Common Loon - Gavia immer
The Common Loon is a large and mainly aquatic bird. Males are generally larger than females. Adult body length ranges from 71 to 92 cm (28 to 36 inches) with wingspans to 147 cm (58 inches). Weight varies ranging from 1.6 to 8 kg (3.5 to 17.6 lb.) with an average of about 3 to 4 kg (6.6 to 8.8 lb.) (McIntyre 1988, McIntyre and Barr 1997). The feet are located far back on the body and are large, webbed, and sweep to the side rather than forward under the belly. This trait makes it difficult for Common Loons to walk on land but allows more efficient swimming underwater.
Sexes are indistinguishable based on plumage. The head and neck of breeding adults are black with a green gloss. The back, wings and sides are also black. Scapulars and wing-coverts have large white markings, which is a distinctive field mark. The eye is red. Common Loons have a broad patch of vertical white stripes on the side of the neck and a smaller patch on the upper foreneck. The breast and belly are white and the bill is straight, heavy and black (McIntyre and Barr 1997). In the non-breeding plumage, the head, neck and upper parts are dark gray to dark brown. The cheeks, throat, and underparts are white. The bill is brownish-gray to pale bluish-gray or horn colored. The iris is brown. The tail is dark brown, tipped with white (Bent 1919, Johnsgard 1987, McIntyre 1986, 1988). Juvenile plumage is similar to the adult non-breeding plumage, although the upperparts have paler and more conspicuous feather margins than those of adults, and the throat and sides of the neck are more finely streaked with brown. This plumage is worn until the following summer when the birds molt into more adult-like basic plumage (Palmer 1962, McIntyre 1988).
Common Loons are known for their distinctive calls, three of which are heard on summer breeding lakes. The wail, a long almost mournful cry, the tremolo, a high pitched, rapid, five-beat call, and probably the best known is the yodel which is given only by males during territorial confrontations. Common Loons generally lay 2 subelliptical to ovoid shaped eggs which vary from deep olive to light brown in color, with irregular dark brown or black spots.
The Common Loon is a large loon with a heavy, black bill and an easily recognizable breeding plumage. The large size of the Common Loon distinguishes it from the Pacific Loon (G. pacifica) and the Red-throated Loon (G. stellata), as well as the Arctic Loon (G. arctica), which has never occurred in Montana. Only the Yellow-billed Loon (G. adamsii) is comparable in size. It, however, has a distinctive yellow bill as well as subtle differences in plumage (McIntyre and Barr 1997).
Western Hemisphere Range
Summary of Observations Submitted for Montana
Number of Observations:
(Click on the following maps and charts to see full sized version)
Map Help and Descriptions
(direct evidence "B")
(indirect evidence "b")
No evidence of Breeding
(regular observations "W")
(at least one obs. "w")
(Records associated with a range of dates are excluded from time charts)
In Montana, spring migration begins in early to mid-March. Fall migration starts in late August and may continue through October in Montana. Transient sightings occur throughout the state during spring migration, especially between April and June, and fall migration, between September and November (Montana Bird Distribution Committee 2012). The species is not known to remain on breeding lakes throughout the year, although there are observations of Common Loons remaining in Montana throughout the winter.
In Montana, Common Loons will not generally nest on lakes less than about 13 acres in size or over 5000 feet in elevation (Skaar 1990). Successful nesting requires both nesting sites and nursery areas. Small islands are preferred for nesting, but herbaceous shoreline areas, especially promontories, are also selected. Nursery areas are very often sheltered, shallow coves with abundant small fish and insects (Skaar 1990). Most Montana lakes inhabited by Common Loons are relatively oligotrophic and have not experienced significant siltation or other hydrological changes.
The quantity and quality of nesting habitat limits the Common Loon population of northwest Montana. Skaar (1990) estimated the state's "carrying capacity" at 185 potential nesting territories, based on the size and number of lakes within the species' breeding distribution. He assumed 100 ha of surface area per pair. Kelly (1992) documented a density of 72.2 surface ha of water per adult Common Loon for the Tobacco, Stillwater, Clearwater, and Swan River drainages.
