Search Field Guide
Advanced Search
MT Gov Logo
Montana Field Guide

Montana Field Guides

Boreal Chorus Frog - Pseudacris maculata

Image Copyright and Usage Information
Native Species

Global Rank: G5
State Rank: S5
(see State Rank Reason below)


Agency Status
USFWS:
USFS:
BLM:


 

External Links


Listen to an Audio Sample
Copyright by Canadian Amphibian and Reptile Conservation Network
State Rank Reason (see State Rank above)
Species is apparently secure and not at risk of extirpation or facing significant threats in all or most of its range.
  • Details on Status Ranking and Review
    Boreal Chorus Frog (Pseudacris maculata) Conservation Status Review
    Review Date = 05/03/2018
    Range Extent

    ScoreG - 200,000-2,500,000 km squared (about 80,000-1,000,000 square miles)

    Comment300,680 square Kilometers from Natural Heritage Program range maps

    Long-term Trend

    ScoreE - Relatively Stable (±25% change)

    CommentAlthough degradation and loss of habitat have occurred, its flexibility in both breeding sites and habitat have allowed it to continue to thrive in developed areas and it is unlikely that populations have changed much since European arrival.

    Short-term Trend

    ScoreE - Stable. Population, range, area occupied, and/or number or condition of occurrences unchanged or remaining within ±10% fluctuation

    CommentPopulations appear stable based on regular detections during lentic surveys conducted over the last 15 years.

    Threats

    ScoreH - Unthreatened. Threats if any, when considered in comparison with natural fluctuation and change, are minimal or very localized, not leading to significant loss or degradation of populations or area even over a few decades’ time. (Severity, scope, and/or immediacy of threat considered Insignificant.)

    CommentNo operational threats in the next 15-20 years identified

    Intrinsic Vulnerability

    ScoreC - Not Intrinsically Vulnerable. Species matures quickly, reproduces frequently, and/or has high fecundity such that populations recover quickly (< 5 years or 2 generations) from decreases in abundance; or species has high dispersal capability such that extirpated populations soon become reestablished through natural recolonization (unaided by humans).

    CommentThis species has high fecundity, matures quickly, and disperses readily

    Environmental Specificity

    ScoreD - Broad. Generalist. Broad-scale or diverse (general) habitat(s) or abiotic and/or biotic factors are used or required by the species, with all key requirements common in the generalized range of the species in the area of interest. If the preferred food(s) or breeding/nonbreeding microhabitat(s) become unavailable, the species switches to an alternative with no resulting decline in numbers of individuals or number of breeding attempts.

    CommentAlthough restricted to lentic sites for breeding, species is found across a range of habitats

    Raw Conservation Status Score

    Score 3.5 + 0 (geographic distribution) + 0.5 (environmental specificity) + 0 (short-term trend) + 0.75 (threats) = 4.75

 
General Description
EGGS:
Although individual females are known to lay 137 to 793 (mean = 455, SE = 47, N = 16 at a site in Colorado) eggs at a time, eggs are usually deposited in several clutches a few centimeters in size. Clusters contain 7 to 190 eggs and are usually less than 25.4 mm (1 in) across and attached to submerged vegetation (Pack 1920, Pettus and Angleton 1967). Each ovum is black above, white to cream below and is surrounded by a single layer of jelly (Maxell et al. 2009). Ovum diameters are 0.8 to 1.3 mm (0.3-0.5 in) (Pettus and Angleton 1967, Maxell et al. 2009), but total egg diameters, including the jelly layer may vary from 4.0 to 6.0 mm (0.16-0.24 in) (Maxell et al. 2009).

LARVAE:
Eyes outside the outline of the body when viewed from above. Mottled with brown and gold dorsally and pale gold to clear ventrally (Maxell et al. 2009). The dorsal tail fin is highly arched and dendritically pigmented with gold while the ventral tail fin is a uniform width and transparent (Maxell et al. 2009). Total length of 4.8-52 mm (0.2-2 in) (Pettus and Angleton 1967, Hammerson 1999). Tadpoles are brown/bronze with eyes located on the sides of the head.

JUVENILES AND ADULTS:
The ends of the toes have minute disks or toe pads and there is little webbing between any of the toes (Maxell et al. 2009). A dark line extends from the snout through to the groin. Basic coloration varies; dorsal color is cream, gray, brown, or green. Typically, there are three green, brown, or gray stripes or rows of spots dorsally and one row laterally (Smith 1956, Corn 1980a, Maxell et al. 2009). Cream colored ventrally, possibly with a few small black spots. Snout-vent length (SVL) of 7.0-38 mm (0.23-1.5 in) (Blair 1951, Pettus and Angleton 1967).

VOICE:
Calls from this species are loud and can be heard up to 400 m (approximately 1,312 ft) (Bryce Maxell, personal communication). The calls are a short chirp that is repeated in quick succession over several seconds resulting in a long sequence similar in sound to the slow running of fingers over the teeth of a comb (Werner et al. 2004).

Diagnostic Characteristics
Except for the Sierran Treefrog (Pseudacris sierra), adults of all other frogs and toads in Montana are much larger and have webbing between the toes of their hind feet. Similarly, the eyes of the tadpoles for all the other frogs and toads in Montana do not stick out beyond the body outline when viewed from above, excluding the Sierran Treefrog (Werner et al. 2004). However, the geographic range of the Sierran Treefrog does not overlap with the geographic range of the Boreal Chorus Frog.

Species Range
Montana Range Range Descriptions

Native

Western Hemisphere Range

 


Range Comments
Although there is debate about whether subspecific or specific status should be assigned, most authorities recognize four geographic varieties of Western Chorus Frog (Pseudacris triseriata) which range from Great Slave Lake and Hudson Bay to the Gulf of Mexico and from New Jersey to western Idaho at elevations up to 3,720 m (12,200 ft) (Hedges 1986, Platz and Forester 1988, Platz 1989, Conant and Collins 1998, Hammerson 1999). Only one variety, the Boreal Chorus Frog, Pseudacris maculata, (full species recognition) or Pseudacris triseriata maculata (subspecies recognition), is recognized as occurring in Montana. Individuals in Montana have been documented east of the Continental Divide and the Big Hole Valley.

Maximum elevation: 2,841 m (9,320 ft) just east southeast of Black Butte in southern Madison County (Matt Bell; MTNHP 2008).

Observations in Montana Natural Heritage Program Database
Number of Observations: 6088

(Click on the following maps and charts to see full sized version) Map Help and Descriptions
Relative Density

Recency

 

(Observations spanning multiple months or years are excluded from time charts)



Migration
Individuals may commonly undergo seasonal migrations of 250 m (820 ft), but apparently do not normally disperse more than 700 m (2,297 ft) from their natal sites (Spencer 1964b).

