present across much of central and eastern Montana during the summer and fall. Tree roosting bat species, including the Eastern Red Bat, are commonly killed at wind farms, which presents a substantial threat to the long-term viability of populations within the state.
- Details on Status Ranking and Review
ScoreG - 200,000-2,500,000 km squared (about 80,000-1,000,000 square miles)
Comment223,758 square Kilometers from Natural Heritage Program range maps
ScoreE - Relatively Stable (±25% change)
CommentHabitat is likely stable within +/- 25% since European settlement. Although deciduous forests have declined since European settlement due to grazing and invasive species, it is unlikely that the habitat for this species has been reduced by more then 25% within the state given that pine forest is also used.
ScoreU - Unknown. Short-term trend in population, range, area occupied, and number and condition of occurrences unknown.
CommentNo data on trends available.
ScoreB - Moderate and imminent threat. Threat is moderate to severe and imminent for a significant proportion (20-60%) of the population or area.
CommentThis species has been shown to be among those commonly killed at wind energy facilities in other areas. There are no documented mortalities of this species at wind energy facilities in Montana, but data on bat mortalities at these facilities are limited. The population effects of mortalities has not been established, but there is the potential for severe declines given the species relatively low fecundity.
SeverityModerate - Major reduction of species population or long-term degradation or reduction of habitat in Montana, requiring 50-100 years for recovery.
CommentIf complete mitigation of mortalities were implemented today, the species would likely recover relatively quickly (Low), however development and implementation of effective mitigation strategies may not occur for a number of years (if at all), so recovery will likely take much longer (Moderate)
ScopeModerate - 20-60% of total population or area affected
CommentAlthough wind energy facilities are not currently present within much of the state-wide range for this species, they are placed in areas that concentrate individuals during migration and the impacted area is increasing as more facilities are built. Turbines also appear to attract individuals, increasing the effective footprint of these facilities and their impacts to tree roosting bat species.
ImmediacyHigh - Threat is operational (happening now) or imminent (within a year).
CommentThreat is ongoing and increasing
ScoreB - Moderately Vulnerable. Species exhibits moderate age of maturity, frequency of reproduction, and/or fecundity such that populations generally tend to recover from decreases in abundance over a period of several years (on the order of 5-20 years or 2-5 generations); or species has moderate dispersal capability such that extirpated populations generally become reestablished through natural recolonization (unaided by humans).
CommentSpecies can have 1-2 pups each year, although individuals may not breed each year. Juvenile mortality may be moderate to high. Species has good dispersal capabilities and reestablishment of extirpated populations through dispersal is possible.
ScoreB - Narrow. Specialist. Specific habitat(s) or other abiotic and/or biotic factors (see above) are used or required by the Element, but these key requirements are common and within the generalized range of the species within the area of interest.
CommentSpecies is dependent on major riparian areas with cottonwoods or other deciduous trees for roosting.
Raw Conservation Status Score
3.5 + 0 (geographic distribution) + 0 (environmental specificity) + 0 (long-term trend) + -0.75 (threats) = 2.75
The Eastern Red Bat is a moderately-sized lasurine (7 to 15 g) with long pointed wings and heavily-furred interfemoral membrane. Pelage overall is reddish, lighter on the belly than the back. Ears are low and rounded, tragus triangular, forearm length about 39 to 41 mm. It has large teeth; the dental formula is I 1/3, C 1/1, P 2/2, M 3/3 (Shump and Shump 1982a, Adams 2003).
Eastern Red Bat has a distinctive pelage: upper parts are brick-red to rusty-red washed with white, under parts are slightly paler. Only one other bat species in Montana, Hoary Bat, has an interfemoral membrane completely furred on the dorsal surface. The Hoary Bat is much larger (2.0-2.5 X in body wieght, 17-20% longer in total length) than the Eastern Red Bat, and its dorsal pelage is mixed grayish and brownish, tinged with while, giving it a frosty or hoary appearance, not uniformly reddish (Shump and Shump 1982a). Definitive Eastern Red Bat calls are also of higher characteristic frequency: 38-50 kHz lasting > 10 milliseconds for Eastern Red versus < 23 kHz lasting up to 20 milliseconds for Hoary.
Western Hemisphere Range
Current distribution for eastern red bats extends in a line from southwestern Texas through northeastern British Columbia, primarily following the eastern edge of the Rocky Mountains. From Montana to Colorado, 20.2% of records for eastern red bats are from the eastern edge of the Rocky Mountains (Solick et al. 2020).
The northernmost range of eastern red bats appears to be the boundary between the Taiga and Northern Forests ecoregions in Canada. Most records for eastern red bats in western North America are from the Great Plains ecoregion, a third of which were captured or collected within 8 km of major rivers and streams (Solick et al. 2020).
Observations in Montana Natural Heritage Program Database
Number of Observations:
(Click on the following maps and charts to see full sized version)
Map Help and Descriptions
(Observations spanning multiple months or years are excluded from time charts)
The Eastern Red Bat is migratory; all verified records in Montana (n = 11) are from the autumn migration period (between 2 August and 5 September); as of 2011 all individuals identified to sex (n = 3) were female.
The Eastern Red Bat migrates through eastern Montana, particularly along wooded and riparian areas. In other parts of its range, it is reported to prefer elm, box elder, wild plum, willow, hawthorn, sumac, and a variety of other woody plants for roosting, and hibernates in woodpecker holes, tree foliage, and under loose bark (Shump and Shump 1982a, Jones et al. 1983, van Zyll de Jong 1985).
Ecological Systems Associated with this Species
- Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
- Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
- Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
- Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
- Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if
point observations were associated with that system.
Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.
If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: mtnhp.org/requests
) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.
- Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
- Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
- Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
- Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
- Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
- Maxell, B.A. 2000. Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
- Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
- Commonly Associated with these Ecological Systems
Forest and Woodland Systems
Recently Disturbed or Modified
Shrubland, Steppe and Savanna Systems
Sparse and Barren Systems
Wetland and Riparian Systems
- Occasionally Associated with these Ecological Systems
Forest and Woodland Systems
Human Land Use
Recently Disturbed or Modified
Wetland and Riparian Systems
No diet information is available from Montana. Elsewhere it is reported that Eastern Red Bat feeds on flying insects in wooded areas, often on moths (Lepidoptera) but also the Orders Homoptera, Coleoptera, Hymenoptera, and Diptera; they also feed on ground-dwelling crickets, flies, bugs, beetles, cicadas and grain moths (Shump and Shump 1982a). East of Montana they are reported to hunt around city street lights or barn flood lights.
No information from Montana. Elsewhere, these bats tend to be solitary, roosting singly or in female-litter groups, usually in foliage or tree cavities (1 to 6 m above ground but also at ground level) near habitat edges or water. Tends not to associate with other bat species, including foliage-roosting species, except during foraging or drinking. They often begin to forage within two hours after sunset, with some feeding throughout the night. During winter they arouse from hibernation on warm days to feed. Several species of mammals and raptorial birds are documented predators of Eastern Red Bat. In the east, Blue Jays are significant predators of young (Shump and Shump 1982a, Adams 2003).