Ecological Systems Associated with this Species
- Details on Creation and Suggested Uses and Limitations
How Associations Were Made
We associated the use and habitat quality (high, medium, or low) of each of the 82 ecological systems mapped in Montana for
vertebrate animal species that regularly breed, overwinter, or migrate through the state by:
- Using personal observations and reviewing literature that summarize the breeding, overwintering, or migratory habitat requirements of each species (Dobkin 1992, Hart et al. 1998, Hutto and Young 1999, Maxell 2000, Foresman 2001, Adams 2003, and Werner et al. 2004);
- Evaluating structural characteristics and distribution of each ecological system relative to the species’ range and habitat requirements;
- Examining the observation records for each species in the state-wide point database associated with each ecological system;
- Calculating the percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system to get a measure of “observations versus availability of habitat”.
Species that breed in Montana were only evaluated for breeding habitat use, species that only overwinter in Montana were only evaluated for overwintering habitat use, and species that only migrate through Montana were only evaluated for migratory habitat use.
In general, species were associated as using an ecological system if structural characteristics of used habitat documented in the literature were present in the ecological system or large numbers of point observations were associated with the ecological system.
However, species were not associated with an ecological system if there was no support in the literature for use of structural characteristics in an ecological system, even if
point observations were associated with that system.
High, medium, and low habitat quality was assigned based on the degree to which the structural characteristics of an ecological system matched the preferred structural habitat characteristics for each species in the literature.
The percentage of observations associated with each ecological system relative to the percent of Montana covered by each ecological system was also used to guide assignments of habitat quality.
If you have any questions or comments on species associations with ecological systems, please contact Bryce Maxell at firstname.lastname@example.org
or (406) 444-3655.
Suggested Uses and Limitations
Species associations with ecological systems should be used to generate potential lists of species that may occupy broader landscapes for the purposes of landscape-level planning.
These potential lists of species should not be used in place of documented occurrences of species (this information can be requested at: http://mtnhp.org/requests/default.asp
) or systematic surveys for species and evaluations of habitat at a local site level by trained biologists.
Users of this information should be aware that the land cover data used to generate species associations is based on imagery from the late 1990s and early 2000s and was only intended to be used at broader landscape scales.
Land cover mapping accuracy is particularly problematic when the systems occur as small patches or where the land cover types have been altered over the past decade.
Thus, particular caution should be used when using the associations in assessments of smaller areas (e.g., evaluations of public land survey sections).
Finally, although a species may be associated with a particular ecological system within its known geographic range, portions of that ecological system may occur outside of the species’ known geographic range.
- Adams, R.A. 2003. Bats of the Rocky Mountain West; natural history, ecology, and conservation. Boulder, CO: University Press of Colorado. 289 p.
- Dobkin, D. S. 1992. Neotropical migrant land birds in the Northern Rockies and Great Plains. USDA Forest Service, Northern Region. Publication No. R1-93-34. Missoula, MT.
- Foresman, K.R. 2001. The wild mammals of Montana. Special Publication No. 12. Lawrence, KS: The American Society of Mammalogists. 278 p.
- Hart, M.M., W.A. Williams, P.C. Thornton, K.P. McLaughlin, C.M. Tobalske, B.A. Maxell, D.P. Hendricks, C.R. Peterson, and R.L. Redmond. 1998. Montana atlas of terrestrial vertebrates. Montana Cooperative Wildlife Research Unit, University of Montana, Missoula, MT. 1302 p.
- Hutto, R.L. and J.S. Young. 1999. Habitat relationships of landbirds in the Northern Region, USDA Forest Service, Rocky Mountain Research Station RMRS-GTR-32. 72 p.
- Maxell, B.A. 2000. Management of Montana’s amphibians: a review of factors that may present a risk to population viability and accounts on the identification, distribution, taxonomy, habitat use, natural history, and the status and conservation of individual species. Report to U.S. Forest Service Region 1. Missoula, MT: Wildlife Biology Program, University of Montana. 161 p.
- Werner, J.K., B.A. Maxell, P. Hendricks, and D. Flath. 2004. Amphibians and reptiles of Montana. Missoula, MT: Mountain Press Publishing Company. 262 p.