Habitat
Typically found within 100 m (328 ft) of permanent or temporary waters in grasslands, shrublands, or forest parklands (Kramer 1974; Roberts and Lewin 1979). Adults are freeze tolerant and are presumed to overwinter in underground rodent burrows, underneath thick vegetation or debris or in the crevices of rocks and logs (Whitaker 1971, Swanson et al. 1996). Inhabits marshes, ponds, small lakes in all life zones including lower alpine (Baxter and Stone 1980). Boreal Chorus Frogs are regularly found in the water only during the breeding period in spring. In eastern Montana, they breed in temporary ponds and small lakes surrounded by prairie (or occasionally open forest) habitats. When not breeding, individuals are generally found in damp grassy/marshy areas or damp forests near water, but they have been found up to 500 m (1,640 ft) from water (Nussbaum et al. 1983, Hammerson 1982).

Ecological Systems Associated with this Species
  • Details on Creation and Suggested Uses and Limitations
    How Associations Were Made
    We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
    1. Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
    2. Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
    3. Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
    4. Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
    Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.  In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.  However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if point observations were associated with that system.  Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.  The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.  If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.

    Suggested Uses and Limitations
    Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.  These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: mtnhp.org/requests) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.  Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.  Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.  Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).  Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.

    Literature Cited
    • Adams, R.A.  2003.  Bats of the Rocky Mountain West; natural history, ecology, and conservation.  Boulder, CO: University Press of Colorado.  289 p.
    • Dobkin, D. S.  1992.  Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34.  Missoula, MT.
    • Foresman, K.R.  2012.  Mammals of Montana.  Second edition.  Mountain Press Publishing, Missoula, Montana.  429 pp.
    • Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998.  Montana atlas of terrestrial vertebrates.  Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT.  1302 p.
    • Hutto, R.L. and J.S. Young.  1999.  Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32.  72 p.
    • Maxell, B.A.  2000.  Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species.  Report to U.S. Forest Service Region 1.  Missoula, MT: Wildlife Biology Program, University of Montana.  161 p.
    • Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath.  2004.  Amphibians and reptiles of Montana.  Missoula, MT: Mountain Press Publishing Company. 262 p.

Food Habits
Tadpoles feed on algae (Whitaker 1971). Adults and juveniles feed on a variety of arthropods such as ants, spiders, flies, beetles, and aphids (Nussbaum et al. 1983). They have also been known to feed on their own shed skins and vegetation (Moore and Strickland 1954, Whitaker 1971).

Ecology
Most common amphibian noted in north-central Montana. In high mountains, males may not breed until their second year and females in their third year. Survival to adulthood may be only around 1% in mountain populations (Hammerson 1982).

Reproductive Characteristics
Breeding takes place in late March to early June in Wyoming (Hammerson 1982) or in late April to June in a variety of shallow waterbodies (Cope 1879, Roberts and Lewin 1979, Maxell et al. 2009). Breeding sites are in shallow, warm, fishless waters which may or may not have emergent vegetation (Maxell et al. 2009). They announce their presence this time of year by calling frequently at night and sporadically during the day. Noted singing in early April in southwest Idaho and as late as early July (Nussbaum et al. 1983).

Females deposit eggs on emergent vegetation at depths usually less than 20 cm (7.9 in) in ponds that do not have a closed canopy (Livezey 1952, Maxell et al. 2009) Eggs usually hatch in 5 to 14 days and tadpoles metamorphose in two or three months during mid-summer (Livezey 1952, Nussbaum et al. 1983). Metamorphosis observed in late June to early August across eastern Montana (Mosimann and Rabb 1952).

Management
The following was taken from the Status and Conservation section for the Boreal Chorus Frog account in Maxell et al. 2009

With larvae being found in most temporary standing waterbodies and in shallower portions of permanent standing water bodies that lack fish Boreal Chorus Frogs may be the most widely distributed and common amphibian species at low to mid elevations east of the Continental Divide. Risk factors relevant to the viability of populations of this species are likely to include all the general risk factors described above except for timber harvest and harvest and commerce. Individual studies that specifically identify risk factors or other issues relevant to the conservation of Boreal Chorus Frog include the following. (1) Sanders (1970a) studied the sensitivities of one-week old tadpoles of the Boreal Chorus Frog to 16 pesticides and herbicides and found most of them to result in high rates of mortality when exposed for 48 or 96 hours. Powell et al. (1982) found that the insecticide fenthion formulated with either water or diesel oil had not bioaccumulated in adult Boreal Chorus Frogs three days after exposure at commonly applied levels. However, as noted by the authors it may be unlikely that the adults would bioaccumulate the pesticide because individuals would not be likely to have eaten insects that had been exposed (frogs do not normally eat dead prey). The authors warn that tadpoles may be more sensitive to bioaccumulation because they ingest algae that would likely be contaminated. The relationship of the inactive and active ingredients in these pesticides to commonly applied pesticides in Montana is not known, but it is likely that both pesticides and herbicides may represent lethal and/or sublethal threats to Boreal Chorus Frog populations. (2) Hecnar (1995) found that acute and chronic toxic effects of ammonium nitrate were observed in tadpoles of the Boreal Chorus Frog at concentrations that are commonly exceeded in agricultural areas. Acute exposures to ammonium nitrate fertilizers at 20 mg/L for 96 hours resulted in 50 percent mortality and significant weight loss in those individuals that survived. Chronic exposures to 10 mg/L for 100 days resulted in significantly lower survivorship. (3) Corn et al. (1997) found that Boreal Chorus Frogs were commonly breeding at sites where trout were present but noted that tadpoles of the species are often only found in heavily vegetated shallow water where they are not likely to be exposed to fish predation. (4) Corn et al. (1989) found that embryos from a clutch of Boreal Chorus Frog eggs did not suffer significantly higher mortality rates until pH dropped below 5.2, but had an LC50 at pH 4.8, and suffered 100% mortality at pH 4.6. However, at the larval stage, Kiesecker (1996) found that survival rate, growth rate, mass, and time to metamorphosis did not change when pH was at 4.5, 5.5, 6.0, and 7.0.


