Gray Comma - Polygonia progne
[From Ferris and Brown 1981; Scott 1986; Opler and Wright 1999; Glassberg 2001; Guppy and Shepard 2001] Forewing 2.3-2.6 cm. Upperside bright orange-brown with a wide dark border; winter form (from 2nd flight) has a border covering only about 1/4 of the wing, both enclosing a few small yellow spots; underside charcoal gray and heavily striated, outer portion of forewing whitish with 3-4 light chevrons in a dark border, silver mark in center of hindwing is small, slender, L-shaped, tapered at both ends.
One flight, mainly June and July north of southern Canada; two flights, late June to early July then August to early September elsewhere (Scott 1986; Glassberg 2001; Guppy and Shepard 2001).
Determined by a combination of upper hindwing black border wide with small yellow spots, underwing surfaces charcoal gray and heavily striated, silver-white "comma" on the under hindwing tapered at both ends.
From central and northeastern British Columbia and southeastern Yukon Territory east through boreal Canada and the northern US to Maine and the Maritimes, south to the central US east of the Rocky Mountains (Scott 1996; Glassberg 2001; Guppy and Shepard 2001). In Montana, reported from at least 11 counties east of the mountains (Stanford and Opler 1993; FLMNH Lepidopterists' Society database; MNHP). Rare to uncommon (Glassberg 2001).
Observations in Montana Natural Heritage Program Database
Number of Observations:
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Map Help and Descriptions
(Observations spanning multiple months or years are excluded from time charts)
Deciduous woodlands, riparian woodlands, forest openings, aspen parkland (Opler and Wright 1999; Glassberg 2001; Guppy and Shepard 2001). Habitat in Montana not described.
Ecological Systems Associated with this Species
- Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (common or occasional) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
- Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2012, Adams 2003, and Werner et al. 2004);
- Evaluating structural characteristics and distribution of each ecological system relative to the species' range and habitat requirements;
- Examining the observation records for each species in the state-wide point observation database associated with each ecological system;
- Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of "observations versus availability of habitat".
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were listed as associated with an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not listed as associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if
point observations were associated with that system.
Common versus occasional association with an ecological system was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species as represented in scientific literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignment of common versus occasional association.
If you have any questions or comments on species associations with ecological systems, please contact the Montana Natural Heritage Program's Senior Zoologist.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: mtnhp.org/requests
) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species' known geographic range.
- Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
- Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
- Foresman, K.R. 2012. Mammals of Montana. Second edition. Mountain Press Publishing, Missoula, Montana. 429 pp.
- Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
- Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
- Maxell, B.A. 2000. Management of Montana's amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
- Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
- Commonly Associated with these Ecological Systems
Forest and Woodland Systems
Shrubland, Steppe and Savanna Systems
Sparse and Barren Systems
Wetland and Riparian Systems
- Occasionally Associated with these Ecological Systems
Sparse and Barren Systems
Larval food plants primarily species of Ribes, also Betula, Rhododendron, rarely Ulmus. Adults feed on tree sap, rotten fruit, mud, rarely flower nectar (Ferris and Brown 1981; Scott 1984, 1986, 1992; Guppy and Shepard 2001).
Females lay eggs singly mostly on the undersides of host plant leaves. Larvae rest on undersides of leaves or on stems; do not build nests. Develop through five larval instars prior to pupation; diapause (overwinter) as adults (Scott 1986). Males perch on small trees at edges of clearings to await passage of females.
- Literature Cited AboveLegend: View Online Publication
- Ferris, C.D. and F.M. Brown (eds). 1981. Butterflies of the Rocky Mountains. Univ. of Oklahoma Press. Norman. 442 pp.
- Glassberg, J. 2001. Butterflies through Binoculars: A Field Guide to the Butterflies of Western North America. Oxford University Press.
- Guppy, C.S. and J.H. Shepard. 2001. Butterflies of British Columbia: including western Alberta, southern Yukon, the Alaska Panhandle, Washington, northern Oregon, northern Idaho, northwestern Montana. UBC Press (Vancouver, BC) and Royal British Columbia Museum (Victoria, BC). 414 pp.
- Opler, P.A. and A.B. Wright. 1999. A field guide to western butterflies. Second edition. Peterson Field Guides. Houghton Mifflin Company, Boston, Massachusetts. 540 pp.
- Scott, J.A. 1984. A review of Polygonia progne (oreas) and P. gracilis (zephyrus) (Nymphalidae), including a new subspecies from the southern Rocky Mountains. Journal of Research on the Lepidoptera. 23(3): 197-210.
- Scott, J.A. 1986. The butterflies of North America: a natural history and field guide. Stanford University Press, Stanford, California.
- Scott, J.A. 1992. Hostplant records for butterflies and skippers (mostly from Colorado) 1959-1992, with new life histories and notes on oviposition, immatures, and ecology. Papilio new series #6. 185 p.
- Stanford, R.E. and P.A. Opler. 1993. Atlas of western USA butterflies: including adjacent parts of Canada and Mexico. Unpubl. Report. Denver and Fort Collins, Colorado 275 pp.
- Additional ReferencesLegend: View Online Publication
Do you know of a citation we're missing?
- Allen, T.J., J.P. Brock, and J. Glassberg. 2005. Caterpillars in the field and garden: a field guide to the butterfly caterpillars of North America. Oxford University Press.
- Brock, J.P. and K. Kaufman. 2003. Kaufman Field Guide to Butterflies of North America. Houghton Mifflin Company, New York, NY 284 pp.
- Layberry, R.A., P.W. Hall, and J.D. LaFontaine. 1998. The Butterflies of Canada. University of Toronto Press. 280 pp. + color plates.
- Additional Sources of Information Related to "Insects"