No food habit data is available for Montana. Generally, Common Loons dive from the surface and feed mainly on fishes but are opportunistic and will eat any suitable prey they can readily see and capture (McIntyre 1988) including amphibians and various invertebrates (Terres 1980). Their primary food on breeding lakes is yellow perch (Perca flavescens), followed by other shallow, warmwater fish and minnows (Cyprinidae) (Olson and Marshall 1952, Palmer 1962, Barr 1973, McIntyre 1986). Salmonids are taken on lakes that have low populations of other fish species (McIntyre 1988). On the Great Lakes, alewives (Alosa pseudoharengus) appear to be the most common prey item (McIntyre 1988). Crustaceans, especially crayfish (Decapoda), are commonly taken, and plant material is occasionally eaten (Palmer 1962, McIntyre 1988). On lakes without fish, Common Loons have been reported feeding on molluscs, insects, amphipods and amphibians (Munro 1945, Parker 1985). Young birds have a diversified diet consisting primarily of small fish and minnows, aquatic insects and crayfish (McIntyre 1988).
Winter foods are reported to include flounder (Pleuronectoidei), rock cod (Gadus morhua), herring (Clupea spp.), menhaden (Brevoortia patronus), sea trout (Salmo spp.), sculpin (Leptocottus armatus), and crabs (Palmer 1962, McIntyre 1988). A detailed study of winter-feeding patterns and preferences has not been conducted.
If nesting on a small lake, they may use an adjacent lake for supplementary foraging (Johnsgard 1987). In Ontario, Common Loons attempting to raise a chick on a fishless, acidic lake fed the chick benthic algae and possibly benthic invertebrates, but flew to other lakes to feed themselves (Alvo et al. 1988). They feed usually in waters less than 5 m deep.
In 1985, Montana's population was estimated at at least 105 birds (Skaar 1986). No other information is available for the state. Other ecological data, from different areas in the species' range, indicate lakes smaller than 80 ha generally support only one breeding pair. Typically, territory size is larger on large lakes than on small lakes. Wintering birds may defend feeding territories during the day, and gather into rafts at night. The ecology of wintering Common Loons is not well studied. McIntyre (1978) found that loons off the Virginia coast maintained individual feeding territories of four to eight ha during the day and rafted together at night. Activity patterns were significantly correlated with tidal changes. Maintenance behavior was greatest during the mid-period of tidal rise. Feeding activities peaked late in the flood tide and during the first half of the ebb tide.
In Rhode Island, no winter-feeding territories, feeding assemblages, or tide-correlated activity patterns were noted by Daub (1989).
Up to 86 lakes in Montana have had at least one pair of Common Loons present during the breeding season and up to 33 lakes have had Common Loon chicks present. On an annual basis, about 160 to 180 Common Loons can be found on Montana lakes. Between 1999 and 2001, 60% to 80% of these adults formed territorial pairs, but less than half produced chicks (Bissell 2002). Chick production in Montana has ranged between 33 and 51 chicks, with an average of 39.5 chicks produced per year. Average clutch size is 1.87 and the average chicks per successful nest (1982 to 1985) was 1.39 (Skaar 1986).
Most other reproductive data collected on Common Loons comes from studies in other areas of the species' range. These studies indicate the timing of spring arrival is correlated with latitude and dictated primarily by ice-out phenology (McIntyre 1988). Males typically return first, especially in southern breeding areas (McIntyre 1975, 1988; Sutcliffe 1980). However, pairs often arrive together at northern lakes (McIntyre 1988). Territories are established immediately after arrival and may change in size as the breeding season progresses, expanding after the chicks hatch and shrinking for failed pairs (McIntyre 1988).
Courtship begins shortly after territory reoccupation and involves quiet, shared displays including simultaneous swimming, head posturing, and short dives. Vocalizations are not extensive. Copulation sequences are stereotyped, typically last from three to ten minutes, and take place on land (McIntyre 1988). Some copulation sites become nest sites (McIntyre 1975). It is believed that pairs re-mate each spring and that courtship serves primarily to renew the pair bond (McIntyre 1988). Nest-building is conducted by both members of the pair and may immediately follow copulation, sometimes lasting over four days (McIntyre 1975, 1988). Egg laying begins one to 4.5 weeks after spring arrival and eggs are typically laid at two-day intervals (McIntyre 1975). Most clutches contain two eggs, and most one-egg clutches result from loss of the first egg (McIntyre 1975, Titus and VanDruff 1981). Both pair members incubate, beginning with the laying of the first egg, for an average period of 28 to 29 days, ranging from 26 to 31 days (Bent 1919, Olson and Marshall 1952, Palmer 1962, McIntyre 1975). An adult is present at the nest 99 percent of the time, and the eggs hatch within a day of one another (McIntyre 1975).