References
  • Literature Cited AboveLegend:   View Online Publication
    • Baxter, G.T. and M.D. Stone. 1980. Amphibians and reptiles of Wyoming. Wyoming Game & Fish Department, Cheyenne. 137 pp.
    • Blair, A.P. 1951. Note on the herpetology of the Elk Mountains, Colorado. Copeia 1951: 239-240.
    • Conant, R. and J.T. Collins 1998. A field guide to reptiles and amphibians of eastern and central North America. Third edition, expanded. Houghton Mifflin Company. Boston, MA. 616 p.
    • Cope, E. D. 1879. A contribution to the zoology of Montana. American Naturalist 13(7): 432-441.
    • Corn, P.S. 1980a. Comment on the occurrence of Pseudacris clarki in Montana. Bulletin of the Chicago Herpetological Society 15(3): 77-78.
    • Corn, P.S., M.L. Jennings, and E. Muths. 1997. Survey and assessment of amphibian populations in Rocky Mountain National Park. Northwest. Nat. 78:34-55.
    • Corn, P.S., W. Stolzenburg, and B.R. Bury. 1989. Acid precipitation studies in Colorado and Wyoming: interim report of surveys of montane amphibians and water chemistry. U.S. Fish and Wildlife Service Biological Report 89(40.26). Air Pollution and Acid Rain Report Number 26. 56 p.
    • Hammerson, G.A. 1982. Amphibians and reptiles in Colorado. Colorado Division of Wildlife. Denver, CO. 131 pp.
    • Hammerson, G.A. 1999. Amphibians and reptiles in Colorado. University Press of Colorado & Colorado Division of Wildlife. Denver, CO. 484 p.
    • Hecnar, S.J. 1995. Acute and chronic toxicity of ammonium nitrate fertilizer to amphibians from southern Ontario. Environmental Toxicology and Chemistry 14(12): 2131-2137.
    • Hedges, S.B. 1986. An electrophoretic analysis of Holarctic Hylid frog evolution. Systematic Zoology 35(1): 1-21.
    • Kiesecker, J. 1996. pH mediated predator-prey interactions between Ambystoma tigrinum and Pseudacris triseriata. Ecological Applications 6(4): 1325-1331.
    • Kramer, D.C. 1974. Home range of the western chorus frog (Pseudacris triserata triserata). Journal of Herpetology 8: 245-246.
    • Livezey, R.L. 1952. Some observations on Pseudacris nigrita triseriata (Wied) in Texas. American Midland Naturalist 47: 372-381.
    • Maxell, B.A., P. Hendricks, M.T. Gates, and S. Lenard. 2009. Montana amphibian and reptile status assessment, literature review, and conservation plan, June 2009. Montana Natural Heritage Program. Helena, MT. 643 p.
    • Moore, J.E. and E.H. Strickland. 1954. Notes on the food of three species of Alberta amphibians. American Midland Naturalist 52: 221-224.
    • Mosimann, J.E. and G.B. Rabb. 1952. The herpetology of Tiber Reservoir Area, Montana. Copeia(1): 23-27.
    • Nussbaum, R.A., E.D. Brodie, Jr. and R.M. Storm. 1983. Amphibians and reptiles of the Pacific Northwest. University of Idaho Press. Moscow, ID. 332 pp.
    • Pack, H.J. 1920. Eggs of the swamp tree frog. Copeia 1920(77): 7.
    • Pettus, D. and G.M. Angleton. 1967. Comparative reproductive biology of montane and piedmont chorus frogs. Evolution 21: 500-507.
    • Platz, J. E. 1989. Speciation within the chorus frog Pseudacris triseriata: morphometric and mating call analyses of the boreal and western subspecies. Copeia 1989:704-712.
    • Platz, J.E. and D.C. Forester. 1988. Geographic variation in mating call among the four subspecies of the chorus frog: Pseudacris triseriata (Wied). Copeia 1988(4): 1062-1066.
    • Powell, G.V.N., L.R. DeWeese, and T.G. Lamont. 1982. A field evaluation of frogs as a potential source of secondary organophosphorus insecticide poisoning. Canadian Journal of Zoology 60: 2233-2235.
    • Roberts, W. and V. Lewin. 1979. Habitat utilization and population densities of the amphibians of northeastern Alberta. Canadian Field Naturalist 93(2): 144-154.
    • Sanders, H.O. 1970a. Pesticide toxicities to tadpoles of thet western chorus frog (Pseudacris triseriata) and Fowler's toad (Bufo woodhousei fowleri). Copeia 1970: 246-251.
    • Smith, P.W. 1956. The status, correct name, and geographic range of the boreal chorus frog. Proceedings of the Biological Society of Washington 69: 169-176.
    • Spencer, A.W. 1964b. Movement in a population of Pseudacris triseriata. Journal of the Colorado-Wyoming Academy of Science 5(5): 44.
    • Swanson, D.L., B.M. Graves and K.L. Koster. 1996. Freezing tolerance/intolerance and cryoprotectant synthesis in terrestrially overwintering anurans in the Great Plains, USA. Journal of Comparative Physiology B Biochemistry, Systematics and Environment
    • Werner, J.K., B.A. Maxell, P. Hendricks and D.L. Flath. 2004. Amphibians and Reptiles of Montana. Mountain Press Publishing Company: Missoula, MT. 262 pp.
    • Whitaker, J.A. 1971. A study of the western chorus frog (Pseudacris triseriata) in Vigo County, Indiana. Journal of Herpetology 5(3-4): 127-150.
  • Additional ReferencesLegend:   View Online Publication
    Do you know of a citation we're missing?
    • [DCC] Decker Coal Company. 1998. 1997 Consolidated annual progress report. Decker Coal Company West, North and East Pits. Decker, MT.
    • [EI] Econ Incorporated. 1984. Terrestrial wildlife inventory for the Lame Jones and Ismay coal lease tracts. Econ Incorporated. Helena, MT.
    • [OEA] Olson Elliot and Associates Research. 1985. 1983-1984 Wildlife monitoring report for the CX Ranch project. Olson Elliot and Associates Research. Helena, MT.
    • [PRESI] Powder River Eagle Studies Incorporated. 1998a. Big Sky Mine 1997 wildlife monitoring studies. Powder River Eagle Studies Incorporated. Gillete, WY.
    • [PRESI] Powder River Eagle Studies Incorporated. 1998b. Spring Creek Mine 1997 wildlife monitoring studies. Powder River Eagle Studies Incorporated. Gillete, WY.
    • [VTNWI] VTN Wyoming Incorporated. No Date. Second year's analysis of terrestrial wildlife on proposed mine access and railroad routes in southern Montana and northern Wyoming, March 1979 - February 1980. VTN Wyoming Incorporated. Sheridan, WY. 62 p.
    • [WESCO] Western Ecological Services Company. 1983a. Wildlife inventory of the Knowlton known recoverable coal resource area, Montana. Western Ecological Services Company, Novato, CA. 107 p.
    • [WESCO] Western Ecological Services Company. 1983b. Wildlife inventory of the Southwest Circle known recoverable coal resource area, Montana. Western Ecological Services Company, Novato, CA. 131 p.
    • [WESTECH] Western Technology and Engineering Incorporated. 1991. Update On the Wildlife Resources of the Little Rocky Mountains Environmental Study Area. Western Technology and Engineering, Inc., Helena, Mt.