Little information available from Montana; two females caught on 9 and 11 August were nulliparous. Mating probably occurs during autumn migration and within the winter range, during August and September, but implantation delayed until spring. Gestation period is 80-90 days, young are born in late June and July, lactating females reported in southern Michigan in early August. Litter size ranges from 1-5, but averages about 3 (Shump and Shump 1982a).
No management measures have been enacted specifically for the protection of Eastern Red Bat in Montana, in part because we have too few data; more surveys are necessary to define habitat needs, dates of occurrence and migration, and population status. In Michigan, numbers captured in paired netting surveys have declined 52-85% over 1-26 years, which corresponds to a 10-fold reduction in numbers tested for rabies during 38 years (Winhold at el. 2008). Population declines, if real, could be related to forest fragmentation, pesticides and environmental pollutants, controlled burning of leaf litter, and collisions with various man-made objects, all of which could expose the species to hazards throughout the year. This species is also vulnerable to collision with wind turbines, and large numbers are killed at wind farms (Hendricks et al 2003, Arnett et al. 2008). Increasing rotor start-up wind speed or changing the pitch angle of blades and lowering the required generator speed for electricity production had the same effect in reducing bat fatalities at an Alberta wind farm by 57-60% (Baerwald et al. 2009), and may be promising mitigation techniques at wind energy facilities.
- Literature Cited AboveLegend: View Online Publication
- Adams, R.A. 2003. Bats of the Rocky Mountain West: natural history, ecology and conservation. University Press of Colorado: Boulder, CO. 289 pp.
- Arnett, E., W.K. Brown, W.P. Erickson, J.K. Fiedler, B.L. Hamilton, T.H. Henry, and A. Jain. 2008. Patterns of bat fatalities at wind energy facilities in North America. Journal of Wildlife Management 72(1): 61-78.
- Baerwald, E.F., J. Edworthy, M. Holder, and R.M. R. Barclay. 2009. A large-scale mitigation experiment to reduce bat fatalities at wind energy facilities. Journal of Wildlife Management 73(7): 1077-1081.
- Hendricks, Paul, C. Currier, S. Lenard, and J. Johnson. August/September 2003. MTNHP Bat Survey Field Form - Sites in central and southern Montana.
- L. Winhold., A. Kurta, R. Foster. 2008. Long-term change in an assemblage of North American bats: are eastern red bats declining. Acta Chiropterologica 359-366
- Shump, K.A., Jr. and A.U. Shump. 1982. (Lasiurus borealis). American Society of Mammalogists, Lawrence, KS. Mammalian Species No. 183:1-6.
- Solick, D.I., R.M.R. Barclay, L. Bishop-Boros, Q.R. Hays, and C.L. Lausen. 2020. Distributions of eastern and western red bats in western North America. Western North American Naturalist 80(1):90-97.
- Van Zyll de Jong, C.G. 1985. Handbook of Canadian mammals. Volume 2. Bats. National Museums of Canada, Ottawa. 212 pp.
- Additional ReferencesLegend: View Online Publication
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- Abernethy, I.M., M.D. Andersen, and D.A. Keinath. 2012. Bats of southern Wyoming: distribution and migration year 2 report. Prepared for the USDI Bureau of Land Management by the Wyoming Natural Diversity Database, University of Wyoming, Laramie, Wyoming.
- Agnarsson, I., C.M. Zambrana-Torrelio, N.P. Flores-Saldana, L.J. May-Collado. 2011. A time-calibrated species-level phylogeny of bats (Chiroptera, Mammalia). PLoS Currents Tree of Life. Edition1. RRN1212.
- Amelon, S.K., F.R. Thompson, and J.J. Millspaugh. 2014. Resource utilization by foraging eastern red bats (Lasiurus borealis) in the Ozark Region of Missouri. Journal of Wildlife Management 78(3): 483-493.
- Amichai, E., G. Blumrosen, and Y. Yovel. 2015. Calling louder and longer: how bats use biosonar under severe acoustic interference from other bats. Proceedings of the Royal Society B: Biological Sciences, The Royal Society 282(1821): 20152064.
- Arnett, E.B. 2007. Presence, relative abundance, and resource selection of bats in managed forest landscapes in western Oregon. Ph.D. dissertation, Oregon State University, Corvallis, Oregon.
- Bachen, D.A., A.L. McEwan, B.O. Burkholder, S.L. Hilty, S.A. Blum, and B.A. Maxell.. 2018. Bats of Montana: Identification and Natural History. Report to Montana Department of Environmental Quality. Montana Natural Heritage Program. Helena, MT. 111pp.
- Bachen, D.A., and B.A. Maxell. 2014. Distribution and status of bird, small mammal, reptile, and amphibian species, South Dakota Field Office-BLM. Report to the Bureau of Land Management, South Dakota Field Office. Montana Natural Heritage Program, Helena, Montana 25 pp. plus appendices.
- Baerwald, E.F. and R.R. Barclay. 2009. Geographic Variation in Activity and Fatality of Migratory Bats at Wind Energy Facilities. Journal Of Mammalogy 90(6): 1341-1349.
- Baker, R.H. and C.J. Phillips. 1965. Mammals from El Nevado de Colima, Mexico. Journal of Mammalogy 46(4): 691-693.
- Barclay, R.M.R. 1984. Observations on the migration, ecology and behaviour of bats at Delta Marsh, Manitoba. Canadian Field-Naturalist. 98(3): 331-336.
- Barclay, R.M.R. 1991. Population Structure of Temperate Zone Insectivorous Bats in Relation to Foraging Behavior and Energy Demand. Journal of Animal Ecology 60(1): 165.
- Bat Conservation International. G4 Harp Trap: Assembly and Advice. 8 p.
- Baxter, D.M., J.M. Psyllakis, M.P. Gillingham, and E.L. O'Brien. 2006. Behavioural response of bats to perceived predation risk while foraging. Ethology 112(10): 977-983.
- Bell, J.F., G.J. Moore, G.H. Raymond, and C.E. Tibbs. 1962. Characteristics of Rabies in Bats in Montana. American Journal of Public Health 52(8): 1293-1301.
- Bell, J.F., W.J. Hadlow and W.L. Jellison. 1957. A survey of chiropteran rabies in western Montana. Public Health Reports. 72(1): 16-18.
- Bender, M.J. and G.D. Hartman. 2015. Bat Activity Increases with Barometric Pressure and Temperature During Autumn in Central Georgia. Southeastern Naturalist 14(2): 231-242.
- Bender, M.J., S.B. Castleberry, D.A. Miller, and T.B. Wigley. 2015. Site occupancy of foraging bats on landscapes of managed pine forest. Forest Ecology & Management 336: 1-10.
- Benedict, R.A. 2004. Reproductive activity and distribution of bats in Nebraska. Western North American Naturalist 64(2): 231-248.
- Berthinussen, A. and J. Altringham. 2012. The effect of a major road on bat activity and diversity. Journal of Applied Ecology 49(1): 82-89.