Chicks leave the nest within 24 hours of hatching and are soon moved to nursery areas (McIntyre 1988). Chicks are fed largely by their parents until eight weeks of age (McIntyre 1988) even though chicks are capable of short dives at the time of nest departure and may capture some fish by the second or third week (McIntyre 1975). Adults aggressively defend chicks underwater and on the surface (McIntyre 1988). Most juveniles are capable of flight at 11 to 12 weeks (Barr 1973, McIntyre 1975), and some leave their small, natal lakes or parental territories shortly afterwards (McIntyre 1975).
Common Loons appear to be faithful to breeding territories. Banded adults have been recaptured on the same breeding territory in subsequent years (McIntyre 1974, Yonge 1981). Yearly reuse of nest sites and nursery areas has been documented (Strong et al. 1987, Jung 1991), but it is not known whether the same individuals were involved. Sonograms of yodel calls suggest that individual males return to the same territory each year (McIntyre 1988, Miller 1989). Little is known about mate fidelity of breeding pairs.
You can access a variety of detailed information on Common Loons and their management in Montana in the Conservation Plan for the Common Loon in Montana (Hammond 2009).
Management of Common Loons and their habitat in Montana should include multiple methods and techniques based on the management intensity level necessary on a particular lake or area. These techniques include, but are not limited to, monitoring, protection from disturbance by people, and protection of nesting and nursery habitat.
The Montana Loon Society monitors Common Loon reproduction each year in Montana. Potential breeding lakes are visited throughout the state and observations are made of adults and chicks. Observing Common Loons on lakes throughout Montana is a productive and valuable method of monitoring, but only reveals limited information; use of the lake and the production of chicks. More intensive monitoring should be implemented in areas where Common Loons are known to occur but have low nest success. This increased monitoring activity would reveal valuable information regarding nesting attempts and the timing and causes of nest failure or chick loss (Dolan 1994). Lakes where no evidence of breeding exists, but Common Loons are present, should be monitored in the spring for any sign of nesting. This added measure would assist in determining the cause of failure at these locations and would also provide a better estimate of total breeding pairs in Montana and rates of success in the population (Dolan 1994). Additional monitoring efforts should be focused on effectiveness of signs in nesting areas and nurseries. Posting floating signs is an effective management method to protect Common Loon nesting sites and nursery areas from disturbance by people. Placing informative signs in key public areas, such as boat ramps, detailing the function and reason for floating signs, and issuing press releases about Common Loons and chick sensitivity during the breeding season, are both important aspects of floating sign success or failure (Dolan 1994).
Management methods that would protect nesting and nursery habitat are varied in scope depending on the location within the state and the particular situation. Primary to the continued use of Montana lakes by Common Loons is the protection of nesting and nursery habitat from destruction resulting from construction, dredging, or filling. Avoiding artificial water level fluctuations is another technique in protecting nest site habitat. Keep water levels consistent and only allow flooding or drawdown practices to occur after the breeding cycle has been completed.
Other ways of protecting Common Loon habitat include implementing no-wake zones to prevent erosion of nesting habitat and working with homeowner associations and other owners, thereby allowing the public to become active in the protection of Common Loons and loon habitat on their lake (Dolan 1994). Common Loons are listed as a sensitive species by the U.S. Fish and Wildlife Service, Region 1 and they are a Species of Management Concern in Region 6 (U.S. Fish and Wildlife Service 1995).
- Literature Cited AboveLegend: View WorldCat Record View Online Publication
- Alvo, R., D. J. T. Hussell, and M. Berril. 1988. The breeding success of common loons (Gavia immer) in relation to alkalinity and other lake characteristics in Ontario. Canadian Journal of Zoology 66:746-752.
- Barr, J. F. 1973. Feeding biology of the common loon (Gavia immer) in oligotrophic lakes of the Canadian shield. University of Guelph, Guelph, Ontario, Canada. Ph.D. dissertation.
- Bent, A.C. 1919. Life histories of North American diving birds. U.S. National Museum Bulletin 107. Washington, D.C.
- Bissell, G. 2002. Second annual common loon report. Montana Fish, Wildlife and Parks. 9 pp. plus appendices.
- Daub, B.C. 1989. Behavior of common loons in winter. Journal of Field Ornithology 60:305-311.
- Dolan, P. M. 1994. The common loon (Gavia immer) in the northern region: biology and management recommendations. Unpublished report. U.S.D.A. Forest Service Region 1. 76 pp.
- Hammond, C. A. 2008. Demographic and landscape analysis for common loons in northwest Montana. M.S. thesis. The University of Montana, Missoula.
