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A Jaffueliobryum Moss - Jaffueliobryum wrightii


Global Rank: G4G5
State Rank: SNR

Agency Status
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BLM:
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General Description
Plants: Growing in small, crowded patches or convex cushions of upright shoots, green with yellow tones to deep olive green, frosted with hyaline awns (FNA 2007), and pale brown proximally (Crum and Anderson et al. 1981). Stems few-branched, 5-15 mm tall, catkin-like (FNA 2007).

Leaves: Crowded, closely-appressed and overlapping when dry (FNA 2007), spreading when wet, cupped but not keeled (Crum and Anderson et al. 1981), 0.4-0.9 mm in length (minus the awn), elliptic to egg-shaped with the broad end distal, ending in an obtuse, rounded apex plus awn, the awn 0.2-1.3 mm in length, to twice as long as the rest of the blade (FNA 2007), finely saw-toothed (Crum and Anderson et al. 1981); margins flat, smooth or sometimes faintly saw-toothed above (FNA 2007); costa disappearing into the bottom of the awn (Crum and Anderson et al. 1981).

Leaf Cells: Upper lamina 1 cell-layer thick, seldom with stretches of 2 layers, the cells somewhat long and diamond-shaped; medial cells egg-shaped to erratically diamond-shaped, shorter than the upper cells; lower cells square to short and somewhat oblong; costa in X-section flat on the ventral surface (FNA 2007).

Phenology
Fruit ripens in spring and summer (FNA 2007).

Diagnostic Characteristics
J. wrightii tends to occur in hotter, drier areas than its close relative, J. raui, extending further south and generally occurring at lower elevations. J. raui has a costa in X-section distinctly longitudinally folded rather than flat on the ventral surface as in J. wrightii. It also has lower leaves spreading rather than usually lying flat against the stem (FNA 2007).

Range Comments
In North America the species ranges widely, from south central Montana and western North American Range

Canada: AB; USA: CA, NV, AZ, MT to NM, SD to TX, MN, IA, WI; Mexico (FNA 2007). Known in Montana from Carter, Custer, Fallon, Flathead, Garfield, Roosevelt, Rosebud, and Valley Counties (Elliott and Pipp 2016).


Habitat
Calcareous rock and sandstone, dry shrublands and prairies, dry, open woods. Occurring from lowlands to about 9190 feet in elevation (FNA 2007).

Reproductive Characteristics
Usually autoicous. Perichaetial bracts longer and with mostly longer awns than the vegetative leaves. Seta straight, 0.2-0.4 mm tall, leaving the capsule hidden amongst the bracts. Capsule ochre, becoming russet over time, 0.7-1 mm (FNA 2007), creased when dry (Crum and Anderson et al. 1981); operculum with a short beak (FNA 2007); peristome teeth split into about 4 filiform divisions or perforate almost to the bottom, with numerous papillae (Crum and Anderson et al. 1981). Calyptra bell-shaped to somewhat cone-shaped (Crum and Anderson et al. 1981), shielding about 3/4 of the capsule (FNA 2007).

References
  • Literature Cited AboveLegend:   View Online Publication
    • Crum, H.A. and L.E. Anderson. 1981. Mosses of Eastern North America. 2 volumes. Columbia University Press, New York. 1328 pp.
    • Elliott, J. and A. Pipp. 2016 (forthcoming). Checklist of Montana Mosses. Revised 2016. Prepared by the Montana Natural Heritage Program, Helena, Montana. 90 pp.
    • Flora of North America Editorial Committee, eds. 2014. Flora of North America North of Mexico. Volume 28. Bryophytes: Mosses, Part 2. Oxford University Press, Inc., NY. xxi + 702 pp.
  • Additional ReferencesLegend:   View Online Publication
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    • Malcolm, B., N. Malcolm, J. Shevock, and D. Norris. 2009. California Mosses. Nelson, New Zealand: Micro-Optics Press. 430 pp.
  • Web Search Engines for Articles on "A Jaffueliobryum Moss"
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A Jaffueliobryum Moss — Jaffueliobryum wrightii.  Montana Field Guide.  .  Retrieved on , from