- Bogan, M.A., and K. Geluso. 1999. Bat roosts and historic structures on National Park Service lands in the Rocky Mountain region. Albuquerque, NM: U.S. Geological Survey, Midcontinent Ecological Science Center, Dept. of Biology, the University of New Mexico. Unpublished report. 25 pp
- British Columbia Ministry of Environment, Lands, and Parks. 1998. Inventory Methods for Bats; Standards for Components of British Columbia's Biodiversity No. 20. Prepared for the Terrestrial Ecosystems Task Force Resources Inventory Committee. Version 2.0. 58 p.
- Butts, T. 1997. Bat surveys Indian Creek Canyon, Elkhorn Mountains, Montana. Continental Divide Wildlife Consulting. Helena, MT 32 pg.
- Butts, T.W. 1993. A preliminary survey of the bats of the Deerlodge National Forest, Montana: 1991. Montana Natural Heritage Program. Helena, MT. 35 pp.
- Butts, T.W. 1993. A survey of the bats of the Deerlodge National Forest Montana: 1992. Montana Natural Heritage Program. Helena, MT. 39 pp.
- Butts, T.W. 1993. A survey of the bats of the Townsend Ranger District, Helena National Forest, Montana. Montana Natural Heritage Program. Helena, Montana. 16 pp.
- Callahan, E.V. and R.D. Drobney. 1997. Selection of summer roosting sites by Indiana bats (Myotis sodalis) in Missouri. Journal of Mammalogy 78(3): 818.
- Carlson, J.C. and P. Hendricks. 2001. A Proposal for: Bat Use of Highway Structures: A Pilot Study. Submitted to the Montana Department of Transportation.
- Carlson, J.C. and S.V. Cooper. 2003. Plant and Animal Resources and Ecological Condition of the Forks Ranch Unit of the Padlock Ranch, Big Horn County, Montana and Sheridan County, Wyoming. Unpublished report to The Nature Conservancy Montana Field Office.
- Carroll, S.K., T.C. Carter, and G.A. Feldhamer. 2002. Placement of nets for bats: Effects on perceived fauna. Southeastern Naturalist 1(2): 193-198.
- Chester, J.M., N.P. Campbell, K. Karsmizki, and D. Wirtz. 1979. Resource inventory and evaluation. Azure Cave, Montana. BLM unpublished report. 55 pp.
- Choate, J. R. and J. M. Anderson. 1997. Bats of Jewel Cave National Monument, South Dakota. Prairie Naturalist 29:39-47.
- Christy, R. E. and S. W. West. 1993. Biology of bats in Douglas-fir forests. U.S.D.A., Forest Serv., Pac. Northw. res. Station, Gen. Tech. Rep. PNW-GTR-308.
- Chung-MacCoubrey, A.L. 2005. Use of pinyon–juniper woodlands by bats in New Mexico. Forest Ecology and Management 204: 209–220.
- Clement, M.J., J.M. O'Keefe, and B. Walters. 2015. A method for estimating abundance of mobile populations using telemetry and counts of unmarked animals. Ecosphere 6(10): 184.
- Clement, M.J., T.J. Rodhouse, P.C. Ormsbee, J.M. Szewczak, and J.D. Nichols. 2014. Accounting for false-positive acoustic detections of bats using occupancy models. Journal of Applied Ecology 51(5): 1460-1467.
- Cockrum, E.L., B. Musgrove, and Y. Peteryszyn. 1996. Bats of Mohave County, Arizona: populations and movements. Occasional Papers of The Museum, Texas Tech University 157: 1-71.
- Coleman, J.L. and R.R. Barclay. 2013. Prey availability and foraging activity of grassland bats in relation to urbanization. Journal of Mammalogy 94(5): 1111-1122.
- Coleman, L.S., W.M. Ford, C.A. Dobony, and E.R. Britzke. 2014. A Comparison of Passive and Active Acoustic Sampling for a Bat Community Impacted by White-Nose Syndrome. Journal of Fish and Wildlife Management 5(2): 217-226.
- Constantine, D. G., February 1959, Ecological Observations of Lasiurine Bats in the North Bay Area of California
- Corbett, J. 2011. Evaluation and management of select natural cave and abandoned mine features of the Lewis & Clark and Helena National Forests, Montana. Bat Conservation International. 18pp plus appendices.
- Cox, M.R., E.V. Willcox, P.D. Keyser, and A.L. Vander Yacht. 2016. Bat response to prescribed fire and overstory thinning in hardwood forest on the Cumberland Plateau, Tennessee. Forest Ecology and Management 359: 221-231.
- Cryan, P. M. 2003. Seasonal distribution of migratory tree bats (Lasiurus and Lasionycteris) in North America. Journal of Mammalogy 84:579-593.
- Cryan, P. M., M. A. Bogan, and J. S. Altenbach. 2000. Effect of elevation on distribution of female bats in the Black Hills, South Dakota. Journal of Mammalogy 81:719-725.
- Cryan, P.M., J.W. Jameson, E.F. Baerwald, C.R. Willis, R.M.R. Barclay, E.A. Snider, and E.G. Crichton. 2012. Evidence of Late-Summer Mating Readiness and Early Sexual Maturation in Migratory Tree-Roosting Bats Found Dead at Wind Turbines. PLoS One 7(10): e47586.
- Cvikel, N., E. Levin, E. Hurme, I. Borissov, A. Boonman, E. Amichai, and Y. Yovel. 2015. On-board recordings reveal no jamming avoidance in wild bats. Proceedings of the Royal Society B: Biological Sciences, The Royal Society 282(1798).
- Davis, R. 1966. Homing performance and homing ability in bats. Ecological Monographs 36(3): 201-237.
- Davis, W.B. 1937. Some mammals from western Montana and eastern Idaho. The Murrelet 18(2): 22-27.
- Department of Health and Environmental Sciences Food and Consumer Safety Bureau. 1981. Montana Bats Part I: Control. Vector Control Bulletin Number 2. 6 p.
- Department of Health and Environmental Sciences Food and Consumer Safety Bureau. 1981. Montana Bats Part II: Identification and Biology. Vector Control Bulletin Number 2A. 10 p.
- Downes, W.J. 1964. Unusual Roosting Behavior in Red Bats. Journal of Mammalogy 45(1): 143-144.
- Dubois, K. 1999. Region 4 bat surveys: 1998 progress report. Unpublished report, Montana Fish, Wildlife & Parks Region 4 Headquarters, Great Falls, Montana. 20 pp.
- DuBois, K. 2000. Species occurrence and Distribution of Bats in North Central Montana: Range Map Changes Resulting from Two Years of Field Surveys. Intermountain Journal of Sciences 6(4): 376.
- Ducummon, S.L. 1997. The North American bats and mines project: a cooperative approach for integrating bat conservation and mine-land reclamation. Paper presented at the 1997 National Meeting of the American Society for Mining and Reclamation, Austin, Texas.