- Johnsgard, P. A. 1987. Diving birds of North America. Univ. Nebraska Press, Lincoln. xii + 292 pp.
- Jung, R. E. 1991. Effects of human activities and lake characteristics on the behavior and breeding success of common loons. The Passenger Pigeon 53:207-218.
- Kelley, L.M. 1992. The effects of human disturbance on common loon productivity in northwestern Montana. M.S. thesis. Montana State University, Bozeman. 65 pp.
- McIntyre, J. W. 1974. Territorial affinity of a common loon. Bird-Banding 45:178.
- McIntyre, J. W. 1978. Wintering behavior of common loons. Auk 95:396-403.
- McIntyre, J. W. 1986. Common loon. Pages 679-95 in R. L. Di Silvestro (editor). Audubon Wildlife Report 1986. National Audubon Society, New York, New York.
- McIntyre, J. W. 1988. The common loon: spirit of northern lakes. University of Minnesota Press, Minneapolis. x + 200 pp.
- McIntyre, J. W. and J. F. Barr. 1997. Common loon (Gavia immer). In: A. Poole and F. Gill, eds. The Birds of North America, Number 313. The Academy of Natural Sciences, Philadelphia, PA and The American Ornithologists' Union, Washington, D.C.
- McIntyre, J.M.W. 1975. Biology and behavior of the common loon (Gavia immer) with reference to its adaptability in a man-altered environment. Ph.D. Dissertation. St. Paul, Minnesota: University of Minnesota. 243 p.
- Miller, E. 1989. Population turnover rate and early-season usage of multi-lake territories by common loons in upper Michigan/Wisconsin. North American Loon Fund, Meredith, New Hampshire. Unpublished report.
- Montana Bird Distribution Committee. 2012. P.D. Skaar's Montana bird distribution. 7th Edition. Montana Audubon, Helena, Montana. 208 pp. + foldout map.
- Munro, J. A. 1945. Observations of the loon in the Cariboo Parklands, British Columbia. Auk 62:38-49.
- Olson, S. T., and W. H. Marshall. 1952. The common loon in Minnesota. Minnesota Museum of Natural History, Occasional Paper 5, University of Minnesota Press, Minneapolis, Minnesota.
- Palmer, R. S. 1962. Handbook of North American birds. Volume 1. Loons through flamingos. Yale University Press, New Haven. 567 pp.
- Parker, K. E. 1985. Foraging and reproduction of the common loon (Gavia immer) on acidified lakes in the Adirondack Park, New York. College of Environmental Science and Forestry, State University of New York, Syracuse, New York. M.S. thesis.
- Skaar, D. 1990. Montana common loon management plan. U.S. Forest Service, Region 1, Missoula, Montana. 57 p.
- Skaar, Don. 1986. The Montana loon study: 4th annual report. Unpublished report.
- Strong, P. I. V., J. A. Bissonette, and J. S. Fair. 1987. Reuse of nesting and nursery areas by common loons. Journal of Wildlife Management 51:123-127.
- Sutcliffe, S. A. 1980. Aspects of the nesting ecology of common loons in New Hampshire. University of New Hampshire, Durham, New Hampshire. M.S. thesis.
- Terres, J.K. 1980. The Audubon Society encyclopedia of North American birds. Alfred A. Knopf, New York. 1109 pp.
- Titus, J. R., and L. W. VanDruff. 1981. Response of the common loon to recreational pressure in the Boundary Waters Canoe Area, northeastern Minnesota. Wildlife Monographs 79:1-59.
- U.S. Fish and Wildlife Service, Office of Migratory Bird Management. 1995. Migratory nongame birds of management concern in the United States: the 1995 list. U.S. Government Printing Office: 1996-404-911/44014. 22 p.
- Yonge, K. S. 1981. The breeding cycle and annual production of the common loon (Gavia immer) in the boreal forest region. Thesis. University of Manitoba, Winnipeg, Manitoba. 131 p.
- Additional ReferencesLegend: View WorldCat Record View Online Publication
Do you know of a citation we're missing?
- Alexander, L. L. 1985. Trouble with loons. Living Bird Quarterly. 4:10-13.
- Alvo, R. 1981. Marsh nesting of common loons (GAVIA IMMER). Can. Field Nat. 95:357.
- American Ornithologists' Union. 1983. Check-list of North American birds. Sixth Edition. American Ornithologists' Union, Washington, D.C.