    • [WESTECH] Western Technology and Engineering Incorporated. 1998. Wildlife Monitoring Absaloka Mine Area 1997. Western Technology and Engineering, Inc., Helena, Mt.
    • Atkinson, E.C. and M.L. Atkinson. 2004. Amphibian and reptile survey of the Ashland and Sioux of the Custer National Forest with special emphasis on the Three-Mile Stewardship Area:2002. Marmot's Edge Conservation. 22 p.
    • Bauer, D. 1997. 1997 wildlife study Savage Mine report. Knife River Corporation, Savage Mine. Richland County, MT.
    • Bauer, Delane, 2002, 2002 Four Seasons Wildlife Study. Savage Mine Report, Richland County, Montana.
    • Baxter, G.T. and M.D. Stone. 1985. Amphibians and reptiles of Wyoming. Second edition. Wyoming Game and Fish Department. Cheyenne, WY. 137 p.
    • Beal, M.D. 1951. The occurrence and seasonal activity of vertebrates in the Norris and Gibbon Geyser Basins of Yellowstone National Park. M.S. Thesis. Utah State Agricultural College. Logan, Utah. 61 pp.
    • Benson, K.R. 1978. Herpetology of the Lewis and Clark expedition 1804-1806. Herpetological Review 9(3): 87-91.
    • Bergeron, D. 1978. Terrestrial Wildlife Survey Coal creek Mine Area, Montana. Unpublished report for Coal Creek Mining Co., Ashland, Montana.
    • Bergeron, D.J. 1978a. Terrestrial wildlife survey Divide Mine area, Montana 1977-1978. Western Technology and Engineering, Inc. Helena, MT.
    • Bergeron, D.J. 1978b. Terrestrial wildlife survey P-M Mine area, Montana 1977-1978. Western Technology and Engineering, Inc. Helena, MT.
    • Black, J.H., and A.N. Bragg. 1968. New additions to the herpetofauna of Montana. Herpetologica 24: 247.
    • BLM. 1982b. Moorhead baseline inventory - wildlife. Bureau of Land Management, Miles City District Office. Miles City, MT. 29 pp.
    • Bramblett, R.G., and A.V. Zale. 2002. Montana Prairie Riparian Native Species Report. Montana Cooperative Fishery Research Unit, Montana State University - Bozeman.
    • Brunson, R.B. 1955. Check list of the amphibians and reptiles of Montana. Proceedings of the Montana Academy of Sciences 15: 27-29.
    • Bureau of Indian Affairs, Department of the Interior. 1981. Draft Environmental Impact Statement. Unpublished report for the Crow/Shell Coal Lease, Crow Indian Reservation, Montana.
    • Burton, S.R., D.A. Patla, and C.R. Peterson. 2002. Amphibians of Red Rock Lakes National Wildlife Refuge: occurrence, distribution, relative abundance, and habitat associations. Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 66 p.
    • Butts, T.W. 1997. Mountain Inc. wildlife monitoring Bull Mountains Mine No. 1, 1996. Western Technology and Engineering. Helena, MT.
    • Confluence Consulting Inc. 2013. Montana Department of Transportation Wetland Mitigation Monitoring Reports (various sites). MDT Helena, MT.
    • Cook, F.R. 1964a. Additional records and a correction of the type locality for the boreal chorus frog in northwestern Ontario. Canadian Field Naturalist 78: 186-192.
    • Cooper, S.V., C. Jean, and P. Hendricks. 2001. Biological survey of a prairie landscape in Montana's glaciated plains. Report to the Bureau of Land Management. Montana Natural Heritage Program, Helena, MT. 24 pp. plus appendices.
    • Cope, E.D. 1872. Report on the recent reptiles and fishes of the survey, collected by Campbell Carrington and C.M. Dawes. pp. 467-469 In: F.V. Hayden, Preliminary report of the United States geological survey of Montana and portions of adjacent territories; being a fifth annual report of progress. 538 pp. 42nd Congress, 2nd Session, House Executive Document Number 326. Serial 1520.
    • Cope, E.D. 1875. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. U.S. Natioanl Museum Bulletin 1: 1-104.
    • Cope, E.D. 1889. The Batrachia of North America. Bulletin of the U.S. National Museum 34: 1-525, figs. 1-119, pls. 1-86.
    • Corn, P.S. and E. Muths. 2004. Variable breeding phenology affects the exposure of amphibian embryos to ultraviolet radation. Ecology 83(11): 2958-2963.
    • Corn, P.S., B.R. Hossack, E. Muths, D.A. Palta, C.R. Peterson, and A.L. Gallant. 2005. Status of amphibians on the Continental Divide: aurveys on a transect from Montana to Colorado, USA. Alytes 22(3-4):85-94.
    • Corn, P.S., E. Muths, and W.M. Iko. 2000. A comparison in Colorado of three methods to monitor breeding amphibians. Northwestern Naturalist 81:22-30.
    • Coues, E. and H. Yarrow. 1878. Notes on the herpetology of Dakota and Montana. Bulletin of the U.S. Geological Geographic Survey of the Territories 4: 259-291.
    • Crother, B.I. (ed.) 2008. Scientific and standard English names of amphibians and reptiles of North America north of Mexico. SSAR Herpetological Circular No. 37:1-84.
    • Day, D. 1989. Montco Terrestrial Wildlife Monitoring Report. Unpublished report for Montco, Billings, Montana.
    • Day, D., P.J. Farmer, and C.E. Farmer. 1989. Montco terrestrial wildlife monitoring report December, 1987 - July, 1989. Montco, Billings, MT, and Western Technology and Engineering, Inc. Helena, MT.
    • Dood, A.R. 1980. Terry Badlands nongame survey and inventory final report. Montana Department of Fish, Wildlife, and Parks and Bureau of Land Management, Helena, MT. 70 pp.
    • Durnin, K.B. and G.R. Smith. 2001. Effects of changing water volume on the tadpoles of two anuran species (Pseudacris triseriata and Rana blairi). Journal of Freshwater Ecology 16(3): 411-414.
    • Econ, Inc. 1988. Wildlife monitoring report, 1987 field season, Big Sky Mine. March 1988. In Peabody Mining and Reclamation Plan Big Sky Mine Area B. Vol. 8, cont., Tab 10 - Wildlife Resources. Appendix 10-1, 1987 Annual Wildlife Report.
    • Farmer, P. 1980. Terrestrial wildlife monitoring study, Pearl area, Montana June, 1978 - May, 1980. Western Technology and Engineering, Inc. Helena, MT.
    • Farmer, P. J. 1980. Terrestrial Wildlife Monitoring Study, Pearl Area, Montana, June, 1978 - May, 1980. Tech. Rep. by WESTECH for Shell Oil Co.
    • Feigley, H. P. 1997. Colonial nesting bird survey on the Bureau of Land Management Lewistown District: 1996. Unpublished report, U.S. Bureau of Land Management, Lewistown, Montana.
    • Fjell, Alan K., 1986, Peabody Coal Company Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1985 field season. March 1986.
    • Fjell, Alan K., and Brian R. Mahan, compilers., 1984, Peabody Coal Company Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1983 field season. February 1984.