- Duff, A.A. and T.E. Morrell. 2007. Predictive Occurrence Models for Bat Species in California. Journal of Wildlife Management 71(3): 693-700.
- Dunbar, M.B. and T.E. Tomasi. 2006. Arousal patterns, metabolic rate, and an energy budget of eastern red bats (Lasiurus borealis) in winter. Journal of Mammalogy 87(6): 1096-1102.
- Easterla, D. A. 1973. Ecology of the 18 species of Chiroptera at Big Bend National Park, Texas. Part I and II. Northwest Missouri State University Studies 34:1-165.
- Elmore, L.W., D.A. Miller, and F.J. Vilella. 2005. Foraging Area Size and Habitat Use by Red Bats (Lasiurus borealis) in an Intensively Managed Pine Landscape in Mississippi. American Midland Naturalist 153(2): 405-417.
- Elmore, L.W., D.A. Miller, F.J. Vilella. 2004. Selection of diurnal roosts by red bats (Lasiurus borealis) in an intensively managed pine forest in Mississippi. Forest Ecology & Management 199(1): 11-20.
- Everette, A.L., T.J. O'Shea, L.E. Ellison, L.A. Stone, and J.L. McCance. 2001. Bat use of a high-plains urban wildlife refuge. Wildlife Society Bulletin 29(3): 967-973.
- Feigley, H.P. 1998. An Examination of the Issues and Feasibility of Conducting Surveys of Abandoned Mines for Bats. Montana Natural Heritage Program, Helena, MT. 12 p.
- Feigley, H.P., M. Brown, S. Martinez, and K. Schletz. 1997.Assessment of mines for importance to bat species of concern, southwestern Montana. Report to: U.S. Geological Survey, Biological Resources Division, Midcontinent Ecological Science Center; 4512 McMurry Ave., Fort Collins, CO 80525-3400. 9pp.
- Feldhamer, G.A., T.C. Carter, and J.O. Whitaker Jr. 2009. Prey Consumed by Eight Species of Insectivorous Bats from Southern Illinois. American Midland Naturalist 162(1): 43-51.
- Fenton, M. B., H. G Merriam, and G L. Holroyd. 1983. Bats of Kootenay, Glacier, and Mount Revelstoke national parks in Canada: identification by echolocation calls, distribution, and biology. Canadian Journal of Zoology 61 :2503-2508.
- Fenton, M.B. 1990. The Foraging Behavior and Ecology of Animal-Eating Bats. Canadian Journal of Zoology 68(3): 411-422.
- Fenton, M.B. 2003. Science and the conservation of bats: where to next? Wildlife Society Bulletin 31(1) 6-15.
- Ford, W.M., E.R. Britzke, C.A. Dobony, J.L. Rodrigue, and L.B. Johnson. 2011. Patterns of Acoustical Activity of Bats Prior to and Following White-Nose Syndrome Occurrence. Journal of Fish and Wildlife Management 2(2): 125-134.
- Ford, W.M., M.A. Menzel, J.M. Menzel, and D.J. Welch. 2002. Influence of Summer Temperature on Sex Ratios in Eastern Red Bats (Lasiurus borealis). American Midland Naturalist 147(1): 179-184.
- Foresman, K.R. 2001. The wild mammals of Montana. American Society of Mammalogists, Special Publication Number 12. Lawrence, KS. 278 pp.
- Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
- Gehrt, S.D. and J.E. Chelsvig. 2004. Species-specific patterns of bat activity in an urban landscape. Ecological Applications 14(2): 625-635.
- Geluso, K. 2006. Bats in a human-made forest of central Nebraska. The Prairie Naturalist 38(1): 13-23.
- Geluso, K. and E.W. Valdez. 2019. First records of the eastern red bat (Lasiurus borealis) in Arizona, Utah, and western New Mexico. Occasional Papers of the Museum of Texas Tech University 361:1-12.
- Genoways, H. H., and J. K. Jones, Jr. 1972. Mammals from southwestern North Dakota. Occasional Papers, The Museum, Texas Tech University 6:1-36.
- Genter, D. L. 1986. Wintering bats of the upper Snake River plain: occurrence in lava-tube caves. Great Basin Naturalist 46(2):241-244.
- Genter, D.L. and K.A. Jurist. 1995. Bats of Montana. Guide for Assessing Mines for Bats Workshop, June 14-15, 1995, Helena, MT, hosted by Montana Department of State Lands and the Abandoned Mine Reclamation Bureau. Montana Natural Heritage Program. 11 pp.
- Gilbert, A.T., G.F. McCracken, L.L. Sheeler, L.I. Muller, D. O’Rourke, W.J. Kelch, and J.C. New Jr. 2015. Rabies surveillance among bats in Tennessee, USA, 1996-2010. Journal of Wildlife Diseases 51(4): 821-832.
- Griscom, H.R. and D.A. Keinath. 2011. Inventory and status of bats at Devils Tower National Monument. Report prepared for the USDI National Park Service by the Wyoming Natural Diversity Database. University of Wyoming, Laramie, Wyoming.
- Griscom, H.R., M.D. Anderson, and D.A. Keinath. 2012. Bats of southern Wyoming: distribution and migration, year 1 report. Prepared for the USDI Bureau of Land Management by the Wyoming Natural Diversity Database. University of Wyoming, Laramie, Wyoming.
- Hanauska-Brown, L., B.A. Maxell, A. Petersen, and S. Story. 2014. Diversity Monitoring in Montana 2008 – 2010 Final Report. Montana Fish, Wildlife & Parks. Helena, MT. 78 pp.
- Harmata, A., D. Flath, R. Hazlewood, and S. Milodragovich. 2002. Initial Site Evaluation for Wind Resource Development in Montana: An Index Relative to Potential Impacts on Vertebrate Wildlife. Intermountain Journal of Sciences 8(4): 253-254.
- Harvey, M.J., J.S. Altenbach, and T.L. Best. 2011. Bats of the United States and Canada. Johns Hopkins University Press. Baltimore, Maryland. p. 202
- Hatch, S.K., E.E. Connelly, T.J. Divoll, I.J. Stenhouse, K.A. Williams, and J.D. Brown. 2013. Offshore observations of eastern red bats (Lasiurus borealis) in the mid-atlantic United States using multiple survey methods. PLoS one 8(12): e83803.
- Hein, C.D., S.B. Castleberry, and K.V. Miller. 2009. Site-occupancy of bats in relation to forested corridors. Forest Ecology and Management 257(4): 1200-1207.
- Hendricks, P. 1997. Mine assessments for bat activity, Garnet Resource Area, BLM: 1997. Unpublished report to USDI, Bureau of Land Management. 17pp.
- Hendricks, P. 1998. Bats surveys of Azure Cave and the Little Rocky Mountains: 1997-1998. November 1998.
- Hendricks, P. 1999. Effect of gate installation on continued use by bats of four abandoned mine workings in western Montana. Unpublished report to Montana Department of Environmental Quality. Montana Natural Heritage Program, Helena, Montana. 13 pp.