- American Ornithologists' Union. 1998. Check-list of North American birds. 7th edition. American Ornithologists' Union, Washington, D.C. 829 pp.
- Atkinson, E.G. 1991. Distribution of harlequin ducks ( Histrionicus histrionicus ) and common loons (Gavia immer) on the Targee National Forest. Coop. Challenge Cost Share Proj. Targee Natl. For. and Id. Dept. Fish Game.
- Barklow, W. E., and J. A. Chamberlain. 1984. The use of the tremolo call during mobbing by the common loon. Journal of Field Ornithology 55:258-9.
- Barr, J.F. 1986. Population dynamics of the Common Loon (Gavia immer) associated with mercury contaminated waters in northwestern Ontario. Canadian Wildife Service Occ. Paper, no. 56. 25 pp.
- Bull, J. 1974. Birds of New York state. Doubleday/Natural History Press, Garden City, New York. Reprint, 1985 (with Supplement, Federation of New York Bird Clubs, 1976), Cornell Univ. Press, Ithaca, New York.
- Casey, D. 2000. Partners in Flight Draft Bird Conservation Plan Montana. Version 1.0. 287 pp.
- Clark, T. W., H.A. Harvey, R.D. Dorn, D.L. Genter, and C. Groves (eds). 1989. Rare, sensitive, and threatened species of the greater Yellowstone ecosystem. Northern Rockies Conservation Cooperative, Montana Natural Heritage Program, The Nature Conservancy, and Mountain West Environmental Services. 153 p.
- Corkran, C. C. 1988. Status and potential for breeding of the common loon in the Pacific Northwest. Pp. 107-116 in: Papers from the 1987 conference on loon research and management. Paul Strong, (ed). North American Loon Fund, Meredith, NH. 214 pp.
- Desgranges, J.L., and P. Laporte. 1979. Preliminary consideration of the status of loons (Gaviidae) in Quebec. Report to Joint Committee of the James Bay and Northern Quebec Agreement. Canadian Wildlife Service, Ontario, Canada.
- Dulin, G. S. 1987. Pre-fledging feeding behavior and sibling rivalry in common loons. Central Michigan University, Mt. Pleasant, Michigan. M.S. thesis.
- ECON, Inc. (Ecological Consulting Service), Helena, MT., 1977, Colstrip 10 x 20 Area wildlife and wildlife habitat annual monitoring report, 1977. Proj. 164-85-A. December 31, 1977.
- Ehrlich, P., D. Dobkin, and D. Wheye. 1988. The birder’s handbook: a field guide to the natural history of North American birds. Simon and Schuster Inc. New York. 785 pp.
- Fair, J. S. 1990. Water level fluctuations and common loon nest failure. Pages 57-63 in S. Sutcliffe (editor). Proceedings of the second North American conference on common loon research and management. National Udubon Sociey, New York New York.
- Fay, L. D. 1966. Type E botulism in Great Lakes water birds. Pages 139-149 in Trans. Thiry-first N. Amer. Wild. and Nat. Res. Conf.
- Fox, G. A., K. S. Yonge, and S. G. Sealy. 1980. Breeding performance, pollutant burden and eggshell thinning in common loons GAVIA IMMER nesting on a boreal forest lake. Ornis Scand. 11:243-8.
- Frank, R., H. Lumsden, J.F. Barr, and H.E. Braun. 1983. Residues of organochlorine insecticides, industrial chemicals, and mercury in eggs and tissues taken from healthy and emaciated common loons, Ontario, Canada, 1968-1980. Archives of Environmental Contamination and Toxicology 12:641-654.
- Gelatt, T.S. and J.D. Kelley. 1995. Western painted turtles, Chrysemys picta bellii, basking on a nesting common loon, Gavia immer. Canadian Field Naturalist 109(4): 456-45
- Haney, J. C. 1990. Winter habitat of common loons on the continental shelf of the southeastern United States. Wilson Bull. 102:253-263.
- Heglund, P.S., J.R. Jones, L.H. Fredrickson, and M.S. Kaiser. 1994. Use of boreal forested wetlands by Pacific Loons (Gavia immer, Lawrence) and Horned Grebes (Podiceps auritus, L.): relations with limnological characteristics. Hydrobiologia 279/280: 171-183.