    • Fjell, Alan K., and Brian R. Mahan., 1983, Peabody Coal Company Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1982 field season. May 1983.
    • Fjell, Alan K., and Brian R. Mahan., 1985, Peabody Coal Company Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1984 field season. February 1985.
    • Fjell, Alan K., and Brian R. Mahan., 1987, Big Sky Mine, Rosebud County, MT. Wildlife monitoring report: 1986 field season. April 1987.
    • Flath, D.L. 2002. Reptile and amphibian surveys in the Madison-Missouri River Corridor, Montana. Annual Progress Report. 14pp.
    • Garber, C. S. 1992. A survey for spotted frogs (Rana pretiosa), wood frogs (Rana sylvatica), and boreal toads (Bufo boreas) in Wyoming. Wyoming Natural Diversity Database, Laramie. 15 pp. + appendix.
    • Gates, M.T. 2005. Amphibian and reptile baseline survey: CX field study area Bighorn County, Montana. Report to Billings and Miles City Field Offices of Bureau of Land Management. Maxim Technologies, Billings, MT. 28pp + Appendices.
    • Hanauska-Brown, L., B.A. Maxell, A. Petersen, and S. Story. 2014. Diversity Monitoring in Montana 2008-2010 Final Report. Montana Fish, Wildlife & Parks. Helena, MT. 78 pp.
    • Hayden, F.V. 1862. On the geology and natural history of the upper Missouri. Transactions of the American Philosophical Society New Series 12(1): 1-218
    • Hendricks, P. 1999a. Amphibian and reptile survey of the Bureau of Land Management Miles City District, Montana. Montana Natural Heritage Program, Helena, MT. 80 p.
    • Hendricks, P. and J.D. Reichel. 1996b. Preliminary amphibian and reptile survey of the Ashland District, Custer National Forest: 1995. Montana Natural Heritage Program, Helena, MT. 79 p.
    • Hendricks, P. and J.D. Reichel. 1998. Amphibian and reptile survey on Montana refuges: 1996. Montana Natural Heritage Program, Helena, MT. 19 p.
    • Hendricks, P. and M. Roedel. 2001. A faunal survey of the Centennial Valley Sandhills, Beaverhead County, Montana. Report to the U.S. Bureau of Land Management and U.S. Fish and Wildlife Service. Montana Natural Heritage Program, Helena, MT. 44 p.
    • Hendricks, P., S. Lenard, D.M. Stagliano, and B.A. Maxell. 2013. Baseline nongame wildlife surveys on the Fort Peck Indian Reservation. Report to the Assiniboine and Sioux Tribes of the Fort Peck Indian Reservation. Montana Natural Heritage Program, Helena, MT. 83 p.
    • Hill, S.R. 1995. Migratory chronology of adult tiger salamanders (Ambystoma tigrinum) and survey of larvae of the tiger salamander in the northern range of Yellowstone National Park. Masters thesis, Montana State University. Bozeman, MT. 72 pp.
    • Hill, S.R. and R.E. Moore. 1994. Herpetological survey in the northern range of Yellowstone National Park. Annual Report, Yellowstone National Park. February 1, 1994. 21 pp.
    • Hill, S.R., Jr. and R.E. Moore. 1994a. Herpetological survey in the northern range of Yellowstone National Park. Investigator's Annual Reports Yellowstone National Park 1993. Yellowstone Center for Resources. pp. 96-97.
    • Hoppe, D.M. and D. Pettus. 1974. Selection factors affcting dorsal color polymorphism in boreal chorus frogs. Journal of the Colorado-Wyoming Academy of Science 12(5): 73.
    • Hoppe, D.M. and D. Pettus. 1984. Developmental features influencing color polymorphism in chorus frogs. Journal of Herpetology 18: 113-120.
    • Hossack, B. and K.J. Yale. 2002. Geographic Distribution. Pseudacris maculata. Herpetological Review 33(1): 63.
    • Hossack, B. and K.J. Yale. 2002. Geographic Distribution. Pseudacris maculata. Herpetological Review 33(1): 63.
    • Hossack, B. and P.S. Corn. 2001. Amphibian survey of Medicine Lake National Wildlife Refuge Complex: 2001. USGS Northern Rocky Mountain Science Center, Aldo Leopold Wilderness Research Institute, Missoula, MT. 13 p.
    • Hossack,B.R., W.R. Gould, D.A. Patla, E. Muths, R. Daley, K. Legg, and P.S. Corn. 2015. Trends in Rocky Mountain amphibians and the role of beaver as a keystone species. Biological Conservation 187:260-269.
    • Humphris, Michael., 1993, Wildlife Monitoring Report. Spring Creek Coal Company 1993 Mining Annual Report. Appendix I. April 11, 1993.
    • Humphris, Michael., 1994, Wildlife Monitoring Report. Spring Creek Coal Company 1994 Mining Annual Report. Appendix I. April 1994.
    • Jacobs, D. 1950. Pseudacris nigrita triseriata on the north shore of Lake Superior. Copeia 1950(2): 154.
    • Jean, C., P. Hendricks, M. Jones, S. Cooper, and J. Carlson. 2002. Ecological communities on the Red Rock Lakes National Wildlife Refuge: inventory and review of aspen and wetland systems. Report to Red Rock Lakes National Wildlife Refuge.
    • Jones, Lawrence L. C., W. P. Leonard and D. H. Olson, eds. 2005. Amphibians of the Pacific Northwest. Seattle Audubon Society: Seattle, WA, 227 pp.
    • Joslin, Gayle, and Heidi B. Youmans. 1999. Effects of recreation on Rocky Mountain wildlife: a review for Montana. [Montana]: Montana Chapter of the Wildlife Society.
    • Koch, E.D. and C.R. Peterson. 1989. A preliminary survey of the distribution of amphibians and reptiles in Yellowstone National Park. pp. 47-49. In: Rare, sensitive and threatened species of the Greater Yellowstone Ecosystem, T.W. Clark, A.H. Harvey, R.D. Dorn, D.C. Genter, and C. Groves (eds.), Northern Rockies Conservation Cooperative , Montana Natural Heritage Program, The Nature Conservancy, and Mountain West Environmental Services. 153 p.
    • Koch, E.D. and C.R. Peterson. 1995. Amphibians and reptiles of Yellowstone and Grand Teton national parks. University of Utah Press, Salt Lake City, UT. 188 p.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Beaverhead Gateway, Dillon, Montana. Proj. No. 130091.011. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Big Spring Creek, Lewistown, Montana. Proj. No. 130091.029. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2001: Fourchette Creek Reservoir Complex, Phillips County, Montana. Proj. No. 130091.023. July 2002. In 2001 Wetland Mitigation Monitoring Reports, Vol. I.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2002: Big Spring Creek, Lewistown, Montana. Proj. No. 130091.029. February 2003. In 2002 Wetland Mitigation Monitoring Reports, Vol. I.
    • Land & Water Consulting, Inc., Missoula, MT., 2002, Montana Dept. of Transportation Wetland Mitigation Monitoring Report, Year 2002: Fourchette Creek Reservoir Complex, Phillips County, Montana. Proj. No. 130091.023. February 2003. In 2002 Wetland Mitigation Monitoring Reports, Vol. I.