- Hendricks, P. 1999. Mine Assessments for Bat Activity, Helena National Forest: 1999. Montana Natural Heritage Program. Helena, MT. 12 pp.
- Hendricks, P. 2000. Assessment of abandoned mines for bat use on Bureau of Land Management lands in the Philipsburg, Montana area: 1999. Montana Natural Heritage Program, Helena, MT.13pp.
- Hendricks, P. 2000. Bat survey along the Norris-Madison Junction Road corridor, Yellowstone National Park, Wyoming, 1999. Montana Natural Heritage Program, Helena, MT. 15pp.
- Hendricks, P. 2000. Preliminary bat inventory of caves and abandoned mines on BLM lands, Judith Mountains, Montana. Montana Natural Heritage Program, Helena, Montana. 21 pp.
- Hendricks, P. 2003. Assessment of Selected Abandoned Mines for Use by Bats in the Garnet and Avon Areas: 2002. Montana Natural Heritage Program. Helena, MT. 12 p.
- Hendricks, P. and B.A. Maxell. 2005. Bat Surveys on USFS Northern Region Lands in Montana: 2005. Report to the USDA Forest Service, Northern Region. Montana Natural Heritage Program, Helena, MT. 12 pp. plus appendices.
- Hendricks, P. and D. Kampwerth. 2001. Roost environments for bats using abandoned mines in southwestern Montana : a preliminary assessment. Report to the U.S. Bureau of Land Management. Montana Natural Heritage Program, Helena, Montana. 19 pp.
- Hendricks, P. and D.L. Genter. 1997. Bat surveys of Azure Cave and the Little Rocky Mountains: 1996. Montana Natural Heritage Program. Helena, MT. 25 pp.
- Hendricks, P. and J.C. Carlson. 2001. Bat use of abandoned mines in the Pryor Mountains. Report to the Montana Department of Environmental Quality, Mine Waste Cleanup Bureau. Montana Natural Heritage Program. Helena, Montana. 8 pp.
- Hendricks, P. and L.M. Hendricks. 2010. Water Aquistion During Daylight by Free-Ranging Myotis Bats. Northwestern Naturalist 91(3): 336-338.
- Hendricks, P., B.A. Maxell, S. Lenard, C. Currier and J. Johnson. 2006. Riparian Bat Surveys in Eastern Montana. A report to the USDI Bureau of Land Management, Montana State Office. Montana Natural Heritage Program, Helena, Montana. 13 pp. plus appendices.
- Hendricks, P., C. Currier, and J. Carlson. 2004. Bats of the BLM Billings Field Office in south-central Montana, with emphasis on the Pryor Mountains. Report to Bureau of Land Management Billings Field Office. Montana Natural Heritage Program, Helena, MT. 19 pp. + appendices.
- Hendricks, P., D. Kampwerth and M. Brown. 1999. Assessment of abandoned mines for bat use on Bureau of Land Management lands in southwestern Montana: 1997-1998. Montana Natural Heritage Program, Helena. 29 pp.
- Hendricks, P., K. Jurist, D.L. Genter, and J.D. Reichel. 1995. Bat survey of the Sioux District, Custer National Forest: 1994. Montana Natural Heritage Program. Helena, Montana. 41 pp.
- Hendricks, P., K.A. Jurist, D.L. Genter and J.D. Reichel. 1996. Bats of the Kootenai National Forest, Montana. Unpublished report. Montana Natural Heritage Program, Helena, Montana. 99 pp.
- Hendricks, P., K.A. Jurist, D.L. Genter, and J.D. Reichel. 1995. Bat survey of the Kootenai National Forest, Montana: 1994. MTNHP report.
- Hendricks, P., S. Lenard, C. Currier and J. Johnson. 2005. Bat Use of Highway Bridges in South-Central Montana. FHWA/MT-05-007/8159. Final Report prepared for the Montana Department of Transportation, in cooperation with the U.S. Department of Transportation Federal Highway Administration, prepared by the Montana Natural Heritage Program.
- Hendricks, P., S. Lenard, C. Currier, and B.A. Maxell. 2007. Filling the distribution gaps for small mammals in Montana. Helena, MT.: Montana Natural Heritage Program.
- Hendricks, P., S. Lenard, D.M. Stagliano, and B.A. Maxell. 2013. Baseline nongame wildlife surveys on the Fort Peck Indian Reservation. Report to the Assiniboine and Sioux Tribes of the Fort Peck Indian Reservation. Montana Natural Heritage Program, Helena, MT. 83 p.
- Hendricks, Paul. 2012. Winter Records of Bats in Montana. Northwestern Naturalist. 93:154-162.
- Hickey, M. C. and M.B. Fenton. 1990. Foraging by Red Bats (Lasiurus borealis): Do Intraspecific Chases Mean Territoriality? Canadian Journal of Zoology 68(12): 2477-2482.
- Hickey, M.B.C., L. Acharya, and S. Pennington. 1996. Resource Partitioning by Two Species of Vespertilionid Bats (Lasiurus cinereus and Lasiurus borealis) Feeding Around Street Lights. Journal of Mammalogy 77(2): 325-334.
- Hill J.E. and J.D. Smith: 1984. Bats: a natural history. Univ. Texas Press Austin. 243 pp.
- Hinman, K.E. and T.K. Snow, eds. 2003. Arizona Bat Conservation Strategic Plan. Nongame and Endangered Wildlife Program Technical Report 213. Arizona Game and Fish Department, Phoenix, Arizona.
- Holroyd, S.L., V.J. Craig, and P. Govindarajulu. 2016. Best Management Practices for Bats in British Columbia. Ministry of Environment, Victoria, BC. 301pp.
- Humphrey, S. R. 1975. Nursery roosts and community diversity of Nearctic bats. Journal of Mammalogy 56:321-346.
- Hutchinson, J.T., and M.J. Lacki. 1999. Foraging behavior and habitat use of Red Bats in mixed mesophytic forests of the Cumberland Plateau, Kentucky. In J.W. Stringer and D.L. Loftis (Eds.). 12th Central Hardwood Forest Conference. General Technical Report SRS-24 USDA Forest Service, Asheville, NC. pp.171-177.
- Jameson, J.W. and C.K. Willis. 2012. Bat Mortality at a Wind Power Facility in Central Canada. Northwestern Naturalist 93(3): 194-202.
- Jameson, J.W. and C.K. Willis. 2014. Activity of tree bats at anthropogenic tall structures: implications for mortality of bats at wind turbines. Animal Behaviour 97: 145-152.
- Jean, C., P. Hendricks, M. Jones, S. Cooper, and J. Carlson. 2002. Ecological communities on the Red Rock Lakes National Wildlife Refuge: inventory and review of aspen and wetland systems. Report to Red Rock Lakes National Wildlife Refuge.
- Jiang, T., H. Wu, J. Feng. 2015. Patterns and causes of geographic variation in bat echolocation pulses. Integrative Zoology 10(3): 241-256.
