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		Spiny-spore Quillwort - Isoetes echinospora
		
			
				Other Names:  
				
				Isoetes tenella
			
		
		
		
		
	 
	
			
            State Rank Reason (see State Rank above)
            Isoetes echinospora is known from 8 occurrences scattered in western Montana.  At one occurrence, the species has been observed in 1940, 1967, and 1998 indicating persistence.  However, current survey work is needed to document locations, population sizes, and threats.
			
               
			
				- Details on Status Ranking and Review
                
					
					    
                    	
                    	    Range ExtentScoreF - 20,000-200,000 sq km (~8,000-80,000 sq mi) Area of OccupancyScoreD - 6-25 4-km2 grid cells Number of PopulationsScoreB - 6 - 20 Number of Occurrences or Percent Area with Good Viability / Ecological IntegrityScoreB - Very few (1-3) occurrences with excellent or good viability or ecological integrity Environmental SpecificityScoreB - Narrow. Specialist or community with key requirements common ThreatsScoreD - Low CommentNo known threats. 
                    	        
                    	     
 
 
		
	 
	
	 
		General Description
		Plants aquatic. Leaves 6–15 cm long with a blunt tip. Sporangia 3–6 mm long. Megaspores 0.4–0.6 mm wide (
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX).
Species Range
	
		
			Montana Range
		Range Descriptions
			 
			
			
					
						 Native
						Native
					 
				
				
					
			 
			
		 
		
	 
	
    		Range Comments
			In MT in Flathead, Lake, Missoula and Ravalli counties; Circumboreal, south to CA, UT, CO, OH, NJ (Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX).
			
		
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 11
		
		
(Click on the following maps and charts to see full sized version)
		Map Help and Descriptions
		
		
			
				Relative Density
				 
			 
		
			
				Recency
				 
			 
		
		 
		
			
				 
			
				 
			
					 (Observations spanning multiple months or years are excluded from time charts)
				
			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Habitat
		Unconsolidated mud in shallow water of lakes and sloughs; valleys to lower subalpine (
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX).
Stewardship Responsibility
		
		
	
		Threats or Limiting Factors
		STATE THREAT SCORE REASON
Threat impact not assigned because threats are not known (MTNHP Threat Assessment 2021).
		
	
	References
	
		
			- Literature Cited AboveLegend:   View Online Publication View Online Publication Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
 MTNHP Threat Assessment. 2021. State Threat Score Assignment and Assessment of Reported Threats from 2006 to 2021 for State-listed Vascular Plants. Botany Program, Montana Natural Heritage Program, Helena, Montana. MTNHP Threat Assessment. 2021. State Threat Score Assignment and Assessment of Reported Threats from 2006 to 2021 for State-listed Vascular Plants. Botany Program, Montana Natural Heritage Program, Helena, Montana.
 
- Additional ReferencesLegend:   View Online Publication View Online Publication
 Do you know of a citation we're missing? Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p. Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p.
 
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