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Montana Field Guides

Giant Golden Moss - Homalothecium megaptilum
Other Names:  Trachybryum megaptilum

Species of Concern

Global Rank: G4
State Rank: S1
* (see State Rank Reason below)

Agency Status
USFWS:
USFS:
BLM:
MNPS Threat Rank:

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State Rank Reason (see State Rank above)
Endemic to western North America. In Montana it occurs on the eastern edge of its distribution.
 
General Description
Plant: Pleurocarpous. Large, scattered individuals to loose mats. Glossy, yellow-green to brown below. Stems 6.0-15.0 cm long and branches 1-3 cm long. Main stem occasionally is regularly pinnate with branches in one plane.

Leaf: Stem leaves: Triangular-ovate to broadly ovate-lanceolate, 3-5 x 1.5-2.0 mm, auriculate, and acuminate to filiform at the apex. Branch leaves: Ovate-lanceolate, 1.7-2.5 x 0.8-1.0, broad & blunt integrating to narrow & filiform at the apex, not auriculate, and deeply plicate with many folds. Margins recurved from near base to apex or partially plain. Apex entire to serrate. Costa strong at base, tapering towards apex, about 2/3 or more length of leaf, and often ends in a spine.

Leaf Cells: Basal cells in a few rows, shorter than median cells (25-35 x 8-10 µm), and strongly pitted. Median cells long and narrow (60-100 x 5-6 µm). Alar cells similar to basal cells and extend up along margins.

Phenology
Capsules ripen in the summer.

Diagnostic Characteristics
The large size, typically very pale green color, and its orderly pinnate branching in one plane when moist make this species distinctive (FNA 2014).

Range Comments
Canada: BC; USA: CA, ID, MT, OR, WA (FNA 2014). In Montana, known from Lake, Lincoln, and Sanders Counties (Elliott 2016).

Habitat
Litter, rock, soil, and humus in coniferous forests (Elliott 2016), and on soil over rock in areas influenced by a Pacific maritime climate. Rare below 1,080 feet. Usually between 1,080-4,300 feet elevation.

Reproductive Characteristics
Dioicous, or phyllodiocous, in which very small males grow on the leaves of the female plants. Seta 2-2.5 cm tall and rough. Capsules inclined to horizontal, zygomorphic, short and thick, 2-2.5 mm, and contracted under the mouth when dry.

References
  • Literature Cited AboveLegend:   View Online Publication
    • Elliott, J. and A. Pipp. 2016 (forthcoming). Checklist of Montana Mosses. Revised 2016. Prepared by the Montana Natural Heritage Program, Helena, Montana. 90 pp.
    • Flora of North America Editorial Committee, eds. 2014. Flora of North America North of Mexico. Volume 28. Bryophytes: Mosses, Part 2. Oxford University Press, Inc., NY. xxi + 702 pp.
  • Additional ReferencesLegend:   View Online Publication
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    • Lawton, E. 1971. Keys for the Identification of the Mosses on the Pacific Northwest. Reprinted from 'Moss Flora of the Pacific Northwest'. Published as Supplement No. 2 of the Journal of the Hattori Botanical Laboratory. Nichinan, Miyazaki, Japan. 66 pp.
    • Lawton, E. 1971. Moss Flora of the Pacific Northwest. Hattori Botanical Laboratory. Japan: Yamabuki-cho, Shinjuku-ku, Tokyo. 362 pages plus appendices.
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Citation for data on this website:
Giant Golden Moss — Homalothecium megaptilum.  Montana Field Guide.  .  Retrieved on , from