    • Lynch, C. 2000. North American Amphibian Monitoring Program's Montana frog-call survey: a report on a pilot program in south-central Montana started April 2000 and completed in June 2000. Zoo Montana Conservation Through Education Program, Billings MT. 39 p.
    • Lynch, C. 2001. North American Amphibian Monitoring Program's Montana frog-call survey: report on year two of a program in south-central Montana started April 2001 and completed in June 2001. Zoo Montana Conservation Through Education Program, Billings MT. 12 p.
    • Lynch, Catherine. 2000. North American Amphibian Monitoring Program's Montana Frog-call Survey, a report on a pilot program in south-central Montana started April, 2000 and completed in June 2000. 36 pp (unnumbered).
    • MacArthur, D.L. and J.W.T. Dandy. 1982. Physiological aspects of overwintering in the boreal chorus frog (Pseudacris triseriata maculata). Comparative Biochemistry and Physiology 72A:137-141.
    • Mackessy, S.P., R. Donoho, J. Hobert, C. Montgomery and K. Waldron. 1996. Pseudacris triseriata maculata (boreal chorus frog). Herpetological Review 27(1): 30.
    • Martin, P.R. 1980a. Terrestrial wildlife habitat inventory in southeastern Montana. Montana Department of Fish, Wildlife and Parks and Bureau of Land Management, Helena MT. 114 p.
    • Martin, P.R., K. Dubois and H.B. Youmans. 1981. Terrestrial wildlife inventory in selected coal areas, Powder River resources area final report. Montana Department of Fish, Wildlife and Parks and Bureau of Land Management, Helena, MT. No. YA-553-CTO- 24. 288 p.
    • Matthews, W.L. 1979. Wibaux-Beach wildlife baseline study - nongame species. Bureau of Land Management, Miles City, MT. 93 p.
    • Matthews, W.L. 1980b. Wildlife of Prairie County Terry study area. Bureau of Land Management, Miles City, MT. 52 p.
    • Matthews, W.L. 1981. Broadus-Pumpkin Creek baseline inventory - wildlife. Bureau of Land Management, Miles City, MT. 83 p.
    • Matthews, W.L. 1982. Bloomfield-North Fork Baseline Inventories - Wildlife. Unpublished report to the Bureau of Land Management, Miles City, Montana. 61 pp.
    • Maxell, B. A. 2000. Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to USFS Region 1, Order Number 43-0343-0-0224. University of Montana, Wildlife Biology Program. Missoula, MT. 161 p.
    • Maxell, B.A. 2016. Flammulated Owl surveys on the Big Timber, Bozeman, Gardiner and Livingston Ranger Districts of the Custer Gallatin National Forest: 2013. Report to Custer Gallatin National Forest. Montana Natural Heritage Program. Helena, MT. 27pp + appendices.
    • Maxell, B.A., J.K. Werner, P. Hendricks, and D.L. Flath. 2003. Herpetology in Montana: a history, status summary, checklists, dichotomous keys, accounts for native, potentially native, and exotic species, and indexed bibliography. Society for Northwestern Vertebrate Biology, Northwest Fauna Number 5. Olympia, WA. 135 p.
    • Maxim Technologies, Inc., 2002, Western Energy Company Rosebud Mine, Colstrip, Montana: 2002 Annual Wildlife Monitoring Report; December 1, 2001 - November 30, 2002. Febr. 24, 2002.
    • McDonald, M. 1982. Terrestrial wildlife inventory, Dominy Project area, July, 1979 - June, 1981. Draft Tech. Rep. for Western Energy Co. by Westech, Helena, MT.
    • McEneaney, T. and J. Jensen. 1974. The reptiles and amphibians of the Charles M. Russell National Wildlife Range - 1974. Charles M. Russell National Wildlife Refuge. Lewistown, MT. 3 p.
    • Muths, E., D.H. Campbell, and P.S. Corn. 2003. Hatching success in salamanders and chorus frogs at two sites in Colordao, USA: effects of acidic deposition and climate. Amphibia-Reptilia 24(1):27-36.
    • OEA Research. 1985. Wildlife Inventory:Monitoring Report for the CX Ranch Project. 1983-1984. Unpublished report for Consolidation Coal Company, Pittsburgh, Pennsylvania.
    • Patla, D.A. 1997a. Changes in a population of spotted frogs in Yellowstone National Park between 1953 and 1995: the effects of habitat modification. Ph.D. dissertation, Idaho State University, Pocatello.
    • Patla, D.A. 1997b. Potential impacts to amphibians and reptiles from the proposed Canyon contractor camp. 10 May 1997 (Supplement, 12 January 1998). Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 16 p.
    • Patla, D.A. 1998a. Amphibians and reptiles in the Old Faithful sewage treatment area. Report to Yellowstone Center for Resources, Yellowstone National Park. 10 September, 1998. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 7 p.
    • Patla, D.A. 1998b. Potential effects of native fish restoration projects on amphibians in Yellowstone National Park Part I. Report to National Park Service, Yellowstone National Park. 20 November 1998. 26 pp.
    • Patla, D.A. 1999a. Amphibians and reptiles along the grand loop road in Yellowstone National Park: Canyon Junction to Fishing Bridge Junction. December 11, 1999. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 24 p.
    • Patla, D.A. 1999b. Amphibians and reptiles of the Madison to Norris road improvement project area, Yellowstone National Park. 11 November, 1999. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 17 p.
    • Patla, D.A. 1999c. Amphibians and reptiles, Tower Junction to Canyon Village, Yellowstone National Park; addendum to previous report. 24 November, 1999. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 10 p.
    • Patla, D.A. 2000. Amphibians in potential native fish restoration areas, Yellowstone National Park Part II. 7 March, 2000. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 22 p.
    • Patla, D.A. and C.R. Peterson. 1996a. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Tower Junction to Canyon Village. 24 February, 1996. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 49 p.
    • Patla, D.A. and C.R. Peterson. 1996b. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Arnica Creek to Little Thumb Creek. 1 August, 1996. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 35 p.
    • Patla, D.A. and C.R. Peterson. 1997. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Mammoth Hot Springs to Norris Junction. 1 February, 1997. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 60 p.
    • Patla, D.A. and C.R. Peterson. 1998. Amphibians of the Greater Yellowstone Ecosystem. Natural Resource Conservation Cooperative News 11(Autumn 1998): 10-11.
    • Patla, D.A. and C.R. Peterson. 1999. Are amphibians declining in Yellowstone National Park? Yellowstone Science 1999 (Winter):2-11.