- Heimburger, M.D., D. Euler, and J. Barr. 1983. The impact of cottage development on common loon reproductive success in central Ontario. Wilson Bulletin 95:431-439.
- Johnsgard, P. A. 1992. Birds of the Rocky Mountains with particular reference to national parks in the northern Rocky Mountain region. Lincoln: University of Nebraska Press. xi + 504 pp.
- Johnsgard, P.A. 1979. Birds of the Great Plains: breeding species and their distribution. University of Nebraska Press, Lincoln. 539 pp.
- Kaveney, D. E., and C. C. Rimmer. 1989. The breeding status of common loons in Vermont--1989. Vermont Inst. Nat. Sci., Woodstock, Vermont. Unpublished report.
- Kerekes, J., R. Tordon, A. Nieuwburg, and L. Risk. 1994. Fish eating bird abundance in oligotrophic lakes in Kejimkujik National Park, Nova Scotia, Canada. Hydrobiologai 279/280: 57-61.
- Kerlinger, P. 1982. The migration of common loons through eastern New York. Condor 84:97-100.
- Lenard, S., J. Carlson, J. Ellis, C. Jones, and C. Tilly. 2003. P. D. Skaar's Montana Bird Distribution, 6th Edition. Montana Audubon: Helena, MT, 144 pp.
- Locke, L. M., S. M. Kerr, and D. Zoromski. 1982. Lead poisoning in common loons (GAVIA IMMER). Avian Diseases 26:392-6.
- McEneaney, T. 1988. Status of the common loon in Yellowstone National Park. Pp. 117-129 in: Paul Strong, (ed.), Papers from the 1987 conference on loon research and management. North American Loon Fund, Meredith, NH. 214 pp.
- McIntyre, J. W. 1983. Nurseries: A Consideration of Habitat Requirements During the Early Chick-Rearing Period in Common Loons. Journal of Field Ornithology 54(3):247-253.
- McIntyre, J. W., and J. F. Barr. 1983. Pre-migratory behavior of common loons on the autumn staging grounds. Wilson Bulletin 95:121-5.
- McIntyre, J.W. 1994. Loons in freshwater lakes. Hydrobiologia 279/280:393-413.
- Mcintyre, Judith W., and Jack F. Barr. 1997. Common Loon (Gavia immer). Species Account Number 313. The Birds of North America Online (A. Poole, Ed.). Ithaca, NY: Cornell Laboratory of Ornithology; Retrieved 3/25/2008 from The Birds of North America Online database
- McPeek, G. A., and D. C. Evers. 1989. Guidelines for the protection and management of the common loon in southwest Michigan: a recovery plan. Kalamazoo Nature Center, Kalamazoo, Michigan. Unpublished report.
- Miller, E., and T. Dring. 1988. Territorial defense of multiple lakes by common loons: a preliminary report. Pages 1-14 in P. I. V. Strong (editor). Conf. on Common Loon Res. and Manage., North American Loon Fund, Meredith, New Hampshire.
- Montana Bird Distribution Committee. 1996. P. D. Skaar's Montana Bird Distribution, Fifth Edition. Special Publication No. 3. Montana Natural Heritage Program, Helena. 130 pp.
- Nelson, J. S. 1983. The tropical fish fauna in Cave and Basin Hotsprings drainage, Banff National Park, Alberta. Canadian Field-Naturalist 97:255-261.
- Okoniewski, J. C., and W. B. Stone. 1987. Some observations on the morbidity and mortality of, and environmental contaminants in, common loons (GAVIA IMMER) in New York, 1972-1986. New York State Department Environmental Conservation, Delmar, New York. Unpublished report.
- Parker, B. J., P. McKee, and R. R. Campbell. 1988 Status of the redside dace, Clinostomus elongatus, in Canada. Canadian Field-Naturalist 102:163-169.
- Parker, K. E., and R. L. Miller. 1988. Status of New York's common loon population--comparison of two intensive surveys. Pages 145-56 in P. I. V. Strong (editor). 1987 Conference on Common Loon Research and Management. North American Loon Fund, Meredith, New Hamphire.
- Paugh, J. 2006. Common loon nesting ecology in northwest Montana.
- Pokras, M.A., and R. Chafel. 1992. Lead toxicosis from ingested fishing sinkers in adult Common Loons (GAVIA IMMER) in New England. Journal of Zoo and Wildlife Medicine 23(1):92-97.
- Powers, K. D., and J. Cherry. 1983. Loon migrations off the northeastern United States. Wilson Bulletin 95:125-32.
