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		Limestone Maidenhair Spleenwort - Asplenium trichomanes-ramosum
		
			
				Other Names:  
				
				Asplenium viride
			
		
		
		
		
	 
	
			
            State Rank Reason (see State Rank above)
            S3 SOC:  Asplenium trichomanes-ramosum plants are never common, grow in habitat that is limited in Montana, and occur where land management (example: national park, wilderness) provides some protections.
			
               
			
				- Details on Status Ranking and Review
                
					
					    
                    	
                    	    Population Size
	                    	
	                    	    ScoreC - 250 - 1,000 individuals
	                    	
                    	    Range Extent
	                    	
                    			ScoreF - 20,000-200,000 sq km (~8,000-80,000 sq mi)
	                    	
                    	    Area of Occupancy
	                    	
                    			ScoreD - 6-25 4-km2 grid cells
	                    	
                    	    Number of Populations
	                    	
                    			ScoreC - 21 - 80
	                    	
                    	    Number of Occurrences or Percent Area with Good Viability / Ecological Integrity
	                    	
                    			ScoreC - Few (4-12) occurrences with excellent or good viability or ecological integrity
	                    	
                    	    Environmental Specificity
	                    	
                    			ScoreB - Narrow. Specialist or community with key requirements common
	                    	
                	        Long-term Trend
	                    	
                				ScoreU - Unknown
	                    	
                	        Trends
	                    	
                				ScoreU - Unknown
	                    	
                    	    Threats
	                    	
                    			ScoreD - Low
	                    	
                        		CommentNo known threats.  Many locations occur in NPS and Wilderness where threats likely don't exist or are very low.
	                    	
                       	    
                    	        
                    	    
                    	
                    	    Intrinsic Vulnerability
	                    	
                    			ScoreU - Unknown
	                    	
					 
				
				 
			
		 
		
	 
	
	 
		General Description
		Plants: Semi-evergreen perennials from short, creeping, scaled rhizomes (McGregor et al. 1986) with few roots (FNA 1993); scales dark, 1-3 mm in length, lanceolate (McGregor 1986).
Leaves: Leaves frequently branched, ascending (FNA 1993), spreading, clustered (McGregor 1986), 4–15 cm long, with 6-20 pairs of pinnae (Lesica 2012); blades arching, once-pinnate, 1.5-12 cm in length, 0.5-1.3 cm in width; petioles slender and wiry, 0.3-4 cm in length, chestnut-colored proximally, becoming green, smooth or with sparse, glandular brown hairs; rachis green; pinnae nearly opposite to alternate, 2-6 mm in length, obovate, short-stalked or nearly sessile, with toothed margins (McGregor et al. 1986) or distal margins crenate (FNA 1993) 
(P. Lesica's contribution adapted from 
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX)
 
		
	
		Phenology
		June-September (McGregor et al. 1986).
		
	
	
	Species Range
	
		
			Montana Range
		Range Descriptions
			
			
			
					
						
						Native
					 
				
				
					
			 
			
		 
		
	 
	
    		Range Comments
			Circumboreal, extending south in scattered localities (Lesica 2012), including Greenland; AB, BC, NB, NL, NT, NS, ON, PE, QC, YT; AK, CA, CO, ID, ME, MI, MT, NV, NY, OR, SD, UT, VT, WA, WI, WY (FNA 1993). Known in Montana from some northwestern counties and is disjunct in Fergus County (Lesica 2012). 
 (Lesica's contribution adapted from Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX)
			
		
		Observations in Montana Natural Heritage Program Database
		Number of Observations: 34
		
		
(Click on the following maps and charts to see full sized version)
		Map Help and Descriptions
		
		
			
				Relative Density
				
			 
		
			
				Recency
				
			 
		
		 
		
			
				
			
				
			
					
				
			(Observations spanning multiple months or years are excluded from time charts)
		 
		
			
		
		
	
		Habitat
		Shaded basic rocks (FNA 1993), including moist to wet limestone crevices; montane to alpine (Lesica 2012).
 (Lesica's contribution adapted from 
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX)
 
		
			
		
		
	
		Reproductive Characteristics
		Sori:  Sori 4 to 8 per pinna (Lesica 2012), blending into each other with age, median, elliptic (McGregor 1986), narrow, 2-3 mm in length (Douglas et al. 2000); the indusia connected to the upper side of the pinna’s veinlet (McGregor et al. 1986).
(P. Lesica's contribution adapted from 
Lesica et al. 2012. Manual of Montana Vascular Plants. BRIT Press. Fort Worth, TX)
 
		
	
		Stewardship Responsibility
		
		
	
		Threats or Limiting Factors
		STATE THREAT SCORE REASON
Threat impact not assigned because threats are not known (MTNHP Threat Assessment 2021).
		
	
	References
	
		
			- Literature Cited AboveLegend:  
 View Online Publication
Douglas, G.W., D. Meidinger, and J. Pojar, editors. 2000. The Illustrated Flora of British Columbia. Volume 5. Dicotyledons (Salicaceae through Zygophyllaceae) and Pteridophytes. British Columbia Ministry of Environment, Lands and Parks and British Columbia Ministry of Forests, Victoria.
Flora of North America Editorial Committee, eds. 1993. Flora of North America North of Mexico. Volume 2. Pteridophytes and Gymnosperms. Oxford University Press, Inc., NY. xvi + 475 pp.
Lesica, P., M.T. Lavin, and P.F. Stickney. 2012. Manual of Montana Vascular Plants. Fort Worth, TX: BRIT Press. viii + 771 p.
McGregor, R.L. (coordinator), T.M. Barkley, R.E. Brooks, and E.K. Schofield (eds). 1986. Flora of the Great Plains: Great Plains Flora Association. Lawrence, KS: Univ. Press Kansas. 1392 pp.
MTNHP Threat Assessment. 2021. State Threat Score Assignment and Assessment of Reported Threats from 2006 to 2021 for State-listed Vascular Plants. Botany Program, Montana Natural Heritage Program, Helena, Montana.
 
			- Additional ReferencesLegend:  
 View Online Publication
Do you know of a citation we're missing?
Britton, N. L. and A. B. Brown. 1913. An Illustrated Flora of the Northern United States, Canada, and the British Possessions. 2nd Edition in 3 Volumes. New York, NY: Charles Scribner's Sons. B13BRI01PAUS.
Lesica, P., M.T. Lavin, and P.F. Stickney. 2022. Manual of Montana Vascular Plants, Second Edition. Fort Worth, TX: BRIT Press. viii + 779 p.
 
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