- Johnson, G.D., W.P. Erickson, M.D. Strickland, M.F. Shepherd, D.A. Shepherd, and S.A. Sarappo. 2003. Mortality of Bats at a Large-scale Wind Power Development at Buffalo Ridge, Minnesota. American Midland Naturalist 150(2): 332-342.
- Johnson, J.S., K.S. Watrous, G.J. Giumarro, T.S. Peterson, S.A. Boyden, and M.J. Lacki. 2011. Seasonal and geographic trends in acoustic detection of tree-roosting bats. Acta Chiropterologica 13(1): 157-168.
- Jones, G.S. and H.H. Thomas. July-October 1983, Distribution and Ectoparasites of Little Brown Bats, Myotis lucifugus, on Prince Edward Island
- Jones, J.K., Jr., R.P. Lampe, C.A. Spenrath, and T.H. Kunz. 1973. Notes on the distribution and natural history of bats in southeastern Montana. Occasional papers (Texas Tech University Museum) 15:1-11.
- Keeley, B. W., and M. D. Tuttle. 1999. “Bats in American Bridges.” Resource Publication No. 4. Bat Conservation International. Austin, TX. 41 p.
- Keinath, D. 2004. Bat and Terrestrial Mammal Inventories in the Greater Yellowstone Network: A progress report. Wyoming Natural Diversity Database, Laramie, WY. 17 pp.
- Keinath, D. 2005. Supplementary Mammal Inventory of Bighorn Canyon National Recreation Area. Final Report. Wyoming Natural Diversity Database, Laramie, WY. 21 pp.
- Keinath, D.A. 2001. Bat Habitat Delineation and Survey Suggestions for Bighorn Canyon National Recreation Area. Unpublished report prepared by the Wyoming Natural Diversity Database for the North American Bat Conservation Partnership.
- Keinath, D.A. 2005. A bat Conservation Evaluation for White Grass Ranch, Grand Teton National Park, Wyoming. Unpublished report for Grand Teton National Park by the Wyoming Natural Diversity Data Base, Laramie, WY.
- Keinath, D.A. 2007. Yellowstone's World of Bats: Taking Inventory of Yellowstone's Night Life. Yellowstone Science 15: 3-13.
- Kingston, T., G. Jones, Z. Akbar, and T.H. Kunz. 2003. Alternation of echolocation calls in 5 species of aerial-feeding insectivorous bats from Malaysia. Journal of Mammalogy 84(1): 205-215.
- Klug, B.J. and E.F. Baerwald. 2010. Incidence and Management of Live and Injured Bats at Wind Energy Facilities. Journal of Wildlife Rehabilitation 30(2): 11-16.
- Korstian, J.M., A.M. Hale, and D.A. Williams. 2015. Genetic Diversity, Historic Population Size, and Population Structure in 2 North American Tree Bats. Journal of Mammalogy 96(5): 972-980.
- Kubista, C.E. and A. Bruckner. 2015. Importance of urban trees and buildings as daytime roosts for bats. Biologia 70(11): 1545-1552.
- Kudray, G.M., P. Hendricks, E. Crowe, and S.V. Cooper. 2004. Riparian forests of the Wild and Scenic Missouri River: ecology and management. Prepared for Lewistown Field Office, Bureau of Land Management, Lewistown, Montana. Montana Natural Heritage Program, Helena, MT. 29 pp. plus appendices.
- Kuenzi, A. J., G. T. Downard, and M. L. Morrison. 1999. Bat distribution and hibernacula use in west central Nevada. Great Basin Naturalist 59:213-220.
- Kunz, T. H. (ed). 1988. Ecological and behavioral methods for the study of bats. Smithsonian Inst., Washington D.C., 533 pp.
- Kunz, T.H. and M.B. Fenton. 2003. Bat Ecology. Chicago, Ill: University of Chicago Press. p. 1-745.
- Kunz, T.H. and P.A. Racey. 1998. Bat biology and conservation. International Bat Research Conference 1995. Boston University. Smithsonian Institution Press.
- Kurta, A. 2010. Reproductive timing, distribution, and sex ratios of tree bats in Lower Michigan. Journal of Mammalogy 91(3): 586-592.
- LaMarr, S. and A.J. Kuenzi. 2011. Bat species presence in southwestern Montana. Intermountain Journal of Sciences. 17:1-4. Pp 14-19.
- Lampe, R.P., J.K. Jones Jr., R.S. Hoffmann, and E.C. Birney. 1974. The mammals of Carter County, southeastern Montana. Occa. Pap. Mus. Nat. Hist. Univ. Kan. 25:1-39.
- LaVal, R.K. and M.L. LaVal. 1979. Notes on reproduction, behavior, and abundance of the red bat, Lasiurus borealis. Journal of Mammalogy, 60(1):209-212.
- Lemke, T. 1991. Big Sky UFOs. Montana Outdoors 22(6):2-7.
- Lenard, S., B.A. Maxell, P. Hendricks, and C. Currier. 2007. Bat Surveys on USFS Northern Region 1 Lands in Montana: 2006. Report to the USDA Forest Service, Northern Region. Montana Natural Heritage Program, Helena, Montana. 23 pp. plus appendices.
- Lenard, S., P. Hendricks, and B.A. Maxell. 2009. Bat surveys on USFS Northern Region lands in Montana: 2007. A report to the USDA Forest Service, Northern Region. Helena, MT. Montana Natural Heritage Program. 21 pp plus appendices.
- Lewis, S. E. 1995. “Roost Fidelity of Bats: a Review.” Journal of Mammalogy 76:481-496.
- Lilley, T.M., J.S. Johnson, L. Ruokolainen, E.J. Rogers, C.A. Wilson, S.M. Schell, K.A. Field, and D.M. Reeder. 2016. White-nose syndrome survivors do not exhibit frequent arousals associated with Pseudogymnoascus destructans infection. Frontiers in Zoology 13(1): 1.
- Limpert, D.L., D.L. Birch, M.S. Scott, M. Andre, and E. Gillam. 2007. Tree Selection and Landscape Analysis of Eastern Red Bat Day Roosts. Journal of Wildlife Management, 71(2), 478-486.
- Loeb, S.C. and J.M. O'Keefe. 2006. Habitat use by forest bats in South Carolina in relation to local, stand, and landscape characteristics. Journal of Wildlife Management 70(5): 1210-1218.
- Loeb, S.C. and T.A. Waldrop. 2008. Bat activity in relation to fire and fire surrogate treatments in southern pine stands. Forest Ecology & Management 255(8/9): 3185-3192.
- Madson, M. and G. Hanson. 1992. Bat hibernaculum search in the Pryor Mountains, southcentral Montana (Draft). Montana Natural Heritage Program. 35 pp.
- Madson. M. 1990. Tate-Potter Cave. Unpublished report, with maps. 6 pp.
- Mager, K.J. and T.A. Nelson. 2001. Roost-site Selection by Eastern Red Bats (Lasiurus borealis). American Midland Naturalist 145(1): 120.