    • Patla, D.A. and C.R. Peterson. 2001. Status and trends of amphibian populations in the Greater Yellowstone Ecosystem, progress report, February 2001. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 8 p.
    • Peterson, C.R. and J.P. Shive. 2002. Herpetological survey of southcentral Idaho. Idaho Bureau of Land Management Technical Bulletin 02-3:1-97.
    • Peterson, C.R., C.J. Askey, and D.A. Patla. 1993. Amphibians and reptiles along the Grand Loop and Fountain Freight Roads between Madison Junction and Biscuit Basin in Yellowstone National Park. 26 July, 1993. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 45 p.
    • Peterson, C.R., D.A. Patla, and S.R. Sullivan. 1995. Amphibians and reptiles along the Grand Loop Highway in Yellowstone National Park: Madison Junction to Norris Campground. 7 July, 1995. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 64 p.
    • Peterson, C.R., E.D. Koch and P.S. Corn. 1992. Monitoring amphibian populations in Yellowstone and Grand Teton National Parks final report to University of Wyoming. National Park Service Research Center, Laramie, WY. 37 p.
    • Pilliod, D.S. and T.C. Esque. 2023. Amphibians and reptiles. pp. 861-895. In: L.B. McNew, D.K. Dahlgren, and J.L. Beck (eds). Rangeland wildlife ecology and conservation. Cham, Switzerland: Springer. 1023 p.
    • Platz, J.E. and A. Lathrop. 1993. Body size and age assessment among advertising male chorus frogs. Journal of Herpetology 27(1): 109-111.
    • Powder River Eagle Studies, Inc., Gillette, WY., 1995, Spring Creek Mine 1994 Wildlife Monitoring Studies. 4/94 to 4/95. Spring Creek Coal Company 1995 Mining Annual Report. Appendix I. May 1995.
    • Powder River Eagle Studies, Inc., Gillette, WY., 1996, Spring Creek Mine 1995 Wildlife Monitoring Studies. Spring Creek Coal Company 1995-1996 Mining Annual Report. Vol. I, App. I. May 1996.
    • Powder River Eagle Studies, Inc., Gillette, WY., 1997, Spring Creek Mine 1996 Wildlife Monitoring Studies. February 1997.
    • Powder River Eagle Studies, Inc., Gillette, WY., 1999, Spring Creek Mine 1998 Wildlife Monitoring. March 1999.
    • Powder River Eagle Studies, Inc., Gillette, WY., 2000, Spring Creek Mine 1999 Wildlife Monitoring. March 2000.
    • Powder River Eagle Studies, Inc., Gillette, WY., 2000, Spring Creek Mine 2000 Wildlife Monitoring. March 2000.
    • Powder River Eagle Studies, Inc., Gillette, WY., 2002, Spring Creek Mine 2001 Wildlife Monitoring. March 2002
    • Rauscher, R.L. 1998. Amphibian and reptile survey on selected Montana Bureau of Reclamation impoundments. Montana Department of Fish, Wildlife and Parks, Nongame Program. Bozeman, MT. 24 pp.
    • Rauscher, R.L. 2000. Amphibian and reptile survey on selected Montana Bureau of Reclamation impoundments. Unpublished report, Montana Fish, Wildlife & Parks Non-game Program. 12 pp + maps, tables.
    • Reichel, J. and D. Flath. 1995. Identification of Montana's amphibians and reptiles. Montana Outdoors 26(3):15-34.
    • Reichel, J. D. In prep. Amphibian and reptile survey in southwest Montana: 1996. Unpublished report.
    • Reichel, J. D. In prep. Amphibian and reptile survey of USFWS lands in north-central Montana: 1996. Unpublished report.
    • Reichel, J.D. 1995a. Preliminary amphibian and reptile survey of the Lewis & Clark National Forest: 1994. Montana Natural Heritage Program. Helena, MT. 92 p.
    • Reichel, J.D. 1995b. Preliminary amphibian and reptile survey of the Sioux District of the Custer National Forest: 1994. Montana Natural Heritage Program, Helena, MT. 75 p.
    • Reichel, J.D. 1996. Preliminary amphibian and reptile survey of the Helena National Forest: 1995. Montana Natural Heritage Program. Helena, MT. 87 pp.
    • Reichel, J.D. 1997. Amphibian and reptile survey in northcentral Montana on the Lewistown District, BLM. Unpublished report.
    • Reichel, J.D. 1997a. Amphibian, reptile and northern bog lemming survey on the Rocky Mountain Front: 1996. Montana Natural Heritage Program, Helena, MT. 81 p.
    • Roedel, M.D. and P. Hendricks. 1998a. Amphibian and reptile survey on the Bureau of Land Management Lewistown District: 1995-1998. Montana Natural Heritage Program, Helena, MT. 53 p.
    • Roedel, M.D. and P. Hendricks. 1998b. Amphibian and reptile inventory on the Headwaters and Dillon Resource Areas in conjunction with Red Rocks Lakes National Wildlife Refuge: 1996-1998. Montana Natural Heritage Program, Helena, MT. 46 p.
    • Russell, A. P. and A. M. Bauer. 1993. The amphibians and reptiles of Alberta. University of Calgary Press. Calgary, Alberta. 264 p.
    • Scow, K.L. 1980. Terrestrial wildlife survey American Colloid study area Phillips County, Montana. Western Technology and Engineering, Inc., Helena, MT.
    • Sestrich, C. 2004. Hebgen Reservoir amphibian survey. USDA Forest Service Annual Progress Report to PPL Montana. 22pp.
    • Sestrich, Clint. 2006. 2006 Hebgen Reservoir Amphibian Survey, USDA Forest Service Annual Progress Report to PPL Montana. Hebgen Lake Ranger District. Gallatin National Forest. West Yellowstone Montana.
    • Skinner, M.P. 1924. The Yellowstone Nature Book. A.C. McClurg Company, Chicago, IL. 221 p.
    • Spring Creek Coal Company., 1992, Wildlife Monitoring Report. Spring Creek Coal Company 1992 Mining Annual Report. Appendix I.
    • Stebbins, R. C. 2003. A field guide to western reptiles and amphibians. 3rd Edition. Houghton Mifflin Company, Boston and New York. 533 p.
    • Sullivan, S.R. and C.R. Peterson. 1996. Amphibians and reptiles along the highway in Yellowstone National Park: Tower Junction to the Northeast Entrance. 25 February, 1996. Greater Yellowstone Ecosystem Amphibian Survey and Monitoring Program, Herpetology Laboratory, Department of Biological Sciences, Idaho State University, Pocatello, ID. 60 p.
    • Thompson, L.S. 1981. Circle West wildlife monitoring study: Third annual report. Technical report No. 8. Montana Department of Natural Resources and Conservation. Helena, Montana.
    • Thompson, L.S. 1982. Circle West Wildlife Monitoring Study. Fourth annual report. Technical report 10. Montana Department of Natural Resources and Conservation, Helena, Montana.
    • Timken, R. No Date. Amphibians and reptiles of the Beaverhead National Forest. Western Montana College, Dillon, MT. 16 p.
    • Tordoff, W., III. 1969. Gene frequency differences among semi-isolated proximate populations of chorus forgs (Pseudacris). Journal of Herpetology 3: 194.
