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A Cosinodon Moss - Coscinodon calyptratus
Other Names:  Grimmia calyptrata


Global Rank: G3G5
State Rank: SNR

Agency Status
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General Description
Plants: Acrocarpous or cladocarpous. Plants 7-10 mm tall, olivaceous, and in cushions to loose mats.

Leaves: Ovate to ovate-lanceolate; 1.4-2.4 x 0.4-0.7 mm; and not plicate. Margins plane or one margin recurved at mid-leaf. Awn 0.4-1.4 mm.

Leaf Cells: Basal laminal cells near costa are long rectangular, 15-80 x 7-10 µm, & evenly thin-walled. Basal laminal cells near margin quadrate to long-rectangular, 16-60 x 7-15 µm, with lateral walls thin and end walls thick or thin; medial laminal cells 1-layer; distal laminal cells 1-layer.

Diagnostic Characteristics
Characteristics of the sporophyte separate Grimmia and Coscinodon. Coscinodon have calyptra that cover at least half the capsule and are large, bell-shaped, and plicate. Awns are often longer as well. Grimmia have calyptra that are smaller, not plicate, and not bell-shaped.

Range Comments
Confined to western North America, reaching its northern limit in the semi-arid interior of British Columbia and extending southward to southern California and Arizona; predominantly at lower elevations but extending occasionally to subalpine sites (Schofield 1992). Canada: AB, BC; USA: AZ, CA, CO, ID, MN, MT, NV, NM, OR, SD, UT, WA, WI, WY (FNA 2007). In Montana: Madison, Mineral, Missoula, Park, Ravalli, Sanders, and Sweet Grass Counties (Elliott 2016).

Habitat
Dry, acidic rocks; sandstone, granitic rock, limestone, and volcanic outcrops. Moderate to high elevations. In western North America it occurs in the mountains of the dry, interior region. It does not occur on the west coast mountains and its eastern boundary is the Rocky Mountain front with “isolated” populations occurring in the Black Hills (SD), WI, and MN.

Reproductive Characteristics
Seta is 1.4-2.4 mm. Capsule is exserted, cylindric, but often constricted at rim; peristome present & solid. Autoicous (FNA 2007).

References
  • Literature Cited AboveLegend:   View Online Publication
    • Elliott, J. and A. Pipp. 2016 (forthcoming). Checklist of Montana Mosses. Revised 2016. Prepared by the Montana Natural Heritage Program, Helena, Montana. 90 pp.
    • Flora of North America Editorial Committee, eds. 2007. Flora of North America North of Mexico. Volume 27. Bryophytes: Mosses, Part 1. Oxford University Press, Inc., NY. xxi + 713 pp.
    • Schofield, W.B. 1992. Some Common Mosses of British Columbia. Royal British Columbia Museum. Queen's Printer, Canada. 394 pp.
  • Additional ReferencesLegend:   View Online Publication
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    • Flowers, S. 1973. Mosses: Utah and the West. Brigham Young University, Provo, Utah. 567 p.
    • Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.
    • Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
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A Cosinodon Moss — Coscinodon calyptratus.  Montana Field Guide.  .  Retrieved on , from