- Martinez, S. 1999. Evaluation of selected bat habitat sites along the Mammoth-Norris grand loop road corridor, Yellowstone National Park, Wyoming, 1997-1998. [Unpublished report submitted to the Montana Natural Heritage Program, Helena, MT]. 16pp.
- Mathews, F., N. Roche, T. Aughney, N. Jones, J. Day, J. Baker, and S. Langton. 2015. Barriers and benefits: implications of artificial night-lighting for the distribution of common bats in Britain and Ireland. Philosophical Transactions of the Royal Society B: Biological Sciences 370(1667): 20140124.
- Matthews, W. L., and J. E. Swenson. 1982. The mammals of east-central Montana. Proc. Mont. Acad. Sci. 41:1-13.
- McGee, M., D.A. Keinath and G.P. Beauvais. 2002. Survey for rare vertebrates in the Pinedale Field Office of the USDI Bureau of Land Management (Wyoming). Unpublished report prepared for USDI Bureau of Land Management. Wyoming State Office by the Wyoming Natural Diversity Database. University of Wyoming, Laramie, Wyoming.
- Miller, D.A. 2003. Species Diversity, Reproduction, and Sex Ratios of Bats in Managed Pine Forest Landscapes of Mississippi. Southeastern Naturalist 2(1): 59-72.
- Montana Bat Working Group. 2020. Recommendations to reduce bat fatalities at wind energy facilities in Montana. Presented to the Montana Fish and Wildlife Commission.
- Mormann, B.M., M. Milam, and L. Robbins. 2004. Red bats do it in the dirt. Bats 22(2):6-9.
- Mumford, R. E. And J. B. Cope. 1964. Distribution and Status of the Chrioptera of Indiana. Am. Midl. Nat. 72(2):473-489.
- Nagorsen, D.W. and B. Paterson. 2012. An Update on the Status of Red Bats, Lasiurus blossevillii and Lasiurus borealis, in British Columbia. Northwestern Naturalist 93(3): 235-237.
- Nagorsen, D.W. and R.M. Brigham. 1993. Bats of British Columbia. Volume I. The Mammals of British Columbia. UBC Press, Vancouver. 164 pp.
- Nagorsen, D.W., A.A. Bryant, D. Kerridge, G. Roberts, A. Roberts, and M.J. Sarell. 1993. Winter bat records for British Columbia. Northwestern Naturalist. 74(3): 61-66.
- Natural Resources Conservation Service and Bat Conservation International. 1998. Bats and mines: Evaluating abandoned mines for bats: recommendations for survey and closure. 6 p.
- Neuweiler, G. 1989. Foraging ecology and audition in echolocating bats. Trends in Ecology and Evolution 4(6): 160-166.
- Neuweiler, G. 1990. Auditory adaptations for prey capture in echolocating bats. Physiological Reviews 70(3): 615-641.
- Nowak, R.M. and E.P. Walker. 1994. Walker's bats of the world. Johns Hopkins University Press. Baltimore, Maryland.
- O'Keefe, J.M., S.C. Loeb, J.D. Lanham, and H.S. Hill. 2009. Macrohabitat factors affect day roost selection by eastern red bats and eastern pipistrelles in the southern Appalachian Mountains, USA. Forest Ecology and Management 257(8): 1757-1763.
- Olson, C.R., D.P. Hobson, and M.J. Pybus. 2011. Changes in Population Size of Bats at a Hibernaculum in Alberta, Canada, in Relation to Cave Disturbance and Access Restrictions. Northwestern Naturalist 92(3): 224-230.
- Patriquin, Krista J. 2004. Red bat (Lasiurus borealis) captured in northeastern Alberta. Northwestern Naturalist. 85:28-30.
- Peck, J. and A. Kuenzi. 2003. Relationship of Orientation on Internal Temperature of Artificial Bat Roosts, Southwestern Montana. Intermountain Journal of Sciences 9(1): 19-25.
- Perkins, J. M., J. M. Barss, and J. Peterson. 1990. Winter records of bats in Oregon and Washington. Northwestern Naturalist 71:59-62.
- Perry, R.W. 2013. Potential energy expenditure by litter-roosting bats associated with temperature under leaf litter during winter. Journal of Thermal Biology 38(8): 467-473.
- Perry, R.W. and V.L. McDaniel. 2015. Temperatures below leaf litter during winter prescribed burns: implications for litter-roosting bats. International Journal of Wildland Fire 24(4): 544-549.
- Perry, R.W., R.E. Thill, and D.M. Leslie Jr. 2008. Scale-Dependent Effects of Landscape Structure and Composition on Diurnal Roost Selection by Forest Bats. Journal of Wildlife Management 72(4): 913-925.
- Perry, R.W., R.E. Thill, and S.A. Carter. 2007. Sex-specific roost selection by adult red bats in a diverse forested landscape. Forest Ecology & Management 253(1-3): 48-55.
- Quay, W.B. 1948. Notes on Some Bats from Nebraska and Wyoming. Journal of Mammalogy 29(2): 181-182.
- Quay, W.B. and J.S. Miller. 1955. Occurrence of the Red Bat (Lasiurus borealis) in Caves. Journal of Mammalogy 36(3): 454-455.
- Rabe, M. J., T. E. Morrell, H. Green, J. C. demos, Jr., and C. R. Miller. 1998. Characteristics of ponderosa pine snag roosts used by reproductive bats in northeastern Arizona. Journal of Wildlife Management 62:612-621.
- Reddy, E. and M. Fenton. 2003. Exploiting vulnerable prey: moths and red bats (Lasiurus borealis; Vespertilionidae). Canadian Journal of Zoology 81(9): 1553-1560.
- Reid, F. 2006. Peterson Field Guide to Mammals of North America, 4th Edition. Houghton Mifflin Company: Boston and New York, 608 pp.
- Roemer, D.M. 1994. Results of field surveys for bats on the Kootenai National Forest and the Lolo National Forest of western Montana, 1993. Unpublished report for the Kootenai National Forest. Montana Natural Heritage Program, Helena. 19 pp.
- Rossillon, M. 1995. The McDonald Mine, west of Ravalli: a cultural resource inventory and evaluation. Renewable Technologies, Inc.. Butte. MT. Unpublished report. 24 pp.
- Sarkozi, D.L. and D.M. Brooks. 2003. Eastern red bat (Lasiurus borealis) impaled by a loggerhead shrike (Lanius ludovicianus). Southwestern Naturalist 48(2): 301-303.
- Sasse, D. 1989. Lick Creek Cave - Survey for Bats. White Sulfur Springs, MT: USDA Forest Service, Lewis & Clark National Forest. Report to the district ranger of Kings Hill Ranger District.
- Sasse, D. C. 1991 . Survey of Tate-Potter Cave. Unpublished report, U.S. Forest Service Belt Creek Information Station, Neihart, MT. 1 3 pp.
- Sasse, D.B., S. Weinstein, and D.A. Saugey. 2014. Interspecific Aggression by a Rabid Eastern Red Bat (Lasiurus borealis). Journal of Wildlife Diseases 50(3): 694-695.