    • Tordoff, W., III. 1980. Selective predation of gray jays (Perisoreus canadensis) upon boreal chorus frogs (Pseudacris triseriata). Evolution 34(5): 1004-1008.
    • Tordoff, W., III., and D. Pettus. 1977. Temporal stability of phenotypic frequencies in Pseudacris triseriata (Amphibia, Anura, Hylidae). Journal of Herpetology 11(2): 161-168.
    • Turner, F.B. 1951. A checklist of the reptiles and amphibians of Yellowstone National Park with incidental notes. Yellowstone Nature Notes 25(3): 25-29.
    • Turner, F.B. 1952a. Peculiar aggregations of toadlets on Alum Creek. Yellowstone Nature Notes 26(5): 57-58.
    • Turner, F.B. 1955. Reptiles and amphibians of Yellowstone National Park. Yellowstone Interpretive Series No. 5. Yellowstone Library and Museum Association. Yellowstone National Park, WY. 40 p.
    • Turner, F.B. 1957. The ecology and morphology of Rana pretiosa pretiosa in Yellowstone Park, Wyoming. Ph.D. dissertation, University of California. Berkeley, CA. 252 pp.
    • Turner, F.B. 1960. Population structure and dynamics of the western spotted frog, Rana pretiosa pretiosa Baird & Girard, in Yellowstone Park, Wyoming. Ecol. Monogr. 30(3): 251-278.
    • Van Kirk, R., L. Benjamin, and D. Patla. 2000. Riparian area assessment and amphibian status in the watersheds of the Greater Yellowstone Ecosystem. Greater Yellowstone Coalition, Bozeman, MT. 102 p.
    • Vitt, L.J., J.P. Caldwell, and D.B. Shepard. 2005. Inventory of amphibians and reptiles in the Billings Field Office Region, Montana. Sam Noble Oklahoma Museum of Natural History and Department of Zoology, University of Oklahoma, Norman, OK. 33 pp.
    • VTN Colorado, Inc. Decker Coal Company., 1975, Draft environmental impact assessment for the proposed North Extension of the West Decker Mine.
    • Waage, B.C. 1998. Western Energy Company Rosebud Mine 1997 annual wildlife monitoring report December 1, 1996 to November 30, 1997 survey period. Western Energy Company, Colstrip, MT.
    • Waage, Bruce C., 1993, Western Energy Company Rosebud Mine, Colstrip, Montana: Annual Wildlife Monitoring Report; 1993 Field Season. April 1993.
    • Waage, Bruce C., 1995, Western Energy Company Rosebud Mine, Colstrip, Montana:1994 Annual Wildlife Monitoring Report; December 1, 1993 - November 30, 1994. February 27, 1995.
    • Waage, Bruce C., 1996, Western Energy Company Rosebud Mine, Colstrip, Montana: 1995 Annual Wildlife Monitoring Report; December 1, 1994 - November 30, 1995. February 28, 1996.
    • Waage, Bruce C., 1998, Western Energy Company Rosebud Mine, Colstrip, Montana: 1997 Annual Wildlife Monitoring Report; December 1, 1996 - November 30, 1997 Survey Period. March 23, 1998.
    • Waage, Bruce C., 1999, Western Energy Company Rosebud Mine, Colstrip, Montana: 1998 Annual Wildlife Monitoring Report; December 1, 1997 - November 30, 1998 Survey Period. February 24, 1999.
    • Waage, Bruce C., 2000, Western Energy Company Rosebud Mine, Colstrip, Montana: 1999 Annual Wildlife Monitoring Report; December 1, 1998 - November 30, 1999. February 2000.
    • Waage, Bruce C., 2001, Western Energy Company Rosebud Mine, Colstrip, Montana: 2000 Annual Wildlife Monitoring Report; December 1, 1999 - November 30, 2000. March 30, 2001.
    • Waage, Bruce C., 2002, Western Energy Company Rosebud Mine, Colstrip, Montana. 2001 Annual Wildlife Monitoring Report; December 1, 2000 - November 30, 2001. Febr. 26, 2002.
    • Western Technology & Engineering, Inc. (WESTECH)., 1991, 1991 Bull Mountains Mine No. 1 Terrestrial Wildlife Monitoring Study. In Meridian Minerals Company Bull Mountains Mine No. 1 Permit Application, Musselshell County, Montana. Vol. 7 of 14: Section 26
    • Western Technology and Engineering, Inc. (WESTECH). 1994. Wildlife Monitoring Absaloka Mine Area Annual Report, 1993. Montana SMP 85005. OSMP Montana 0007c. Mar. 12, 1994.
    • Western Technology and Engineering, Inc. (WESTECH)., 1991, Wildlife Monitoring and Additional Baseline Inventory: Absaloka Mine Area Annual Report, 1991. Montana SMP 85005 R1. OSMP Montana 0007B. Febr. 25, 1991.
    • Western Technology and Engineering, Inc. (WESTECH)., 1996, Wildlife Monitoring Absaloka Mine Area Annual Report, 1995. Montana SMP 85005. OSMP Montana 0007D. Febr. 23, 1996.
    • Western Technology and Engineering, Inc. (WESTECH)., 1997, Wildlife Monitoring Absaloka Mine Area Annual Report, 1996. Montana SMP 85005. OSMP Montana 0007D. Mar. 1997.
    • Western Technology and Engineering, Inc. (WESTECH)., 1999, Wildlife Monitoring Absaloka Mine Area Annual Report, 1998. SMP 85005. OSMP Montana 0007E. April 1999.
    • Western Technology and Engineering, Inc. (WESTECH)., 2000, Wildlife Monitoring Absaloka Mine Area Annual Report, 1999. Montana SMP 85005. OSMP Montana 0007E. February 2000.
    • Western Technology and Engineering, Inc. (WESTECH)., 2001, Wildlife Monitoring Absaloka Mine Area Annual Report, 2000. Montana SMP 85005. OSMP Montana 0007E. February 2001.
    • Wheeler, G.C. and J. Wheeler. 1966. The amphibians and reptiles of North Dakota. University of North Dakota, Grand Forks, ND. 104 pp.
    • Wied, M.P. 1843. Travels in the interior of North America by Maximilian, Prince of Wied. With numerous engravings on wood, and a large map. Translated from the German by H. Evans Lloyd. To accompany the original series of eighty-one elaborately-coloured plates. Size, imperial folio. Ackerman and Co., London, England. 520 p.
    • Wied, M.P. 1865. Verzeichniss der reptilien, welche auf einer reise in nordlichen America beobachtet wurden, von Maximilian, Prinzen zu Wied. Eigengangen bei der Akademie am 1, Juni 1865. Druck von E. Blochmann und Sohn, Dresden, Germany. 141 p.
  • Web Search Engines for Articles on "Boreal Chorus Frog"
  • Additional Sources of Information Related to "Amphibians"
Login Logout

Take us with you! PDF icon

Look for this PDF icon as you search and browse.  Click it to build a custom PDF with species shown on that page or for all the species within that taxonomic level.  To create your own list of species, use the PDF Creator tool at the bottom of the Advanced Search page.

Here's some links if you want to download a whole group.

Mammals
Birds
Reptiles
Amphibians
Fish
Invertebrates
Butterflies / Moths
Dragonflies / Damselflies
Ferns and allies
Conifers
Flowering Plants (Dicots)
Flowering Plants (Monocots)
Bryophytes
Lichens
Ecological Systems
Noxious Weeds
Citation for data on this website:
Boreal Chorus Frog — Pseudacris maculata.  Montana Field Guide.  .  Retrieved on , from