- Ream, C.H. 1976. Loon productivity, human disturbance and pesticide residues in northern Minnesota. Wilson Bulletin 88:427-432.
- Reidman, M. L., and J. A. Estes. 1988. Predation on seabirds by sea otters. Can. J. Zool. 66:1396-1402.
- Reiser, M. H. 1988. Productivity and nest site selection of common loons in a regulated lake system. Pages 15-6 in P. I. V. Strong (editor). 1987 Conference on Common Loon Research and Management. North American Loon Fund, Meredith, New Hampshire
- Rimmer, C. C. 1992. Common loon, GAVIA IMMER. Pages 3-30 in K. J. Schneider and D. M. Pence, editors. Migratory nongame birds of management concern in the Northeast. U.S. Fish and Wildlife Service, Newton Corner, Massachusetts. 400 pp.
- Rimmer, C. C., and D. E. Kaveney. 1988. The breeding status of common loons in Vermont--1988. Vermont Inst. Nat. Sci., Woodstock, Vermont. Unpublished report.
- Root, T. 1988. Atlas of Wintering North American Birds: An Analysis of Christmas Bird Count Data. The Univ. of Chicago Press, Chicago, IL. 312 pp.
- Ruggles, A.K. 1994. Habitat selection by loons in southcentral Alaska. Hydrobiologia 279/280: 421-430.
- Salt, W.R. and J.R. Salt. 1976. The birds of Alberta. Hurtig Publishers, Edmonton, Alberta. xv + 498 pp.
- Schladweiler, Philip, and John P. Weigand., 1983, Relationships of endrin and other chlorinated hydrocarbon compounds to wildlife in Montana, 1981-1982. September 1983.
- Skaar, D. 1988. Creation of a management plan for the common loon in Montana. Pages 101-102 in: North American Loon Fund conference report (B88NOR01).
- Smith, E. L. 1981. Effects of canoeing on common loon production and survival on the Kenai National Wildlife Refuge, Alaska. Thesis. Fort Collins, Colorado: Colorado State University. 53 p.
- St. John, P. 1993. Yodelers of the north. American Birds 47(2):202-209.
- Stewart, R.E. 1975. Breeding birds of North Dakota. Tri-College Center for Environmental Studies, Fargo, North Dakota. 295 pp.
- Strong, P. I. V. 1985. Habitat selection by common loons. University of Maine, Orono, Maine. Ph.D. dissertation.
- Strong, P. I. V. 1989. Feeding and chick-rearing areas of common loons. Journal of Wildlife Management 53:72-6.
- Stroud, R. K., and R. E. Lange. 1983. Information summary: Common loon die-off winter and spring of 1983. Natl. Wildl. Health Lab., U.S. Fish and Wildlife Service, Madison, Wisconsin.
- Sutcliffe, S. A. 1978. Pesticide levels and shell thickness of common loon eggs in New Hampshire. Wilson Bulletin 90:637-40.
- U.S. Fish and Wildlife Service (USFWS). 1987. Migratory Nongame Birds of Management Concern in the United States: The 1987 List. Office of Migratory Bird Management, U.S. Fish and Wildlife Service, Washington, D.C. 63 pp.
- U.S. Forest Service. 1991. Forest and rangeland birds of the United States: Natural history and habitat use. U.S. Department of Agriculture, Forest Service Agricultural Handbook 688. 625 pages.
- Vermeer, K. 1973. Some aspects of the breeding and mortality of common loons in east-central Alberta. Canadian Field-Naturalist 87:403-8.
- Vermeer, K. 1973. Some aspects of the nesting requirements of Common Loons in Alberta. Wilson Bulletin 85:429-435.
- Williams, L. E. 1973. Spring migration of common loons from the Gulf of Mexico. Wilson Bulletin 85:238.
- Wood, R. L. 1979. Management of breeding loon populations in New Hampshire. Pages 141-6 in S. Sutcliffe (editor). Proceedings of the Second North American Conference on Common Loon Research and Management. National Audubon Society, New York, New York.
- Zicus, M. C., R. H. Hier, and S. J. Maxson. 1983. A common loon nest from Minnesota containing four eggs. Wilson Bulletin 95:672-3.
- Zimmer, G. E. 1979. The status and distribution of the common loon in Wisconsin. University of Wisconsin, Stevens Point, Wisconsin. M.S. thesis.