- Schmidt, U. and G. Joermann. 1986. The Influence of Acoustical Interferences on Echolocation in Bats. Mammalia 50(3): 379-390.
- Schmidt-French, B., E. Gillam, and M.B. Fenton. 2006. Vocalizations emitted during mother-young interactions by captive eastern red bats Lasiurus borealis (Chiroptera: Vespertilionidae). Acta Chiropterologica 8(2): 477-484.
- Schwab, N.A. 2004. Bat Conservation Strategy and plan for the State of Montana. Intermountain Journal of Sciences 10(1-4): 80.
- Schwab, Nathan. 2003. Mine Assessments for Bat Activity on Lands Managed by the BLM, Missoula Field Office 2003. Report to USDI BLM Missoula FO. Missoula, MT. 10pp.
- Schwab, Nathan. 2004. Mine Assessment for Bat Activity on Lands Managed by the BLM, Missoula Field Office 2004. USDI BLM Missoula FO. Missoula, MT. 16 pp.
- Sherwin, R.E., J.S. Altenbach, and D.L. Waldien. 2009. Managing abandoned mines for bats. Bat Conservation International.
- Silvis, A., S.D. Gehrt, R.A. Williams. 2016. Effects of shelterwood harvest and prescribed fire in upland Appalachian hardwood forests on bat activity. Forest Ecology and Management 360:205-212.
- Starbuck, C.A., S.K. Amelon, and F.R. Thompson. 2015. Relationships between bat occupancy and habitat and landscape structure along a savanna, woodland, forest gradient in the Missouri Ozarks. Wildlife Society Bulletin 39(1): 20-30.
- Swenson, J. E. and J. C. Bent. 1977. The bats of Yellowstone County, southcentral Montana. Proceedings of the Montana Academy of Sciences 37:82-84.
- Swenson, J.E. and G.F. Shanks, Jr. 1979. Noteworthy records of bats from northeastern Montana. Journal of Mammalogy. 60(3): 650-652
- Taylor, D.A.R. and M.D. Tuttle. 2007. Water for wildlife: a handbook for ranchers and range managers. Bat Conservation International. 20 p.
- Thomas, D.W. 1995. Hibernating Bats Are Sensitive to Nontactile Human Disturbance. Journal of Mammalogy 76(3): 940-946.
- Tigner, J. and E.D. Stukel. 2003. Bats of the Black Hills: a description of status and conservation needs. South Dakota Department of Game, Fish, and Parks. Wildlife Division Report 2003-05. 94 p.
- Tigner, Joel. 2005. Active Season Bat Surveys of Select Abandoned Mines in the Thompson River Valley, Sanders County, MT. BATWORKS, Rapid City, SD. 16pp.
- Tigner, Joel. 2006. Bat Hibernacula Surveys of Select Abandoned Mines in the Thompson River Valley, Sanders County, Montana. BATWORKS, Rapid City, SD. 9pp.
- Tigner, Joel. 2007. Bat hibernacula surveys (in) gated mines, Pryor Mountains, Carbon County, Montana - Report to BLM. Batworks 2416 Cameron Drive, Rapid City, SD 57702.
- Timm, R.M. 1989. Migration and Molt Patterns of Red Bats (Lasiurus borealis) (Chiroptera: Vespertilionidae) in Illinois. Bulletin of the Chicago Academy of Sciences 14: 1-7.
- Turner, R. W. 1974. Mammals of the Black Hills of South Dakota and Wyoming. Mus. Nat. Hist. Univ. Kansas Misc. Pub. No. 60. 178 pp.
- Tuttle, M.D. and D.A.R. Taylor. 1998. Bats and mines. Bat Conservation International, Inc. Resource Publication No. 3. 52 p.
- Twente, J. W., Jr. 1955. Some aspects of habitat selection and other behavior of cavern-dwelling bats. Ecology 36(4): 706-732.
- Van Gelder, R.G. 1956. Echo-location failure in migratory bats. Transactions of the Kansas Academy of Science 59(2): 220-222.
- Van Zyll de Jong, C.G. and P. Geraghty. 1985. Handbook of Canadian Mammals: Volume 2: Bats. Canadian Museum of Nature. 212 p.
- Vonhof, M. J., and R.M.R. Barclay. 1996. Roost-site selection and roosting ecology of forest-dwelling bats in southern British Columbia. Canadian Journal of Zoology 74:1797-1805.
- Vonhof, M.J. and A.L. Russell. 2015. Genetic approaches to the conservation of migratory bats: a study of the eastern red bat (Lasiurus borealis). PeerJ 3: e983.
- Walters, B.L., C.M. Ritzi, D.W. Sparks, and J.O. Whitaker Jr. 2007. Foraging behavior of eastern red bats (Lasiurus borealis) at an urban-rural interface. American Midland Naturalist, 157(2), 365-373.
- Weller, T.J. and D.C. Lee. 2007. Mist Net Effort Required to Inventory a Forest Bat Species Assemblage. Journal of Wildlife Management 71(1): 251-257.
- Whitaker Jr., J.O. and T.J. O'Shea. 2004. Prey selection in a temperate zone insectivorous bat community. Journal of Mammalogy 85(3): 460-469.
- Whitaker, J.O., H.K. Dannelly, and D.A. Prentice. 2004. Chitinase in Insectivorous Bats. Journal of Mammalogy 85(1): 15-18.
- Willis, C.K. 2015. Conservation Physiology and Conservation Pathogens: White-Nose Syndrome and Integrative Biology for Host–Pathogen Systems. Integrative Comparative Biology 55(4): 631-641
- Wilson, D. E., F. R. Cole, J. D. Nichols, R. Rudran, and M. S. Foster, (eds.). 1996. Measuring and monitoring biological diversity: standard methods for mammals. Smithsonian Institution, U.S.A. 409 pp.
- Winhold, L., A. Kurta, and R. Foster. 2008. Long-term change in an assemblage of North American bats: are eastern red bats declining?. Acta Chiropterologica 10(2): 359-366.
- Wolfe, M.L. and A. Kozlowski. 2006. Bat inventories at Grant-Kohrs Ranch National Historic Site and Little Bighorn Battlefield National Monument, FInal Report. Rocky Mountains Cooperative Ecosystems Studies Unit. Utah State University. Logan, UT. 26 pp.
- Worthington, D.J. 1991. Abundance and distribution of bats in the Pryor Mountains of south central Montana and north eastern Wyoming. Montana Natural Heritage Program, Helena, MT.
- Worthington, D.J. and H.N. Ross. 1990. Abundance and distribution of bats in the Pryor Mountains of south central Montana. Unpublished report for the Montana Natural Heritage Program. 20 p.
- Yates, M.D. and R.M. Muzika. 2006. Effect of Forest Structure and Fragmentation on Site Occupancy of Bat Species in Missouri Ozark Forests. Journal of Wildlife Management 70(5): 1238-1248.
- Zukal, J., J. Pikula, and H. Bandouchova. 2015. Bats as bioindicators of heavy metal pollution: history and prospect. Mammalian Biology 80(3): 220-227.
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